Plagiaulacida

Plagiaulacidans
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Multituberculata
Suborder:	†Plagiaulacida
Families
Albionbaataridae Allodontidae Eobaataridae Hahnodontidae Paulchoffatiidae Pinheirodontidae Plagiaulacidae Zofiabaataridae

Plagiaulacida is a group of extinct multituberculate mammals. Multituberculates were among the most common mammals of the Mesozoic, "the age of the dinosaurs". Plagiaulacids, an informal suborder, are the most basal of this order, and ranged from the Middle Jurassic Period to the Lower Cretaceous Period of the northern hemisphere.

Kielan-Jaworowska and Hurum (2001) divides “Plagiaulacida” into three informal lineages, the paulchoffatiids, the plagiaulicids, and the allodontids.

Allodontid line

The Allodontid line may be a superfamily, Allodontoidea.

Both allodontids and paulchoffatiids (below) were among the most basal of the plagiaulacids. The Allodontid line contains:

The family Allodontidae is known from two genera from the Upper Jurassic Morrison Formation of North America.

The family Zofiabaataridae contains a single genus, Zofiabaatar and is also from the Morrison Formation. The affinities of a further Morrison Formation genus, Glirodon, are unclear, but it’s also within the Allodontid line.

Paulchoffatiid line

The Paulchoffatiid line may be a superfamily, Paulchoffatioidea.

Some remains from the Middle Jurassic of England might belong within this group. Representatives are best known from the Upper Jurassic, (especially from Guimarota, Portugal), though some were still extant during the Lower Cretaceous.

The genera of the family Paulchoffatiidae are divided into two of subfamilies, plus a couple of harder-to-place individuals:

• Subfamily Paulchoffatiinae includes Paulchoffatia and its relatives. This taxon contains nine genera.
• Subfamily Kuehneodontinae consists solely of the genus Kuehneodon, though there are half-a-dozen named species.

Other genera include Galveodon and Sunnyodon, both based on teeth from the Lower Cretaceous of Spain and England respectively.

Also referable to the paulchoffatiid line, but not the family itself, are the following:

• Family Hahnodontidae, which is restricted to a single lower tooth from the Upper Cretaceous of Morocco, (Hahnodon).
• Family Pinheirodontidae is known from Lower Cretaceous teeth found in Iberia and England.

Plagiaulacid line (possibly Superfamily Plagiaulacoidea?)

Family Plagiaulacidae is known from the Upper Jurassic (North America) to Lower Cretaceous (Europe and Asia).

Plagiaulax, Bolodon, 'Ctenacodon' brentbaatar. 'Ctenacodon requires renaming, while as of 2001 material from China has yet to be described.

Family Albionbaataridae is known from the Upper Jurassic to Lower Cretaceous of Europe and Asia, (China – undescribed, 2001). These were shrew-sized Multituberculates, with some similarities to the paulchoffis.

Members of the family Eobaataridae display dental similarities with some of the Paracimexomys group, (Cimolodonta). They are known from the Upper Jurassic to Lower Cretaceous of Europe and Asia, (China – undescribed, 2001).

Sinobaatar was described after the study by Kielan-Jaworowska and Hurum (2001). It might be based on some of the Chinese material they mentioned as undescribed. The Mongolian word ‘baatar’ is frequently employed in the nomenclature of Multituberculates. This reflects the fact that many of the most complete fossils have been recovered from sites in Mongolia, though this more applies to members of the more derived Cimolodonta.

A couple of further genera possibly fit somewhere within “Plagiaulacida”. This has been tentatively proposed for Janumys of the Middle Cretaceous. Its contemporary, Ameribaatar, is of uncertain affinities. Both were first described late in 2001.

Taxonomy

Subclass †Allotheria Marsh, 1880

• Order †Multituberculata Cope, 1884:
  • suborder †Plagiaulacida Simpson 1925
    • Family †Paulchoffatiidae Hahn, 1969
      • subfamily †Paulchoffatiinae Hahn, 1971
        • Genus †Paulchoffatia Kühne, 1961
          • Species †P. delgador Kühne, 1961
        • Genus †Pseudobolodon Hahn, 1977
          • Species †P. oreas Hahn, 1977
          • Species †P. krebsi Hahn & Hahn, 1994
        • Genus †Henkelodon Hahn, 1987
          • Species †H. naias Hahn, 1987
        • Genus †Guimarotodon Hahn, 1969
          • Species †G. leiriensis Hahn, 1969
        • Genus †Meketibolodon (Hahn, 1978) Hahn, 1993
          • Species †M. robustus (Hahn, 1978) Hahn, 1993
        • Genus †Plesiochoffatia Hahn & Hahn, 1999
          • Species †P. thoas Hahn & Hahn, 1998
          • Species †P. peparethos Hahn & Hahn, 1998
          • Species †P. staphylos Hahn & Hahn, 1998
        • Genus †Xenachoffatia Hahn & Hahn, 1998
          • Species †X. oinopion Hahn & Hahn, 1998
        • Genus †Bathmochoffatia Hahn & Hahn, 1998
          • Species †B. hapax Hahn & Hahn, 1998
        • Genus †Kielanodon Hahn, 1987
          • Species †K. hopsoni Hahn, 1987
        • Genus †Meketichoffatia Hahn, 1993
          • Species †M. krausei Hahn, 1993
        • Genus †Galveodon Hahn & Hahn, 1992
          • Species †G. nannothus Hahn & Hahn, 1992
        • Genus †Sunnyodon Kielan-Jaworowska & Ensom, 1992
          • Species †S. notleyi Kielan-Jaworowska & Ensom, 1992
      • Subfamily †Kuehneodontinae Hahn, 1971
        • Genus †Kuehneodon Hahn, 1969
          • Species †K. dietrichi Hahn, 1969
          • Species †K. barcasensis Hahn & Hahn, 2001
          • Species †K. dryas Hahn, 1977
          • Species †K. guimarotensis Hahn, 1969
          • Species †K. hahni Antunes, 1988
          • Species †K. simpsoni Hahn, 1969
          • Species †K. uniradiculatus Hahn, 1978
      • Family †Hahnodontidae Sigogneau-Russell, 1991
        • Genus †Hahnodon Sigogneau-Russell, 1991
          • Species †H. taqueti Sigogneau-Russell, 1991
        • Genus †Denisodon Hahn & Hahn,2003
          • Species †D. moroccensis Hahn & Hahn,2003
      • Family †Pinheirodontidae Hahn & Hahn, 1999
        • Genus †Pinheirodon Hahn & Hahn, 1999
          • Species †P. pygmaeus Hahn & Hahn, 1999
          • Species †P. vastus Hahn & Hahn, 1999
        • Genus †Bernardodon Hahn & Hahn, 1999
          • Species †B. sp. Hahn & Hahn, 1999
        • Genus †Gerhardodon Kielan-Jaworowska & Ensom, 1992
          • Species †G. purbeckensis Kielan-Jaworowska & Ensom, 1992
        • Genus †Iberodon Hahn & Hahn, 1999
          • Species †I. quadrituberculatus Hahn & Hahn, 1999
        • Genus †Lavocatia Canudo & Cuenca-Bescós, 1996
          • Species †L. alfambrensis Canudo & Cuenca-Bescós, 1996
        • Genus †Ecprepaulax Hahn & Hahn, 1999
          • Species †E. sp. Hahn & Hahn, 1999
      • Family †Allodontidae Marsh, 1889
        • Genus †Ctenacodon Marsh, 1879
          • Species †C. serratus Marsh, 1879
          • Species †C. nanus Marsh, 1881
          • Species †C. laticeps Marsh, 1881
          • Species †C. scindens Simpson, 1928
        • Genus †Psalodon Simpson, 1926
          • Species †P. potens Marsh, 1887
          • Species †P. fortis Marsh, 1887
          • Species †P. marshi Simpson, 1929
      • Family †Zofiabaataridae Bakker, 1992
        • Genus †Zofiabaatar Bakker & Carpenter, 1990
          • Species †Z. pulcher Bakker & Carpenter, 1990
      • Family Incertae sedis
        • Genus †Glirodon Engelmann & Callison, 2001
        • Species †G. grandis Engelmann & Callison, 2001
      • Family †Plagiaulacidae Gill, 1872
        • Genus? †Ctenacodon Bakker, 1998
          • Species? †C. brentbaatar Bakker, 1998
        • Genus †Plagiaulax Falconer, 1857
          • Species †P. becklesii Falconer, 1857
        • Genus †Bolodon Owen, 1871
          • Species †B. crassidens Owen, 1871
          • Species †B. falconeri Owen, 1871
          • Species †B. minor Falconer, 1857
          • Species †B. osborni Simpson, 1928
          • Species †B. elongatus Simpson, 1928
      • Family †Eobaataridae Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
        • Genus †Eobaatar
          • Species †E. magnus Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
          • Species †E. minor Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
          • Species †E. hispanicus Hahn & Hahn, 1992
          • Species †E. pajaronensis Hahn & Hahn, 2001
        • Genus †Loxaulax Simpson, 1928
          • Species †L. valdensis Simpson, 1928
        • Genus †Monobaatar Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
          • Species †M. mimicus Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
        • Genus †Parendotherium Crusafont Pairó & Adrover, 1966
          • Species †P. herreroi Crusafont Pairó & Adrover, 1966
        • Genus †Sinobaatar Hu & Wang, 2002
          • Species †S. lingyuanensis Hu & Wang, 2002
        • Genus †Heishanobaatar Kusuhashi et al., 2010
          • Species †H. triangulus Kusuhashi et al., 2010
      • Family †Albionbaataridae Kielan-Jaworowska & Ensom, 1994
        • Genus †Albionbaatar Kielan-Jaworowska & Ensom, 1994
          • Species †A. denisae Kielan-Jaworowska & Ensom, 1994
        • Genus †Proalbionbaatar Hahn & Hahn, 1998
          • Species †P. plagiocyrtus Hahn & Hahn, 1998
        • Genus †Kielanobaatar Kusuhashi et al., 2010
          • Species †K. badaohaoensis Kusuhashi et al., 2010
      • Family †Arginbaataridae Hahn & Hahn, 1983
        • Genus †Arginbaatar Trofimov, 1980
          • Species †A. dmitrievae Trofimov, 1980

References

• Hahn G & Hahn R (2000), Multituberculates from the Guimarota mine, p. 97-107 in
• Martin T & Krebs B (eds), Guimarota - A Jurassic Ecosystem, Published by Dr Friedrich Pfeil, Münich, Germany.
• Kielan-Jaworowska Z & Hurum JH (2001), Phylogeny and Systematics of multituberculate mammals. Paleontology 44, p. 389-429.
• Much of this information has been derived from Multituberculata Cope, 1884.