Andesaurus

Andesaurus
Temporal range: Cenomanian
99.6–97 Ma
↓
Restoration of Andesaurus and Giganotosaurus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Superorder: Dinosauria
Order: Saurischia
Suborder: Sauropodomorpha
Infraorder: Sauropoda
(unranked): Titanosauria
Family: Andesauridae
Calvo & Bonaparte, 1991
Genus: Andesaurus
Binomial name
Andesaurus delgadoi
Calvo & Bonaparte, 1991

Andesaurus (/ˌændᵻˈsɔːrəs/ AN-də-SAWR-əs; "Andes lizard") is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, it would have had a small head on the end of a long neck and an equally long tail. Andesaurus was a very large animal, as were many others of its relatives, which included the largest animals ever to walk the Earth.

History of discovery

In 1991, paleontologists Jorge Orlando Calvo and José Fernando Bonaparte named Andesaurus, which refers to the Andes and also includes the Greek word sauros ("lizard"), because of the proximity of this animal's remains to the Andes. Andesaurus fossils were found by Alejandro Delgado, after whom the single known species (A. delgadoi) is named.

The only known material of Andesaurus is a partial skeleton consisting of a series of four vertebrae from the lower back, as well as 27 tail vertebrae, divided up into two series from separate parts of the tail. The vertebrae from the middle part of its tail had elongated centra.[1] Elements of the pelvis were also discovered, including two ischia and a pubis bone, along with rib fragments and an incomplete humerus and femur. Size estimates place Andesaurus at roughly 15 metres (49 ft)[2] to 18 metres (59 ft) in length,[3] and 7 metric tons (7.7 short tons) in weight.[2]

Classification

Several plesiomorphic (primitive) features characterize Andesaurus as the most basal known member of Titanosauria. In fact, this clade has been defined to contain Andesaurus, Saltasaurus, their most recent common ancestor, and all of its descendants.[4] The most prominent plesiomorphy is the articulations between its tail vertebrae. In most derived titanosaurs, the tail vertebrae articulate with ball-and-socket joints, with the hollowed-out socket end on the front (procoelous caudal vertebrae), while in Andesaurus, both ends of the vertebrae are flat (amphiplatyan caudals), as seen in many non-titanosaurian sauropods. Andesaurus itself is only characterized by a single feature, the tall neural spines on top of its back vertebrae, and needs further study.[5]

Some other basal titanosaurs from Argentina, including Argentinosaurus and Puertasaurus, were also sauropods of enormous size. The most derived group of titanosaurs, the Saltasauridae, included some of the smallest known sauropods, including Saltasaurus itself. Thus it is possible that the largest sizes were attained among the more basal members of the clade[6]

Palaeoecology

These fossils were discovered in the Candeleros Formation, the oldest formation within the Neuquén Group of Neuquén Province, Argentina. This formation dates to the Cenomanian stage of the Late Cretaceous period, about 99.6 to 97 million years ago.[3] For the most part, the Candeleros represents an ancient braided river system, and besides Andesaurus, also contains fossils of theropods like Buitreraptor and the enormous Giganotosaurus, as well as other non-related sauropods such as Limaysaurus.

References

  1. ↑ "Caudal Vertebrae," Tidwell, Carpenter, and Meyer (2001). Page 145.
  2. 1 2 Paul, G.S., 2010, The Princeton Field Guide to Dinosaurs, Princeton University Press p. 205
  3. 1 2 Holtz, Thomas R. Jr. (2012) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2011 Appendix.
  4. ↑ Salgado, et al. (1997). Wilson and Upchurch (2003).
  5. ↑ Upchurch, et al. (2004).
  6. ↑ Novas, et al. (2005).

Bibliography

Further reading

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