Chlaenius

Chlaenius

A: C. velutinus; B: C. spoliatus; C: C. festivus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Carabidae
Subfamily:	Harpalinae
Genus:	Chlaenius Bonelli, 1810

Chlaenius velutinus

Chlaenius purpuricollis

Chlaenius is a large and diverse genus of ground beetle. It is native to the Palearctic (including Europe), the Near East, North Africa, Afrotropical region, and the Nearctic. Worldwide, roughly 1,000 species are currently recognized ^[1] with the majority of known species occurring in the Oriental and Afrotropical regions.^[2] The genus is divided into many subgenera.^[3]

Species

Select species

Chlaenius elegans (Schmidt-Goebel, 1846)^[4]
Chlaenius festivus Panzer, 1796
Chlaenius purpuricollis Randall 1838
Chlaenius spoliatus P. Rossi, 1792

Other species include

Chlaenius abacoides Alluaud, 1933
Chlaenius aberanus Sternberg, 1908
Chlaenius aberrans Bates, 1882
Chlaenius abstersus Bates, 1873
Chlaenius accedens Chaudoir, 1876
Chlaenius acroxanthus Chaudoir, 1876
Chlaenius acutecostatus Alluaud, 1933
Chlaenius adagidensis Sternberg, 1908
Chlaenius adametzi Kuntzen, 1913
Chlaenius adonis Andrewes, 1923
Chlaenius aeneocephalus Dejean, 1826
Chlaenius aeratus Quensel, 1806
Chlaenius aestivus Say, 1823
Chlaenius aethiopicus Chaudoir, 1876
Chlaenius afganus Jedlicka, 1956
Chlaenius africanicus Kirschenhofer, 2009
Chlaenius agiloides Jedlicka, 1935
Chlaenius albissoni Reitter, 1908
Chlaenius alesi Jedlicka, 1935
Chlaenius alexanderdostali Kirschenhofer, 2008
Chlaenius allacteus Alluaud, 1919
Chlaenius allardi (Basilewsky, 1967)
Chlaenius allardianus Lorenz, 1998
Chlaenius alluaudi Fairmaire, 1901
Chlaenius alluaudianus Jedlicka, 1957
Chlaenius alternans (Imhoff, 1843)
Chlaenius alternatus G. Horn, 1871
Chlaenius alutaceus Gebler, 1829
Chlaenius amabilis Chaudoir, 1876
Chlaenius amarae Andrewes, 1920
Chlaenius amauropterus Chaudoir, 1856
Chlaenius amazonicus Chaudoir, 1876
Chlaenius ammon (Fabricius, 1801)
Chlaenius amoenus Dejean, 1831
Chlaenius amplians Bates, 1891
Chlaenius amplicollis (Basilewsky, 1953)
Chlaenius amplipennis Chaudoir, 1876
Chlaenius amydrus Alluaud, 1934
Chlaenius analis (Olivier, 1795)
Chlaenius analisimilis Pomeroy, 1932
Chlaenius anchomenoides Bates, 1889
Chlaenius andamanensis Andrewes, 1920
Chlaenius androyanus Jeannel, 1949
Chlaenius angolanus (Basilewsky, 1953)
Chlaenius angustatus Dejean, 1831
Chlaenius anischenkoi Kirschenhofer, 2010
Chlaenius annulipes Bates, 1892
Chlaenius antennatus Chaudoir, 1876
Chlaenius anthraceus Alluaud, 1918
Chlaenius anthracoderus Laferte-Senectere, 1851
Chlaenius apicalis Wiedemann, 1819
Chlaenius apollo Andrewes, 1919
Chlaenius argentinicus Jedlicka, 1946
Chlaenius armenus Jedlicka, 1950
Chlaenius arnoldi Basilewsky, 1949
Chlaenius aruwimius Bates, 1890
Chlaenius aspericollis Bates, 1873
Chlaenius assamensis Kirschenhofer, 2003
Chlaenius assecla Laferte-Senectere, 1851
Chlaenius aterrimus (Basilewsky, 1956)
Chlaenius athleta Kryzhanovskij, 1976
Chlaenius atratulus Mandl, 1983
Chlaenius atratus Chaudoir, 1876
Chlaenius atripes Chaudoir, 1876
Chlaenius atropos Andrewes, 1941
Chlaenius attenuatus Klug, 1833
Chlaenius augustus Newman, 1838
Chlaenius aurifex Basilewsky, 1949
Chlaenius auripilis Andrewes, 1936
Chlaenius australis Dejean, 1831
Chlaenius azurescens Chaudoir, 1876
Chlaenius azureulus Mandl, 1983
Chlaenius baehri (Kirschenhofer, 1998)
Chlaenius baeticus Rambur, 1837
Chlaenius balthasari Jedlicka, 1935
Chlaenius bandjermasinensis Kirschenhofer, 2008
Chlaenius bangkokensis Kirschenhofer, 2004
Chlaenius barkeri Csiki, 1931
Chlaenius basilimbatus (Grundmann, 1956)
Chlaenius baxii (Gory, 1833)
Chlaenius beatus Bates, 1891
Chlaenius bengalensis Chaudoir, 1856
Chlaenius benyovszkyi (Csiki, 1931)
Chlaenius berndjaegeri Kirschenhofer, 2008
Chlaenius bicolor Chaudoir, 1876
Chlaenius bifenestratus Klug, 1832
Chlaenius bilyi Kirschenhofer, 2009
Chlaenius bimaculatus Dejean, 1826
Chlaenius binghami Andrewes, 1919
Chlaenius bioculatus Chaudoir, 1856
Chlaenius bipustulatus Boheman, 1848
Chlaenius birmanicus Chaudoir, 1876
Chlaenius bisignatus Dejean, 1826
Chlaenius bivulnerus Motschulsky, 1858
Chlaenius bocandei (Laferte-Senectere, 1852)
Chlaenius bohemani Chaudoir, 1856
Chlaenius boisduvalii Dejean, 1831
Chlaenius borellyi Burgeon, 1941
Chlaenius borgouensis Kirschenhofer, 2007
Chlaenius bottegoi Alluaud, 1933
Chlaenius boueti (Jeannel, 1949)
Chlaenius boukali Kirschenhofer, 2005
Chlaenius braminus Chaudoir, 1876
Chlaenius brasiliensis Dejean, 1831
Chlaenius breuningi Jedlicka, 1931
Chlaenius brevilabris Le Conte, 1848
Chlaenius brevior Fairmaire, 1901
Chlaenius breviusculus Chaudoir, 1876
Chlaenius brunnescens Kirschenhofer, 2009
Chlaenius bulirschi Kirschenhofer, 2010
Chlaenius buquetii (Dejean & Boisduval, 1830)
Chlaenius buriensis Kirschenhofer, 1998
Chlaenius caecus Dejean, 1831
Chlaenius caeruleiceps (Bates, 1892)
Chlaenius caeruleicollis Chaudoir, 1876
Chlaenius caesitius Andrewes, 1923
Chlaenius caffer Boheman, 1848
Chlaenius callichloris Bates, 1873
Chlaenius cambodiensis Bates, 1889
Chlaenius camerunensis Kirschenhofer, 2003
Chlaenius canariensis Dejean, 1831
Chlaenius carbonatus Chaudoir, 1876
Chlaenius caurinus (G. Horn, 1885)
Chlaenius cavilabrum Barker, 1922
Chlaenius cebuensis Kirschenhofer, 2008
Chlaenius cecrops Kirschenhofer, 2009
Chlaenius celer Chaudoir, 1876
Chlaenius ceramensis Kirschenhofer, 2003
Chlaenius chagga Alluaud, 1927
Chlaenius chainatensis Kirschenhofer, 2011
Chlaenius chalcoderus Chaudoir, 1876
Chlaenius chalcothorax Wiedemann, 1823
Chlaenius cham Chaudoir, 1876
Chlaenius championi Andrewes, 1923
Chlaenius changwatensis (Kirschenhofer, 1998)
Chlaenius chapanus Andrewes, 1919
Chlaenius chaudoiri G. Horn, 1876
Chlaenius chengduensis Kirschenhofer, 2004
Chlaenius chiangmaiensis Kirschenhofer, 2008
Chlaenius chitwanensis Kirschenhofer, 2005
Chlaenius chlorochrous Chaudoir, 1876
Chlaenius chlorodius Dejean, 1826
Chlaenius chrysocephalus P. Rossi, 1790
Chlaenius chrysoderus Chaudoir, 1876
Chlaenius chuji Jedlicka, 1946
Chlaenius cinctus (Fabricius, 1781)
Chlaenius circumdatus Brulle, 1835
Chlaenius circumductus A. Morawitz, 1862
Chlaenius circumscriptus Duftschmid, 1812
Chlaenius clarksoni Barker, 1922
Chlaenius clypeopatens (Basilewsky, 1949)
Chlaenius coeruleus (Steven, 1809)
Chlaenius colasi Kirschenhofer, 2008
Chlaenius collarti Alluaud, 1933
Chlaenius colmanti Alluaud, 1933
Chlaenius colombensis Jedlicka, 1964
Chlaenius columbinus Dejean, 1831
Chlaenius comans Andrewes, 1919
Chlaenius comes Peringuey, 1896
Chlaenius communimacula Chaudoir in Oberthur, 1883
Chlaenius conformis Dejean, 1831
Chlaenius congoanus Alluaud, 1934
Chlaenius congoensis Burgeon, 1935
Chlaenius conradsi Kuntzen, 1913
Chlaenius consobrinus Peringuey, 1896
Chlaenius consors Peringuey, 1896
Chlaenius contractus Chaudoir, 1876
Chlaenius controversus Peringuey, 1926
Chlaenius convexus Fairmaire, 1886
Chlaenius coquereli Fairmaire, 1868
Chlaenius corbetti Andrewes, 1919
Chlaenius cordicollis Kirby, 1837
Chlaenius cordifer Bates, 1891
Chlaenius corrosulus Bates, 1892
Chlaenius coscinioderus Chaudoir,
Chlaenius coscinioderus Chaudoir, 1856
Chlaenius cosciniophorus Chaudoir, 1876
Chlaenius costiger Chaudoir, 1856
Chlaenius costipennis Boheman, 1848
Chlaenius costulatus Motschulsky, 1859
Chlaenius coxalis Fischer Von Waldheim, 1844
Chlaenius crebrepunctatus Chaudoir, 1856
Chlaenius crenistriatus Chaudoir, 1876
Chlaenius cribellatus Chaudoir, 1876
Chlaenius cribellicollis Chaudoir, 1876
Chlaenius cribricollis Dejean, 1831
Chlaenius croesus (Fabricius, 1801)
Chlaenius croyi Kirschenhofer, 2003
Chlaenius cruciatus Dejean, 1831
Chlaenius crudelis Peringuey, 1896
Chlaenius cruralis Fischer Von Waldheim, 1829
Chlaenius cubanus Chaudoir, 1876
Chlaenius cumatilis Leconte, 1851
Chlaenius cupreocinctus Reiche, 1847
Chlaenius cupreolineatus Chaudoir, 1876
Chlaenius cupreolus Fairmaire, 1901
Chlaenius cupreopurpureus Kirschenhofer, 2009
Chlaenius cupripennis Chaudoir, 1876
Chlaenius cuprithorax Quedenfeldt, 1883
Chlaenius cursor Chevrolat, 1835
Chlaenius curtulus Kirschenhofer, 2004
Chlaenius cyaneus Brulle, 1835
Chlaenius cyanipennis Boheman, 1848
Chlaenius cyanostolus Andrewes, 1924
Chlaenius cylindricollis Dejean, 1831
Chlaenius daer Darlington, 1968
Chlaenius dajuensis Kirschenhofer, 2002
Chlaenius dalibaiensis Kirschenhofer, 2008
Chlaenius daressalaami Jedlicka, 1957
Chlaenius darfurensis Kirschenhofer, 2007
Chlaenius darlingensis Castelnau, 1867
Chlaenius darlingtoni (Basilewsky, 1951)
Chlaenius davidsoni Mandl, 1978
Chlaenius decellei (Basilewsky, 1968)
Chlaenius decipiens L. Dufour, 1820
Chlaenius decorsei Alluaud, 1916
Chlaenius dejeanii Dejean, 1831
Chlaenius delicatus Bates, 1892
Chlaenius deliciolus Bates, 1873
Chlaenius delkeskampi (Basilewsky, 1956)
Chlaenius densaticollis Fairmaire, 1901
Chlaenius denticulatus Dejean, 1831
Chlaenius deplanatus (Laferte-Senectere, 1851)
Chlaenius deserticola Raffray, 1885
Chlaenius deuvei Kirschenhofer, 2008
Chlaenius deyrollei (Laferte-Senectere, 1851)
Chlaenius dhawalagiriensis Kirschenhofer, 2002
Chlaenius diabolicus (Basilewsky, 1961)
Chlaenius dianus Jedlicka, 1957
Chlaenius diephelus Alluaud, 1934
Chlaenius differens Peyron, 1858
Chlaenius difficilis Sternberg, 1908
Chlaenius dilatatus (Motschulsky, 1864)
Chlaenius dilutipes Reitter, 1894
Chlaenius dimidiatus Chaudoir, 1842
Chlaenius dinodoides Chaudoir, 1876
Chlaenius discicollis (Laferte-Senectere, 1851)
Chlaenius discopictus Fairmaire, 1893
Chlaenius dispreticus (Basilewsky, 1968)
Chlaenius dissidens Peringuey, 1926
Chlaenius distigma Chaudoir, 1876
Chlaenius ditulus Peringuey, 1904
Chlaenius dives Dejean, 1826
Chlaenius djaina Maindron, 1899
Chlaenius dohrnii (Bertoloni, 1857)
Chlaenius dondoensis Kirschenhofer, 2007
Chlaenius doriae Chaudoir, 1876
Chlaenius dorsalis Dejean, 1831
Chlaenius dostojevskji Tschitscherine, 1895
Chlaenius drescheri Louwerens, 1951
Chlaenius ducalis Chaudoir, 1876
Chlaenius dureli Maindron, 1899
Chlaenius dusaultii (L. Dufour, 1821)
Chlaenius duvaucelii (Dejean, 1831)
Chlaenius ecuadoricus Jedlicka, 1946
Chlaenius effugiens Peringuey, 1908
Chlaenius elgonensis Alluaud, 1939
Chlaenius elisabethanus Burgeon, 1935
Chlaenius elongatulus Kirschenhofer, 2009
Chlaenius elongatus (Klug, 1833)
Chlaenius emarginatus Say, 1823
Chlaenius eneides Bates, 1892
Chlaenius epicosmoides (Basilewsky, 1956)
Chlaenius episcopalis Dejean, 1831
Chlaenius epistrophus Alluaud, 1927
Chlaenius eritreaensis Kirschenhofer, 2007
Chlaenius erythrocnemis Chaudoir, 1876
Chlaenius erythropus Germar, 1824
Chlaenius eugrammus Basilewsky, 1947
Chlaenius eurybates Bates, 1891
Chlaenius euryscopus Bates, 1886
Chlaenius exaratus (Basilewsky, 1949)
Chlaenius exilis Andrewes, 1923
Chlaenius extensus Mannerheim, 1825
Chlaenius eyeni Basilewsky, 1949
Chlaenius fairmairei Murray, 1858
Chlaenius fallax (Olivier, 1795)
Chlaenius farai Jedlicka, 1949
Chlaenius farkaci Kirschenhofer, 2005
Chlaenius fasciger Chaudoir in Oberthur, 1883
Chlaenius feanus Bates, 1892
Chlaenius femoratus Dejean, 1826
Chlaenius fenestratus Chaudoir, 1876
Chlaenius feronioides Murray, 1858
Chlaenius festivus Panzer, 1796
Chlaenius fimbriatus (Klug, 1833)
Chlaenius finitimus Peringuey, 1908
Chlaenius fizianus Basilewsky, 1961
Chlaenius flavicornis Fischer Von Waldheim, 1842
Chlaenius flaviguttatus W. S. Macleay, 1825
Chlaenius flavipes Menetries, 1832
Chlaenius flavofemoratus Laporte, 1834
Chlaenius flavomaculatus (Kolbe, 1889)
Chlaenius flavoscriptus Quedenfeldt, 1891
Chlaenius fletcheri Andrewes, 1919
Chlaenius formosanus Jedlicka, 1935
Chlaenius forreri Bates, 1884
Chlaenius frater Chaudoir, 1876
Chlaenius fraterculus Maindron, 1899
Chlaenius fraternus Kolbe, 1889
Chlaenius freyellus Jedlicka, 1959
Chlaenius freyi Jedlicka, 1960
Chlaenius fugax Chaudoir, 1876
Chlaenius fulgidicollis L. Dufour, 1820
Chlaenius fulvipes (Chaudoir, 1835)
Chlaenius fuscicornis Dejean, 1831
Chlaenius gabonensis Kirschenhofer, 2009
Chlaenius gabonicus Gemminger & Harold, 1868
Chlaenius gansuensis Jedicka, 1935
Chlaenius geayi Jeannel, 1949
Chlaenius geisthardti Kirschenhofer, 2007
Chlaenius gemmingeri Ballion, 1869
Chlaenius geniculatus (Basilewsky, 1956)
Chlaenius germanus Chaudoir, 1876
Chlaenius gestroi Chaudoir, 1876
Chlaenius ghesquierei Burgeon, 1935
Chlaenius gikondo Kirschenhofer, 2009
Chlaenius glabratus Dejean, 1831
Chlaenius glabricollis Dejean, 1831
Chlaenius glaucinus Kirschenhofer, 2007
Chlaenius gonioderus Laferte-Senectere, 1851
Chlaenius gonospilus Walker, 1871
Chlaenius goossensi Alluaud, 1933
Chlaenius goryi Gory, 1833
Chlaenius gotschii Chaudoir, 1846
Chlaenius greensladei Darlington, 1971
Chlaenius greyanus White, 1841
Chlaenius grundmanni (Basilewsky, 1956)
Chlaenius guatemalenus Bates, 1882
Chlaenius guineensis Kolbe, 1883
Chlaenius gundlachi Chaudoir, 1876
Chlaenius guttula Chaudoir, 1856
Chlaenius hamatus Eschscholtz, 1829
Chlaenius hamifer Chaudoir, 1856
Chlaenius harpalinus Eschscholtz, 1833
Chlaenius heidenfelderi Kirschenhofer, 2008
Chlaenius helvetorum Burgeon, 1935
Chlaenius hemichlorus Fairmaire, 1888
Chlaenius henryi Andrewes, 1919
Chlaenius herbaceus Chevrolat, 1834
Chlaenius heyrovskyi Jedlicka, 1949
Chlaenius hildebrandti Harold, 1880
Chlaenius himalayicus Andrewes, 1923
Chlaenius holomelas (Alluaud, 1915)
Chlaenius hornburgi Kirschenhofer, 2008
Chlaenius horni Sternberg, 1908
Chlaenius hostilis Putzeys, 1880
Chlaenius howa Kunckel Dherculais, 1891
Chlaenius huedepohli Mandl, 1983
Chlaenius humeralis Chaudoir, 1856
Chlaenius humphreyi Alluaud, 1927
Chlaenius hunanensis Kirschenhofer, 2003
Chlaenius hypocrita Peringuey, 1896
Chlaenius imitatus Reitter, 1895
Chlaenius immaculatus Peringuey, 1885
Chlaenius immunitus Murray, 1858
Chlaenius imperialis Sternberg, 1908
Chlaenius impictus Alluaud, 1933
Chlaenius impunctifrons Say, 1823
Chlaenius inaequalis Fairmaire, 1901
Chlaenius inderiensis Motschulsky, 1850
Chlaenius indicus Jedlicka, 1956
Chlaenius indutus Klug, 1833
Chlaenius infantulus Chaudoir, 1876
Chlaenius infersus Peringuey, 1926
Chlaenius inops Chaudoir, 1856
Chlaenius insignis Chaudoir, 1876
Chlaenius instabilis Raffray, 1886
Chlaenius insulanus Louwerens, 1956
Chlaenius insularis (Ueno, 1964)
Chlaenius intermedius Chaudoir, 1856
Chlaenius interruptus G. Horn, 1876
Chlaenius irakensis Jedlicka, 1959
Chlaenius iranensis Kirschenhofer, 1998
Chlaenius itombwanus Basilewsky, 1961
Chlaenius jactus Kirschenhofer, 2004
Chlaenius jaechi (Kirschenhofer, 1991)
Chlaenius jakeschi Jedlicka, 1967
Chlaenius jamaicae Darlington, 1935
Chlaenius janthinus L. Redtenbacher, 1844
Chlaenius jeanneli Basilewsky, 1949
Chlaenius jordani (Basilewsky, 1955)
Chlaenius jucundulus (Basilewsky, 1970)
Chlaenius junceus Andrewes, 1923
Chlaenius jureceki Jedlicka, 1935
Chlaenius juvencus Dejean, 1831
Chlaenius kafakumbae Burgeon, 1935
Chlaenius kanarae Andrewes, 1919
Chlaenius kapangae Burgeon, 1935
Chlaenius kashmiricus (Grundmann, 1955)
Chlaenius kaszabi Jedlicka, 1951
Chlaenius katanganus Burgeon, 1935
Chlaenius keniaensis Kirschenhofer, 2008
Chlaenius kenyerii Kirschenhofer, 2003
Chlaenius kerkvoordeae Burgeon, 1935
Chlaenius kilimanus (Basilewsky, 1956)
Chlaenius kindermanni Chaudoir, 1856
Chlaenius kira Alluaud, 1919
Chlaenius kivuanus Basilewsky, 1949
Chlaenius kivuensis Alluaud, 1933
Chlaenius klapperichi Jedlicka, 1956
Chlaenius koenigi Semenov, 1888
Chlaenius kolariensis Maindron, 1898
Chlaenius kolbei (Duvivier, 1892)
Chlaenius kotys Kirschenhofer, 2004
Chlaenius kraatzi Sternberg, 1908
Chlaenius kryzhanovskyi Basilewsky, 1968
Chlaenius kulti Basilewsky, 1949
Chlaenius kurosawai Kasahara, 1986
Chlaenius labroexcisus Basilewsky, 1948
Chlaenius lacunosus Andrewes, 1920
Chlaenius ladon Kirschenhofer, 1998
Chlaenius laetiusculus Chaudoir, 1856
Chlaenius laetoides Burgeon, 1935
Chlaenius laetus (Fabricius, 1794)
Chlaenius laevicollis Peringuey, 1926
Chlaenius laevipennis Chaudoir, 1876
Chlaenius laeviplaga Chaudoir, 1876
Chlaenius laeviusculus Chaudoir, 1856
Chlaenius lafertei Guerin-Meneville, 1843
Chlaenius lamottei (Basilewsky, 1951)
Chlaenius langsonensis Kirschenhofer, 2008
Chlaenius laosensis Kirschenhofer, 2008
Chlaenius laotinus Andrewes, 1919
Chlaenius lapillus Basilewsky, 1949
Chlaenius lastii Bates, 1886
Chlaenius lateralis Brulle, 1838
Chlaenius latesternalis (Basilewsky, 1956)
Chlaenius laticollis Say, 1823
Chlaenius latipalpis Mandl, 1992
Chlaenius latipennis Sternberg, 1908
Chlaenius latithorax Mannerheim, 1844
Chlaenius lativittis Chaudoir, 1876
Chlaenius latreillei (Laferte-Senectere, 1852)
Chlaenius latro Laferte-Senectere, 1851
Chlaenius lederi Reitter, 1888
Chlaenius leishanensis Kirschenhofer, 2005
Chlaenius leleupi (Basilewsky, 1952)
Chlaenius leopoldi Burgeon, 1935
Chlaenius leprieuri Gory, 1833
Chlaenius leptopus Basilewsky, 1956
Chlaenius leucops (Wiedemann, 1823)
Chlaenius leucoristus Chaudoir, 1876
Chlaenius leucoscelis Chevrolat, 1835
Chlaenius leytensis Kirschenhofer, 2008
Chlaenius limbatus Wiedemann, 1821
Chlaenius limbicollis Chaudoir, 1876
Chlaenius limbipennis Boheman, 1860
Chlaenius lineatus Putzeys, 1880
Chlaenius lineellus Motschulsky, 1859
Chlaenius liocephalus Basilewsky, 1952
Chlaenius lioderus Andrewes, 1923
Chlaenius liothorax Alluaud, 1934
Chlaenius lirifer Andrewes, 1941
Chlaenius lissoderus Chaudoir, 1876
Chlaenius lithophilus Say, 1823
Chlaenius litongaensis Kirschenhofer, 2009
Chlaenius loango Kirschenhofer, 2009
Chlaenius lobozianus Alluaud, 1934
Chlaenius loeblianus Kirschenhofer, 2004
Chlaenius lombokensis Kirschenhofer, 2008
Chlaenius lomii G. Muller, 1941
Chlaenius lomsakensis Kirschenhofer, 1998
Chlaenius longeantennatus (Basilewsky, 1951)
Chlaenius longicornis Chaudoir, 1843
Chlaenius longulus (Jeannel, 1949)
Chlaenius louwerensi Andrewes, 1936
Chlaenius loveridgei (Basilewsky, 1951)
Chlaenius loxias Kirschenhofer, 2004
Chlaenius lucasi Payron, 1858
Chlaenius lucasi Peyron, 1858
Chlaenius lucidicollis Laferte-Senectere, 1851
Chlaenius lucidulus (Boheman, 1848)
Chlaenius luculentus Andrewes, 1920
Chlaenius lugens Chaudoir, 1876
Chlaenius luisae Gestro, 1895
Chlaenius lujai Burgeon, 1935
Chlaenius lulengae Burgeon, 1935
Chlaenius lunatus Dejean, 1826
Chlaenius luteicauda Chaudoir, 1876
Chlaenius luzonicus Chaudoir, 1856
Chlaenius lyperus (Jeannel, 1949)
Chlaenius macropus Chaudoir, 1876
Chlaenius maculatus Dejean, 1826
Chlaenius maculiceps Boheman, 1848
Chlaenius maculiger Castelnau, 1867
Chlaenius madecassus Csiki, 1931
Chlaenius madrasensis Kirschenhofer, 2004
Chlaenius makalolo Bates, 1886
Chlaenius malachinus (Motschulsky, 1865)
Chlaenius malcheri Emden, 1937
Chlaenius mandli Grundmann, 1956
Chlaenius manowianus Jedlicka, 1957
Chlaenius manus Louwerens, 1969
Chlaenius maowensis Kirschenhofer, 2008
Chlaenius maputoensis Kirschenhofer, 2007
Chlaenius marginipennis Gory, 1833
Chlaenius marianensis Darlington, 1970
Chlaenius marleyi Barker, 1922
Chlaenius martinbaehri Kirschenhofer, 2008
Chlaenius masoni Andrewes, 1923
Chlaenius maxillosus G. Horn, 1876
Chlaenius medioguttatus Chaudoir, 1876
Chlaenius mediornatus (Basilewsky, 1956)
Chlaenius melampus Menetries, 1849
Chlaenius melancholicus Laferte-Senectere, 1851
Chlaenius melanocnemis Basilewsky, 1949
Chlaenius melanopterus Chaudoir, 1876
Chlaenius melanopus Andrewes, 1923
Chlaenius merkli Kirschenhofer, 2003
Chlaenius meteorus Alluaud, 1939
Chlaenius meticulosus Laferte-Senectere, 1851
Chlaenius micans (Fabricius, 1792)
Chlaenius microspilus (Bates, 1892)
Chlaenius milleti Antoine, 1932
Chlaenius milloti Jeannel, 1949
Chlaenius moaboensis Kirschenhofer, 2009
Chlaenius moestus Csiki, 1931
Chlaenius moheliensis Kirschenhofer. 2009
Chlaenius moluccensis Kirschenhofer, 2003
Chlaenius momboensis Kirschenhofer, 2007
Chlaenius monardi Alluaud, 1934
Chlaenius montanus Lucas, 1859
Chlaenius montivagus Andrewes, 1923
Chlaenius morettoi Kirschenhofer, 2008
Chlaenius morio Boheman, 1860
Chlaenius morosus Laferte-Senectere, 1851
Chlaenius mpanga Kirschenhofer, 2009
Chlaenius muehlei Kirschenhofer, 2004
Chlaenius multicolor Andrewes, 1919
Chlaenius mutatus Gemminger & Harold, 1868
Chlaenius naeviger A. Morawitz, 1862
Chlaenius nanlingensis Deuve & Tian, 2005
Chlaenius nanpingensis Kirschenhofer, 1999
Chlaenius natalensis Chaudoir, 1876
Chlaenius nebraskensis Lecont
Chlaenius nebrioides Antoine, 1961
Chlaenius nemoralis Say, 1823
Chlaenius neocaledonicus Chaudoir in Oberthur, 1883
Chlaenius neochloodes Lorenz, 1998
Chlaenius nepalensis Hope, 1831
Chlaenius nepos Chaudoir, 1876
Chlaenius newdehliensis Kirschenhofer, 2002
Chlaenius niger Randall, 1838
Chlaenius nigerrimus Jedlicka, 1958
Chlaenius nigrans Kirschenhofer, 2003
Chlaenius nigratus Kirschenhofer, 2007
Chlaenius nigricans Wiedemann, 1821
Chlaenius nigricornis Fabricius, 1787
Chlaenius nigripennis Chaudoir, 1856
Chlaenius nigrita Dejean, 1826
Chlaenius nigroscelis Chaudoir, 1856
Chlaenius nigrosuturatus Mandl, 1978
Chlaenius nilgiricus Andrewes, 1919
Chlaenius nimbanus (Basilewsky, 1951)
Chlaenius nitidiceps Dejean, 1826
Chlaenius nitidicollis Dejean, 1826
Chlaenius nitidifrons Fairmaire, 1901
Chlaenius nitidulus Schrank, 1781
Chlaenius noeli Alluaud, 1933
Chlaenius noguchii Bates, 1873
Chlaenius notabilis Laferte-Senectere, 1851
Chlaenius nubicus Chaudoir, 1876
Chlaenius nyassaensis Jedlicka, 1957
Chlaenius nyika Kirschenhofer, 2009
Chlaenius obenbergeri Jedlicka, 1935
Chlaenius oberthuri Sternberg, 1908
Chlaenius obesus Laferte-Senectere, 1851
Chlaenius obliquatus Barker, 1922
Chlaenius oblongus Dejean, 1826
Chlaenius obscuripennis Chevrolat, 1835
Chlaenius obscurus Klug, 1832
Chlaenius obsidianoides Alluaud, 1935
Chlaenius obsidianus Alluaud, 1935
Chlaenius obtusus Dejean, 1831
Chlaenius occultus Sloane, 1907
Chlaenius ochroperas Bates, 1892
Chlaenius ocreatus Bates, 1873
Chlaenius oculatus (Fabricius, 1801)
Chlaenius oligochrysus Alluaud, 1934
Chlaenius olivieri Crotch, 1871
Chlaenius olthofi Darlington, 1968
Chlaenius omochlorus Andrewes, 1931
Chlaenius oneili Barker, 1922
Chlaenius oodioides Chaudoir, 1876
Chlaenius opacipennis Chaudoir, 1876
Chlaenius ophonoides Fairmaire, 1843
Chlaenius orbicollis Chaudoir, 1876
Chlaenius orbiculicollis Barker, 1922
Chlaenius orbus G. Horn, 1871
Chlaenius orphanus Peringuey, 1908
Chlaenius ostrinus Andrewes, 1924
Chlaenius ovalipennis Quedenfeldt, 1883
Chlaenius ovampo Peringuey, 1892
Chlaenius overlaeti Burgeon, 1935
Chlaenius pacholatkoi Kirschenhofer, 2003
Chlaenius pachys Chaudoir, 1876
Chlaenius paenulatus Erichson, 1843
Chlaenius pallidicornis Ballion, 1871
Chlaenius pallipes (Gebler, 1823)
Chlaenius palpalis Laferte-Senectere, 1851
Chlaenius paluensis Kirschenhofer, 2011
Chlaenius pan Darlington, 1968
Chlaenius panagaeoides Chaudoir, 1876
Chlaenius panjabensis Kirschenhofer, 1998
Chlaenius parallelus Dejean, 1831
Chlaenius parepus Jedlicka, 1957
Chlaenius paromius Basilewsky, 1949
Chlaenius patricius Harold, 1879
Chlaenius patrizii (Basilewsky, 1953)
Chlaenius patruelis Leconte, 1844
Chlaenius pauxillus Kirschenhofer, 2002
Chlaenius paykulli Crotch, 1870
Chlaenius pectinipes Bates, 1892
Chlaenius peltastes Jedlicka, 1935
Chlaenius pennsylvanicus Say, 1823
Chlaenius pericallus L. Redtenbacher, 1867
Chlaenius peringueyi Kuntzen, 1919
Chlaenius perpunctatus Kuntzen, 1913
Chlaenius perrieri (Fairmaire, 1899)
Chlaenius persimilis Chaudoir, 1876
Chlaenius perspicillaris Erichson, 1843
Chlaenius peruanus Erichson, 1847
Chlaenius peterseni (Louwerens, 1967)
Chlaenius phaenoderus Chaudoir, 1876
Chlaenius pharaonis Motschulsky, 1865
Chlaenius phenax Basilewsky, 1949
Chlaenius philemon Andrewes, 1936
Chlaenius piceus Chaudoir, 1876
Chlaenius pictus Chaudoir, 1856
Chlaenius piligenys Liu & Liang, 2010
Chlaenius pilosicoerulans Mandl, 1989
Chlaenius planipennis Chaudoir, 1876
Chlaenius planulatus Bates, 1884
Chlaenius platensis G. R. Waterhouse, 1841
Chlaenius platyderus Chaudoir, 1856
Chlaenius platynoides Alluaud, 1934
Chlaenius plausibilis Basilewsky, 1949
Chlaenius pleuroderus Chaudoir in Oberthur, 1883
Chlaenius poecilinus Bates, 1892
Chlaenius porinus Alluaud, 1929
Chlaenius porphyrius Bates, 1891
Chlaenius porrectus Chaudoir, 1876
Chlaenius posticalis Motschulsky, 1854
Chlaenius posticus (Fabricius, 1798)
Chlaenius postmaculatus (Basilewsky, 1952)
Chlaenius postscriptus Bates, 1873
Chlaenius pradieri Chaudoir, 1876
Chlaenius praefectus Bates, 1873
Chlaenius prasinus Dejean, 1826
Chlaenius pratensis Chaudoir, 1876
Chlaenius pretiosus Chaudoir, 1856
Chlaenius principalis Sternberg, 1908
Chlaenius privatus Bates, 1892
Chlaenius privatus Bates, 1892
Chlaenius probsti Kirschenhofer, 2004
Chlaenius prolixus Erichson, 1843
Chlaenius propinquus Csiki, 1931
Chlaenius prostenus Bates, 1873
Chlaenius prostenus Bates, 1873
Chlaenius protensus Chaudoir, 1876
Chlaenius pseudocruciatus Kirschenhofer, 2009
Chlaenius pseudocupreolus Kirschenhofer, 2010
Chlaenius pseudoglabratus Kirschenhofer, 2009
Chlaenius pseudolulengae Kirschenhofer, 2009
Chlaenius pseudopalpalis Basilewsky, 1949
Chlaenius pseudoraffrayi Basilewsky, 1949
Chlaenius pterostichoides Andrewes, 1941
Chlaenius ptuchodes Andrewes, 1923
Chlaenius puberulus Boheman, 1848
Chlaenius pubifer Chaudoir, 1876
Chlaenius pubipennis Chaudoir, 1856
Chlaenius puchneri Kirschenhofer, 2007
Chlaenius pudicus (Fabricius, 1801)
Chlaenius pugni Camerano, 1879
Chlaenius pulchellus Boheman, 1848
Chlaenius pulcher Nietner, 1857
Chlaenius pulchriceps Fairmaire, 1901
Chlaenius pumilio Kolbe, 1889
Chlaenius punctatostriatus Chaudoir, 1856
Chlaenius puncticollis Dejean, 1826
Chlaenius purpureus Chaudoir, 1876
Chlaenius purpuricollis Randall, 1838
Chlaenius pusillus Say, 1823
Chlaenius putzeysi Chaudoir, 1876
Chlaenius pyrrhopodus Fairmaire, 1903
Chlaenius pyrrhos Kirschenhofer, 2004
Chlaenius quadricolor (Olivier, 1790)
Chlaenius quadrimaculatus (Boheman, 1848)
Chlaenius quadrinotatus Dejean, 1826
Chlaenius quadriornatus Basilewsky, 1956
Chlaenius quadrisignatus Boheman, 1860
Chlaenius quadrisulcatus Paykull, 1790
Chlaenius quadrivittatus Jedlicka, 1958
Chlaenius radama Kunckel, 1891
Chlaenius raffrayi Chaudoir, 1876
Chlaenius rambouseki Lutshnik, 1933
Chlaenius ranavalonae Csiki, 1931
Chlaenius rebellis Peringuey, 1926
Chlaenius reichei (Laferte-Senectere, 1851)
Chlaenius reitteri Jakobson, 1906
Chlaenius retropictus Fairmaire, 1901
Chlaenius rhodesianus Peringuey, 1899
Chlaenius richardsi Ali, 1967
Chlaenius riparius Lorenz, 1998
Chlaenius ripicola Andrewes, 1937
Chlaenius robustus Boheman, 1848
Chlaenius rodriguezi Chaudoir, 1876
Chlaenius roeschkei Sternberg, 1908
Chlaenius rotundatulus Lorenz, 1998
Chlaenius rotundithorax Liu & Kavanaugh, 2010
Chlaenius rotundus Andrewes, 1920
Chlaenius rubenticollis Kirschenhofer, 2009
Chlaenius rubricrus Alluaud, 1916
Chlaenius rudesculptus Chaudoir, 1876
Chlaenius rudicollis Chaudoir, 1876
Chlaenius ruficauda Chaudoir, 1856
Chlaenius rufifemoratus Macleay, 1825
Chlaenius rufithorax Wiedemann, 1821
Chlaenius rufomarginatus Dejean, 1831
Chlaenius rugulosus Nietner, 1857
Chlaenius rukwaensis Kirschenhofer, 2007
Chlaenius ruvumaensis Kirschenhofer, 2008
Chlaenius rysonotus Fairmaire, 1901
Chlaenius sabahensis Kirschenhofer, 1998
Chlaenius saginatus Laferte-Senectere, 1851
Chlaenius sagittarius Dejean, 1831
Chlaenius salisburiensis Barker, 1922
Chlaenius sallei Chaudoir, 1876
Chlaenius samoensis Csiki, 1915
Chlaenius sangaicus Alluaud, 1929
Chlaenius sankuruensis Burgeon, 1935
Chlaenius savanicola (Basilewsky, 1968)
Chlaenius scabricollis Chevrolat, 1833
Chlaenius scapularis Chaudoir, 1876
Chlaenius scheerpeltzi (Basilewsky, 1956)
Chlaenius schillhammeri Kirschenhofer, 2003
Chlaenius schmidti Gestro, 1895
Chlaenius schoenherri (Dejean, 1831)
Chlaenius schoutedeni Burgeon, 1935
Chlaenius schuelei Kirschenhofer, 2007
Chlaenius sciakyi Kirschenhofer, 2004
Chlaenius scotti Alluaud, 1937
Chlaenius sculptilis Bates, 1886
Chlaenius scutellaris Harold, 1880
Chlaenius sehnali Kirschenhofer, 2008
Chlaenius seiferti Kirschenhofer, 2005
Chlaenius sellatus Dejean, 1831
Chlaenius semenowi Tschitscherine, 1895
Chlaenius semicupreus Basilewsky, 1949
Chlaenius semicyaneus Solsky, 1874
Chlaenius semipurpureus Motschulsky, 1865
Chlaenius senegalensis Dejean, 1831
Chlaenius sennaariensis Chaudoir, 1856
Chlaenius sericeus (Forster, 1771)
Chlaenius sericimicans Chaudoir, 1876
Chlaenius sexguttatus Andrewes, 1919
Chlaenius sexmaculatus Dejean, 1831
Chlaenius seyrigi Alluaud, 1935
Chlaenius siccus Darlington, 1968
Chlaenius sichuanensis Kirschenhofer, 2008
Chlaenius sierraleonensis Kirschenhofer, 2008
Chlaenius siffointei (Basilewsky, 1968)
Chlaenius silvestrii G. Muller, 1942
Chlaenius simba Alluaud, 1929
Chlaenius simbabwensis Kirschenhofer, 2008
Chlaenius similatus Boheman, 1848
Chlaenius similis Chaudoir, 1856
Chlaenius simplex Wiedemann, 1821
Chlaenius simulatorius Barker, 1922
Chlaenius sinensis Chaudoir, 1856
Chlaenius sivorii Chaudoir, 1876
Chlaenius sobrinus Dejean, 1826
Chlaenius soccatus Say, 1830
Chlaenius sokotranus Csiki, 1931
Chlaenius solitarius Say, 1823
Chlaenius sollicitus Laferte-Senectere, 1851
Chlaenius somaliae Basilewsky, 1956
Chlaenius somereni Alluaud, 1929
Chlaenius soricinus Gerstaecker, 1867
Chlaenius sparsepunctatus Chaudoir, 1876
Chlaenius spathulifer (Bates, 1873)
Chlaenius specularis Emden, 1937
Chlaenius splendidus Dejean, 1831
Chlaenius spoliatus P. Rossi, 1792
Chlaenius startellus Basilewsky, 1949
Chlaenius steveni Quensel, 1806
Chlaenius stichai Jedlicka, 1949
Chlaenius straneoi Basilewsky, 1949
Chlaenius stschukini Menetries, 1837
Chlaenius stungtrengensis Kirschenhofer, 2004
Chlaenius stygius (Laferte-Senectere, 1851)
Chlaenius suavis Alluaud, 1939
Chlaenius subcostatus W. J. Macleay, 1864
Chlaenius suberbiei (Jeannel, 1949)
Chlaenius subferrugineus (Kirschenhofer, 2003)
Chlaenius subovatus Chaudoir, 1876
Chlaenius subsulcatus Dejean, 1831
Chlaenius suensoni Mandl, 1992
Chlaenius sulcicollis (Paykull, 1798)
Chlaenius sulcipennis Dejean, 1826
Chlaenius sumptuosus Alluaud, 1934
Chlaenius superbus Sternberg, 1908
Chlaenius superstes Peringuey, 1926
Chlaenius suppletor Bates, 1891
Chlaenius surdipennis (Jeannel, 1949)
Chlaenius suvorovi Semenov, 1912
Chlaenius swahilius Bates, 1886
Chlaenius syangyaensis Kirschenhofer, 2004
Chlaenius sykesi Hope, 1835
Chlaenius synaptus Alluaud, 1918
Chlaenius syriacus Chaudoir, 1876
Chlaenius systolocranioides Alluaud, 1933
Chlaenius tamdaoensis Kirschenhofer, 2003
Chlaenius tanahrataensis Kirschenhofer, 2008
Chlaenius teani Gestro, 1881
Chlaenius tecospilus Alluaud, 1933
Chlaenius tenellus Klug, 1832
Chlaenius tenuicollis (Fabricius, 1801)
Chlaenius tenuilimbatus Ballion, 1871
Chlaenius tenuis Fairmaire, 1901
Chlaenius terminatus Dejean, 1826
Chlaenius terminatus Dejean, 1826
Chlaenius testaceicrus Fairmaire, 1891
Chlaenius tetracelis Alluaud, 1933
Chlaenius tetragonoderus Chaudoir, 1876
Chlaenius tetraphacus Alluaud, 1918
Chlaenius tetraspilus Alluaud, 1918
Chlaenius thieleni (Kirschenhofer, 1998)
Chlaenius thiesensis Kirschenhofer, 2007
Chlaenius tibialis Dejean, 1826
Chlaenius timorensis Darlington, 1971
Chlaenius tinantae Burgeon, 1935
Chlaenius titschacki Jedlicka, 1946
Chlaenius tomentosus Say, 1823
Chlaenius toubaensis Kirschenhofer, 2008
Chlaenius touzalini Andrewes, 1920
Chlaenius trachys Andrewes, 1923
Chlaenius transfuga Chaudoir, 1876
Chlaenius transversalis Dejean, 1831
Chlaenius trapezicollis Chaudoir, 1856
Chlaenius treichi Alluaud, 1916
Chlaenius trichrous Alluaud, 1918
Chlaenius tricolor Dejean, 1826
Chlaenius trigonotomoides Emden, 1928
Chlaenius trinotatus Laferte-Sinectere, 1851
Chlaenius tripustulatus (Dejean, 1831)
Chlaenius tristis Schaller, 1783
Chlaenius tschitscherini Jedlicka, 1952
Chlaenius tshibindensis Burgeon, 1935
Chlaenius tudicus Andrewes, 1919
Chlaenius tuky Basilewsky, 1949
Chlaenius uluguruanus Basilewsky, 1956
Chlaenius umtalianus (Peringuey, 1904)
Chlaenius unicolor Chaudoir, 1856
Chlaenius uninotatus Andrewes, 1919
Chlaenius unyorensis Alluaud, 1934
Chlaenius uzungwensis Basilewsky, 1951
Chlaenius vadoni (Basilewsky, 1950)
Chlaenius validicornis Boheman, 1848
Chlaenius varians Chaudoir, 1876
Chlaenius variicornis A. Morawitz, 1863
Chlaenius variipes Chaudoir, 1856
Chlaenius variolosus Kirschenhofer, 2008
Chlaenius vartianorum Mandl, 1989
Chlaenius velocipes Chaudoir, 1876
Chlaenius venustulus Dejean, 1831
Chlaenius vertagoides (Laferte-Senectere, 1851)
Chlaenius veselyi Kirschenhofer, 2004
Chlaenius vestitus (Paykull, 1790)
Chlaenius vethi Bates, 1889
Chlaenius viangchanensis Kirschenhofer, 1998
Chlaenius victor Andrewes, 1928
Chlaenius victoriae Kirschenhofer, 2009
Chlaenius viduus G. Horn, 1871
Chlaenius vietnamensis Kirschenhofer, 2008
Chlaenius vietnami Jedlicka, 1966
Chlaenius violaceipennis Chaudoir, 1876
Chlaenius violatus Gemminger & Harold, 1868
Chlaenius virens Rambur, 1837
Chlaenius virescens Chaudoir, 1835
Chlaenius virgulatus (Jeannel, 1949)
Chlaenius virgulifer Chaudoir, 1876
Chlaenius viridicollis Reiche, 1843
Chlaenius viridis Menetries, 1832
Chlaenius vitalisi Andrewes, 1919
Chlaenius vitticollis Boheman, 1848
Chlaenius vividus Chaudoir, 1876
Chlaenius vulneratus Dejean, 1831
Chlaenius waddellii Murray, 1858
Chlaenius wallacei Chaudoir, 1876
Chlaenius wegneri Louwerens, 1953
Chlaenius wenchuanensis Kirschenhofer, 2008
Chlaenius wittei Burgeon, 1937
Chlaenius wittmerianus Mandl, 1978
Chlaenius wrasei (Kirschenhofer, 1997)
Chlaenius xanthacrus Wiedemann, 1823
Chlaenius xanthomerus Alluaud, 1918
Chlaenius xanthospilus (Wiedemann, 1821)
Chlaenius yangonensis Kirschenhofer, 2005
Chlaenius yunnanulus Mandl, 1992
Chlaenius zanzibaricus Chaudoir in Oberthur, 1883
Chlaenius ziloensis Basilewsky, 1970
Chlaenius zygogrammus Laferte-Senectere, 1851

References

↑ Lorenz, W. 2005. Systematic list of extant ground beetles of the world. Second Edition. Tizing, Germany. 530pp.
↑ Bousquet, Y. 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. Zookeys 245:1-1722.
↑ "Chlaenius Bonelli, 1810". Carabidae of the World. 2011. Retrieved 13 Jul 2011.
↑ Faunula coleopterorum Birmaniae, adjectis nonnullis Bengaliae indiginis. Schmidt-Göbel, H. M. (Hermann Maximilian) 1846

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