Anomodont

Anomodonts
Temporal range: Middle Permian-Late Triassic, 270–201 Ma (Possible Early Cretaceous record.)[1]
Mounted skeleton of Lystrosaurus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Order: Therapsida
Clade: Neotherapsida
Suborder: Anomodontia
Owen, 1859
Subgroups

See text

Anomodontia is an extinct group of non-mammalian therapsids containing many species from the Permian and Triassic periods (possibly continuing into the Early Cretaceous),[2] most of which were toothless, possibly endothermic[3][4][5] herbivores.[6] Anomodonts were very diverse during the Middle Permian, including primitive forms like Anomocephalus and Patranomodon and groups like Venyukovioidea, Dromasauria, and Dicynodontia. Of these, only the dicynodonts survived beyond the Middle Permian. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian and Triassic, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage evolved into large, stocky forms that remained the dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline.

Classification

Taxonomy

Phylogeny

Cladogram modified from Liu et al. (2009):[2]

Therapsida 

unnamed


Biarmosuchia



Gorgonopsia



unnamed


Dinocephalia


 Anomodontia 


Biseridens


unnamed


Anomocephalus


unnamed

 Venyukovioidea 


Otsheria


unnamed


Ulemica



Suminia




 Chainosauria 


Patranomodon


unnamed


Galeops



Eodicynodon









Below is a cladogram from Kammerer et al. (2013).[7] The data matrix of Kammerer et al. (2013), a list of characteristics that was used in the analysis, was based on that of Kammerer et al. (2011), which followed a comprehensive taxonomic revision of Dicynodon.[8] Because of this, many of the relationships found by Kammerer et al. (2013) are the same as those found by Kammerer et al. (2011). However, several taxa were added to the analysis, including Tiarajudens Eubrachiosaurus, Shaanbeikannemeyeria, Zambiasaurus and many "outgroup" taxa (positioned outside Anomodontia), while other taxa were re-coded. As in Kammerer et al. (2011), the interrelationships of non-kannemeyeriiform dicynodontoids are weakly supported and thus vary between the analyses.[7]

1 


Biseridens





Anomocephalus



Tiarajudens





Patranomodon



 2 


Suminia




Otsheria



Ulemica




 3 



Galepus



Galechirus





Galeops


 4 


"Eodicynodon" oelofseni





Eodicynodon oosthuizeni



Colobodectes





Lanthanostegus




Chelydontops




Endothiodon




Pristerodon


 5 

 6 

 7 


Diictodon




Eosimops




Prosictodon



Robertia





 8 


Emydops


 9 

 10 


Dicynodontoides



Kombuisia





Myosaurus


 11 


Cistecephalus




Cistecephaloides



Kawingasaurus









 12 














1 Anomodontia, 2 Venyukovioidea, 3 Chainosauria, 4 Dicynodontia, 5 Therochelonia, 6 Diictodontia, 7 Pylaecephalidae, 8 Emydopoidea, 9 Kistecephalia, 10 Kingoriidae, 11 Cistecephalidae, 12 Bidentalia


12 

 13 


Keyseria




Daqingshanodon



 14 


Oudenodon




Tropidostoma



Australobarbarus







Odontocyclops



Idelesaurus




 15 


Rhachiocephalus



Kitchinganomodon



 16 


Syops


 17 


Aulacocephalodon




Pelanomodon




Geikia elginensis



Geikia locusticeps











 18 



Interpresosaurus




Elph



Katumbia






Gordonia





Basilodon



Sintocephalus





Dicynodon lacerticeps




"Dicynodon" huenei





Delectosaurus



Vivaxosaurus






Daptocephalus




Dinanomodon



Peramodon






Jimusaria




Turfanodon



 19 


Euptychognathus




Lystrosaurus murrayi




"Lystrosaurus" declivus




"Lystrosaurus" curvatus




"Lystrosaurus" maccaigi




"Lystrosaurus" hedini




TSK 2



Kwazulusaurus










 20 












12 Bidentalia, 13 Cryptodontia, 14 Oudenodontidae, 15 Rhachiocephalidae, 16 Geikiidae, 17 Geikiinae, 18 Dicynodontoidea, 19 Lystrosauridae, 20 Kannemeyeriiformes


20 


Angonisaurus



 21 



Tetragonias



Vinceria





Shansiodon



Rhinodicynodon






Dinodontosaurus





Shaanbeikannemeyeria




Kannemeyeria lophorhinus



Kannemeyeria simocephalus







Parakannemeyeria



Xiyukannemeyeria






Dolichuranus




Rechnisaurus



Uralokannemeyeria






Rabidosaurus




Sinokannemeyeria





Rhadiodromus



Wadiasaurus



 22 

 23 


Zambiasaurus




Moghreberia



Placerias




 24 


Stahleckeria





Eubrachiosaurus



Sangusaurus





Jachaleria



Ischigualastia














20 Kannemeyeriiformes, 21 Shansiodontidae, 22 Stahleckeriidae, 23 Placeriinae, 24 Stahleckeriinae


See also

References

  1. 1 2 Liu, J.; Rubidge, B.; Li, J. (2009). "A new specimen of Biseridens qilianicus indicates its phylogenetic position as the most basal anomodont". Proceedings of the Royal Society B. 277 (1679): 285–292. PMC 2842672Freely accessible. PMID 19640887. doi:10.1098/rspb.2009.0883.
  2. Bakker 1975
  3. BOTHA-BRINK, Jennifer; ANGIELCZYK, Kenneth D. "Do extraordinarily high growth rates in Permo-Triassic dicynodonts (Therapsida, Anomodontia) explain their success before and after the end-Permian extinction?". Zoological Journal of the Linnean Society. 160 (2): 341–365. doi:10.1111/j.1096-3642.2009.00601.x.
  4. "Microbiota and food residues including possible evidence of pre-mammalian hair in Upper Permian coprolites from Russia". Lethaia. doi:10.1111/let.12156.
  5. Chinsamy-Turan, A. (2011) Forerunners of Mammals: Radiation - Histology - Biology, p.39. Indiana University Press, ISBN 0253356970. Retrieved May 2012
  6. 1 2 Kammerer, C. F.; Fröbisch, J. R.; Angielczyk, K. D. (2013). Farke, Andrew A, ed. "On the Validity and Phylogenetic Position of Eubrachiosaurus browni, a Kannemeyeriiform Dicynodont (Anomodontia) from Triassic North America". PLoS ONE. 8 (5): e64203. PMC 3669350Freely accessible. PMID 23741307. doi:10.1371/journal.pone.0064203.
  7. Kammerer, C.F.; Angielczyk, K.D.; Fröbisch, J. (2011). "A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia) and its implications for dicynodont phylogeny, biogeography, and biostratigraphy". Journal of Vertebrate Paleontology. 31 (Suppl. 1): 1–158. doi:10.1080/02724634.2011.627074.
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