List of Hypericum species

Hypericum perforatum, the type species of the genus Hypericum[1]

The genus Hypericum contains the following 490 species, according to The Plant List,[2] divided by sections.[3] Hypericum species are found most densely in Europe and parts of the Middle East.[4] They are a rather unusual genus in the fact that Norman Robson created a genus-wide monograph in 1977, dividing the genus into distinct sections that are still used today despite the genus's size [5] A phylogenetic study was completed for Hypericum, showing that the genus Triadenum is likely a clade in Hypericum. It also showed that Thornea is sister to Hypericum and that 60% of the sections of Hypericum are monophyletic.[6]

Adenosepalum

Adenosepalum Spach contains shrubs, shrublets, and soft herbs. Its species grow to 2.5 meters tall and can be found in most of Europe and parts of Northern Africa.

Adenotrias

Adenotrias (Jaub. & Spach) R. Keller contains shrubs and shrublets. Its species grow up to 2 meters tall and are found in West Africa and Eastern Europe.

  • Hypericum aciferum (Greuter) N. Robson
  • Hypericum aegypticum L.
  • Hypericum russeggeri (Fenzl) R. Keller

Androsaemum

The species in Androsaemum (Duhamel) Godron grow up to 3 meters tall and are shrubs. They are found in the Middle East and around the Mediterranean.

Arthrophyllum

The species in Arthrophyllum Jaub. & Spach grow to be less than a meter tall. They are found in parts of the Middle East.

  • Hypericum cardiophyllum Boiss.
  • Hypericum nanum Poir.
  • Hypericum pamphylicum N. Robson & P.H.Davis
  • Hypericum rupestre Jaub. & Spach
  • Hypericum vacciniifolium Hayek & Siehe

Ascyreia

Ascyreia Choisy contains shrubs, shrublets, and rarely trees. Most species grow to be 4–5 m tall and can be found across Europe.

  • Hypericum acmosepalum N.Robson
  • Hypericum augustinii N.Robson
  • Hypericum beanii N.Robson
  • Hypericum bellum H.L.Li
  • Hypericum calycinum L.
  • Hypericum choisianum Wall. ex N.Robson
  • Hypericum cohaerens N.Robson
  • Hypericum cordifolium Choisy
  • Hypericum curvisepalum N.Robson
  • Hypericum gaitii Haines
  • Hypericum forrestii (Chitt.) N.Robson
  • Hypericum gracilipes Stapf ex C.E.C.Fisch.
  • Hypericum griffithii Hook.f. & Thomson ex Dyer
  • Hypericum henryi H.Lév. & Vaniot
  • Hypericum hookerianum Wight & Arn.
  • Hypericum kouytchense H.Lév.
  • Hypericum lacei N.Robson
  • Hypericum lagarocladum N.Robson
  • Hypericum lancasteri N.Robson
  • Hypericum leschenaultii Choisy
  • Hypericum lobbii N.Robson
  • Hypericum maclarenii N.Robson
  • Hypericum monogynum L.
  • Hypericum mysurense Wall. ex Wight & Arn.
  • Hypericum oblongifolium Choisy
  • Hypericum pachyphyllum Collett & Hemsl.
  • Hypericum patulum Thunb.
  • Hypericum podocarpoides N.Robson
  • Hypericum prattii Hemsl.
  • Hypericum pseudohenryi N.Robson
  • Hypericum reptans Hook.f. & Thomson ex Dyer
  • Hypericum sherriffii N.Robson & D.G.Long
  • Hypericum siamense N.Robson
  • Hypericum stellatum N.Robson
  • Hypericum subsessile N.Robson
  • Hypericum tenuicaule Hook.f. & Thomson ex Dyer
  • Hypericum uralum Buch.-Ham. ex D.Don
  • Hypericum wardianum N. Robson
  • Hypericum williamsii N.Robson
  • Hypericum wilsonii N.Robson

Brathys

Brathys (Mutis ex L.f.) Choisy is one of the largest sections of Hypericum. It contains mainly evergreen trees, shrubs, shrublets, and wiry herbs that grow up to 6 m tall.

  • Hypericum acostanum Steyerm. ex N.Robson
  • Hypericum addingtonii N.Robson
  • Hypericum andinum Gleason
  • Hypericum arbuscula Standl. & Steyerm.
  • Hypericum asplundii N. Robson
  • Hypericum baccharoides Cuatrec.
  • Hypericum beamanii N. Robson
  • Hypericum bolivaricum N. Robson
  • Hypericum brathys Sm.
  • Hypericum bryoides Gleason
  • Hypericum callacallanum N. Robson
  • Hypericum caracasanum Willd.
  • Hypericum cardonae Cuatrec.
  • Hypericum carinosum R.Keller
  • Hypericum cassiopiforme N. Robson
  • Hypericum castellanoi N.Robson
  • Hypericum chamaemyrtus Triana & Planch.
  • Hypericum costaricense N.Robson
  • Hypericum cuatrecasii Gleason
  • Hypericum cymobrathys N. Robson
  • Hypericum decandrum Turcz.
  • Hypericum diosmoides Griseb.
  • Hypericum drummondii (Grev. & Hook.) Torr. & A. Gray
  • Hypericum eastwoodianum I.M.Johnst.
  • Hypericum elatoides R.Keller
  • Hypericum espinalii N. Robson
  • Hypericum fuertesii Urb.
  • Hypericum galinum S.F. Blake
  • Hypericum garciae Pierce
  • Hypericum gentianoides (L.) Britton, Sterns & Poggenb.
  • Hypericum gladiatum N. Robson
  • Hypericum gleasonii N. Robson
  • Hypericum gnidioides Seem.
  • Hypericum goyanesii Cuatrec.
  • Hypericum harlingii N. Robson
  • Hypericum hartwegii Benth.
  • Hypericum horizontale N. Robson
  • Hypericum humboldtianum Steud.
  • Hypericum irazuense Kuntze ex N.Robson
  • Hypericum jaramilloi N.Robson
  • Hypericum juniperinum Kunth
  • Hypericum lancifolium Gleason
  • Hypericum lancioides Cuatrec.
  • Hypericum laricifolium Juss.
  • Hypericum laricoides Gleason
  • Hypericum llanganaticum N. Robson
  • Hypericum loxense Benth.
  • Hypericum lycopodioides Triana & Planch.
  • Hypericum magdalenicum N.Robson
  • Hypericum magniflorum Cuatrec.
  • Hypericum maguirei N. Robson
  • Hypericum marahuacanum N.Robson
  • Hypericum martense N. Robson
  • Hypericum matangense N. Robson
  • Hypericum mexicanum L.
  • Hypericum millefolium Urb. & Ekman
  • Hypericum myricariifolium Hieron.
  • Hypericum papillosum N.Robson
  • Hypericum parallelum N. Robson
  • Hypericum paramitanum N.Robson
  • Hypericum peninsulare Eastw.
  • Hypericum phellos Gleason
  • Hypericum pimeleoides Planch. & Linden ex Triana & Planch.
  • Hypericum piriai Arechav.
  • Hypericum prietoi N. Robson
  • Hypericum prostratum Cuatrec.
  • Hypericum pseudobrathys Turcz.
  • Hypericum pycnophyllum Urb.
  • Hypericum quitense R.Keller
  • Hypericum radicans N. Robson
  • Hypericum recurvum N. Robson
  • Hypericum roraimense Gleason
  • Hypericum rubritinctum N. Robson
  • Hypericum ruscoides Cuatrec.
  • Hypericum sabiniforme Trevis.
  • Hypericum selaginella N. Robson
  • Hypericum simonsii N.Robson
  • Hypericum sprucei N. Robson
  • Hypericum stenopetalum Turcz.
  • Hypericum strictum Kunth
  • Hypericum struthiolifolium Juss.
  • Hypericum stuebelii Hieron.
  • Hypericum styphelioides A.Rich.
  • Hypericum terrae-firmae Sprague & L.Riley
  • Hypericum tetrastichum Cuatrec.
  • Hypericum thuyoides Kunth
  • Hypericum valleanum N. Robson
  • Hypericum woodianum N. Robson
  • Hypericum wurdackii N. Robson

Bupleuroides

Bupleuroides Stef. contains only a single species that is a small perennial herb.

Campylopus

Campylopus Boiss. contains only Hypericum cerastioides.

  • Hypericum cerastioides (Spach) N. Robson

Camplyosporus

Campylosporus (Spach) R. Keller

Concinna

Coridium

  • Hypericum asperuloides Czern. ex Turcz.
  • Hypericum coris L.
  • Hypericum empetrifolium Willd.
  • Hypericum ericoides L.
  • Hypericum jovis Greuter

Crossophyllum

Drosocarpium

Drosocarpium Spach contains small perennial herbs that are found near the Mediterranean.

  • Hypericum alpigenum Kit.
  • Hypericum ambiguum Elliott
  • Hypericum barbatum Jacq.
  • Hypericum bithynicum Boiss.
  • Hypericum confusum Rose
  • Hypericum montbretii Spach
  • Hypericum perfoliatum L.
  • Hypericum × reinosae A.Ramos
  • Hypericum richeri Vill.
  • Hypericum rumeliacum Boiss.
  • Hypericum spruneri Boiss.
  • Hypericum trichocaulon Boiss. & Heldr.
  • Hypericum umbellatum A.Kern.
  • Hypericum vesiculosum Griseb.

Elodeoida

Elodeoida N. Robson contains 1 m tall perennial herbs.

  • Hypericum austroyunnanicum L.H. Wu & D.P. Yang
  • Hypericum elodeoides Choisy
  • Hypericum hubeiense L.H. Wu & D.P. Yang
  • Hypericum kingdonii N.Robson
  • Hypericum petiolulatum Hook.f. & Thomson ex Dyer
  • Hypericum seniawinii Maxim.

Graveolentia

Graveolentia N. Robson is a diverse section whose type species is H. graveolens.

  • Hypericum collinum Schltdl. & Cham.
  • Hypericum epigeium R.Keller
  • Hypericum formosum Kunth
  • Hypericum graveolens Buckley
  • Hypericum macvaughii N.Robson
  • Hypericum × mitchellianum Rydb.
  • Hypericum oaxacanum R. Keller
  • Hypericum pringlei S.Watson
  • Hypericum pseudomaculatum Bush
  • Hypericum punctatum Lam.

Heterophylla

Heterophylla N. Robson contains a single shrublets measuring up to 25 cm tall.

  • Hypericum heterophyllum

Hirtella

Hirtella Stef. contains perennial herbs averaging 80 cm in height.

  • Hypericum amblysepalum Hochst.
  • Hypericum apiculatum (N.Robson) Sennikov
  • Hypericum apricum Kar. & Kir.
  • Hypericum asperulum Jaub. & Spach
  • Hypericum callithyrsum Coss.
  • Hypericum davisii N.Robson
  • Hypericum elongatum Ledeb. ex Rchb.
  • Hypericum hedgei N.Robson
  • Hypericum helianthemoides (Spach) Boiss.
  • Hypericum hirtellum (Spach) Boiss.
  • Hypericum karjaginii Rzazade
  • Hypericum libanoticum N.Robson
  • Hypericum lysimachioides Boiss. & Noë
  • Hypericum microcalycinum Boiss. & Heldr.
  • Hypericum olivieri (Spach) Boiss.
  • Hypericum pseudolaeve N.Robson
  • Hypericum retusum Aucher ex Jaub. & Spach
  • Hypericum salsolifolium Hand.-Mazz.
  • Hypericum scabroides N.Robson & Poulter
  • Hypericum scabrum L.
  • Hypericum sorgerae N.Robson
  • Hypericum spectabile Jaub. & Spach
  • Hypericum thymbrifolium Boiss. & Noë
  • Hypericum thymopsis Boiss.
  • Hypericum tymphresteum Boiss. & Spruner
  • Hypericum vermiculare Boiss. & Hausskn.

Humifusoideum

Humifusoideum R. Keller, also called Pulogensia, contains shrubs, subshrubs, and perennial herbs.

  • Hypericum bojerianum H. Perrier
  • Hypericum natalense J.M. Wood & M.S. Evans
  • Hypericum nokoense Ohwi
  • Hypericum peplidifolium A.Rich.
  • Hypericum wilmsii R. Keller

Hypericum

Hypericum, sometimes referred to as the "type section" of the genus, contains perennial herbs and very few suffrutices.

  • Hypericum asahinae Makino
  • Hypericum attenuatum Fisch. ex Choisy
  • Hypericum × desetangsii Lamotte
  • Hypericum elegans Stephan ex Willd.
  • Hypericum enshiense L.H. Wu & F.S. Wang
  • Hypericum erectum Thunb.
  • Hypericum faberi R.Keller
  • Hypericum furusei N.Robson
  • Hypericum gracillimum Koidz.
  • Hypericum hachijyoense Nakai
  • Hypericum hakonense Franch. & Sav.
  • Hypericum × hyugamontanum Kimura
  • Hypericum kamtschaticum Ledeb.
  • Hypericum kawaranum N.Robson
  • Hypericum kinashianum Koidz.
  • Hypericum kitamense (Y.Kimura) N.Robson
  • Hypericum kiusianum Koidz.
  • Hypericum kurodakeanum N.Robson
  • Hypericum × laschii A.Fröhl.
  • Hypericum maculatum Crantz
  • Hypericum × medium' Peterm.
  • Hypericum momoseanum Makino
  • Hypericum nakaii H.Koidz.
  • Hypericum nikkoense Makino
  • Hypericum nuporoense N.Robson
  • Hypericum oliganthum Franch. & Sav.
  • Hypericum ovalifolium Koidz.
  • Hypericum perforatum L.
  • Hypericum pibairense (Miyabe & Y.Kimura) N.Robson
  • Hypericum pseudoerectum N.Robson
  • Hypericum pseudopetiolatum R.Keller
  • Hypericum scouleri Hook.
  • Hypericum senanense Maxim.
  • Hypericum sikokumontanum Makino
  • Hypericum taihezanense Sasaki ex S.Suzuki
  • Hypericum tetrapterum Fr.
  • Hypericum triquetrifolium Turra
  • Hypericum undulatum Schousb. ex Willd.
  • Hypericum uniglandulosum Hausskn. ex Bornm.
  • Hypericum vulcanicum Koidz.
  • Hypericum watanabei N.Robson
  • Hypericum yamamotoanum H.Koidz.
  • Hypericum yamamotoi Miyabe & Kimura
  • Hypericum yezoense Maxim.

Inodora

Inodora Stef. contains one species, a shrub that grows from 1 to 5 meters tall.

Monanthema

Monanthema N. Robson contains perennial herbs that grow up to 40 cm tall.

  • Hypericum daliense N.Robson
  • Hypericum himalaicum N.Robson
  • Hypericum ludlowii N.Robson
  • Hypericum monanthemum Hook.f. & Thomson ex Dyer
  • Hypericum subcordatum (R.Keller) N.Robson
  • Hypericum trigonum Hand.-Mazz.
  • Hypericum wightianum Wall. ex Wight & Arn.

Myrianda

Myriandra (Spach) R. Keller contains shrubs, shrublets, and perennial herbs that grow to be up to 4.5 m.

  • Hypericum adpressum W.P.C.Barton
  • Hypericum apocynifolium Small
  • Hypericum brachyphyllum (Spach) Steud.
  • Hypericum buckleyi M.A. Curtis
  • Hypericum chamaenerium (Spach) Steud.
  • Hypericum chapmanii W.P. Adams
  • Hypericum cistifolium Lam.
  • Hypericum crux-andreae (L.) Crantz
  • Hypericum densiflorum Pursh
  • Hypericum edisonianum (Small) W.P. Adams & N. Robson
  • Hypericum ellipticum Hook.
  • Hypericum exile W.P. Adams
  • Hypericum fasciculatum Lam.
  • Hypericum frondosum Michx.
  • Hypericum galioides Lam.
  • Hypericum glomeratum Small
  • Hypericum hypericoides (L.) Crantz
  • Hypericum kalmianum L.
  • Hypericum lissophloeus W.P. Adams
  • Hypericum lloydii (Svenson) P.B.Adams
  • Hypericum lobocarpum Gatt.
  • Hypericum longistylum Oliv.
  • Hypericum lydium Boiss.
  • Hypericum myrtifolium Lam.
  • Hypericum nitidum Lam.
  • Hypericum nudiflorum Michx. ex Willd.
  • Hypericum opacum Torr. & A. Gray
  • Hypericum prolificum L.
  • Hypericum suffruticosum W.P. Adams & N. Robson
  • Hypericum tetrapetalum Lam.

Oligostema

Oligostema (Boiss.) Stef. consista of perennial and annual herbs up to 75 cm tall.

  • Hypericum andjerinum Font Quer & Pau
  • Hypericum australe Ten.
  • Hypericum × caesariense Druce ex N.Robson
  • Hypericum humifusum L.
  • Hypericum kelleri Bald.
  • Hypericum linariifolium Vahl
  • Hypericum sphaerocarpum Michx.

Olympia

Olympia (Spach) Nymam contains four dwarf shrubs. They are found in Southeastern Europe and Turkey.

  • Hypericum auriculatum (N.Robson & Hub.-Mor.) N.Robson
  • Hypericum lycium (N.Robson & Hub.-Mor.) N.Robson
  • Hypericum olympicum L.
  • Hypericum polyphyllum Boiss. & Balansa

Origanifolia

Origanifolia Spach contains unique shrub-like perennial herbs.

  • Hypericum albiflorum (Hub.-Mor.) N.Robson
  • Hypericum aviculariifolium Jaub. & Spach
  • Hypericum bourgaei (Boiss.) N.Robson
  • Hypericum cymbiferum Boiss. & Balansa
  • Hypericum ichelense N.Robson
  • Hypericum imbricatum Poulter
  • Hypericum laxiflorum N.Robson
  • Hypericum leprosum Boiss.
  • Hypericum origanifolium Willd.
  • Hypericum papillare Boiss. & Heldr.
  • Hypericum salsugineum N.Robson & Hub.-Mor.
  • Hypericum trachyphyllum Griseb.
  • Hypericum uniflorum Boiss. & Heldr.

Psorophytum

Psorophytum (Spach) Nyman contains a single evergreen shrub. It grows from 1–5 meters tall.

Roscyna

Roscyna (Spach) R. Keller contains perennial herbs that grow up to 2 meters tall.

Sampsonia

Sampsonia N. Robson contains mainly perennial herbs that grow to be 80 cm.

  • Hypericum assamicum S.N.Biswas
  • Hypericum sampsonii Hance

Santomasia

Santomasia (N. Robson) N. Robson contains a single species.

  • Hypericum steyermarkii Standl.

Taeniocarpium

Taeniocarpium Jaub. & Spach contains small wiry perennial herbs up to 1.1 meters tall.

  • Hypericum armenum Jaub. & Spach
  • Hypericum crenulatum Boiss.
  • Hypericum fissurale Woronow
  • Hypericum havvae Güner
  • Hypericum hirsutum L.
  • Hypericum kotschyanum Boiss.
  • Hypericum linarioides Bosse
  • Hypericum malatyanum Peșmen
  • Hypericum marginatum Woronow
  • Hypericum neurocalycinum Boiss. & Heldr.
  • Hypericum nummularioides Trautv.
  • Hypericum nummularium L.
  • Hypericum peshmenii Yıld.
  • Hypericum pruinatum Boiss. & Balansa
  • Hypericum pseudorepens N.Robson
  • Hypericum pulchrum L.
  • Hypericum pumilio Bornm.
  • Hypericum saxifragum N.Robson & Hub.-Mor.
  • Hypericum taygeteum Quézel & Contandr.
  • Hypericum theodori Woronow
  • Hypericum thymifolium Banks & Sol.
  • Hypericum vaccinioides N.Robson
  • Hypericum venustum Fenzl

Takasagoya

Takasagoya (Y.Kimura) N. Robson contains deciduous shrubs and shrublets that grow up to 1.5 m tall.

  • Hypericum formosanum Maxim.
  • Hypericum geminiflorum Hemsl.
  • Hypericum nakamurai (Masam.) N.Robson
  • Hypericum senkakuinsulare Hatus.
  • Hypericum subalatum Hayata

Triadenoides

Triadenoides Jaub. & Spach contains shrubs and shrublets that can grow to be 0.8 m tall.

Trigynobrathys

Trigynobrathys (Y. Kimura) N. Robson contains shrubs and subshrubs as well as annual and perennial herbs that are very diverse in size and shape.

  • Hypericum anagalloides Cham. & Schltdl.
  • Hypericum aphyllum Lundell
  • Hypericum arenarioides A. Rich.
  • Hypericum boreale (Britton) H. P. Bicknell
  • Hypericum brasiliense Choisy
  • Hypericum brevistylum Choisy
  • Hypericum caespitosum Cham. & Schltdl.
  • Hypericum campestre Cham. & Schltdl.
  • Hypericum canadense L.
  • Hypericum caprifoliatum Cham. & Schltdl.
  • Hypericum carinatum Griseb.
  • Hypericum cavernicola L.B. Sm.
  • Hypericum connatum Lam.
  • Hypericum cordatum (Vell.) N. Robson
  • Hypericum cumulicola (Small) P.B. Adams
  • Hypericum denticulatum Walter
  • Hypericum denudatum A. St.-Hil.
  • Hypericum × dissimulatum E.P. Bicknell
  • Hypericum dolabriforme Vent.
  • Hypericum erythreae (Spach) Steud.
  • Hypericum globuliferum R. Keller
  • Hypericum gramineum G.Forst.
  • Hypericum gymnanthum Engelm. & A.Gray
  • Hypericum harperi R.Keller
  • Hypericum humbertii Staner
  • Hypericum japonicum Thunb.
  • Hypericum killipii N. Robson
  • Hypericum lalandii Choisy
  • Hypericum legrandii L.B. Sm.
  • Hypericum linoides A. St.-Hil.
  • Hypericum lorentzianum Gilg ex R. Keller
  • Hypericum majus (A.Gray) Britton
  • Hypericum microlicioides L.B. Sm.
  • Hypericum microsepalum (Torr. & A. Gray) A. Gray ex S. Watson
  • Hypericum monadenum N.Robson
  • Hypericum moranense Kunth
  • Hypericum mutilum L.
  • Hypericum myrianthum Cham. & Schltdl.
  • Hypericum oligandrum Milne-Redh.
  • Hypericum parvulum Greene
  • Hypericum pauciflorum Kunth
  • Hypericum paucifolium S. Watson
  • Hypericum pedersenii N.Robson
  • Hypericum philonotis Schltdl. & Cham.
  • Hypericum pleiostylum C. Rodr. Jim.
  • Hypericum polyanthemum Klotzsch ex Reichardt
  • Hypericum pratense Schltdl. & Cham.
  • Hypericum pumilum Sessé & Moc.
  • Hypericum relictum N. Robson
  • Hypericum rigidum A. St.-Hil.
  • Hypericum salvadorense N. Robson
  • Hypericum scioanum Chiov.
  • Hypericum setosum L.
  • Hypericum silenoides Juss.
  • Hypericum teretiusculum A. St.-Hil.
  • Hypericum ternum A. St.-Hil.
  • Hypericum thesiifolium Kunth
  • Hypericum virgatum Lam.

Tripentas

Tripentas (Casp.) N. Robson contains long-stemmed perennial herbs.

Umbraculoides

Umbraculoides N. Robson contains a single species by its own name.

Webbia

Webbia (Spach) R. Keller contains a large deciduous shrub that can grow up to 4 meters tall.

Unspecified

  • Hypericum callinum Rose
  • Hypericum × inodorum Mill.
  • Hypericum iwate-littorale H.Koidz.
  • Hypericum nagasawae Hayata
  • Hypericum rudiflorum Michx.
  • Hypericum tubulosum Walter

Bibliography

References

  1. Le Roy Abrams & Roxana Stinchfield Ferris (1923). An Illustrated Flora of the Pacific States: Geraniaceae to Scrophulariaceae, geraniums to figworts. An Illustrated Flora of the Pacific States: Washington, Oregon, and California. 3. Stanford University Press. p. 116. ISBN 9780804700054.
  2. "Hypericum". The Plant List. Retrieved 19 February 2016.
  3. Hypericum MySpecies Retrieved March 4, 2016
  4. CABI.org Retrieved 9/7/16
  5. Robson, N.K.B. (1977) Studies in the genus Hypericum L. (Guttiferae) 1. Infrageneric classification Bulletin of the British Museum (Natural History), Botany 5: 293–355.
  6. Molecular phylogenetics and morphological evolution of St. John's wort (Hypericum; Hypericaceae) by Nürk NM, Madriñán S, Carine MA, Chase MW, Blattner FR
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