Fagus grandifolia

American beech
Scientific classification
Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
(unranked):	Rosids
Order:	Fagales
Family:	Fagaceae
Genus:	Fagus
Species:	F. grandifolia
Binomial name
Fagus grandifolia Ehrh.
Natural range of Fagus grandifolia

Fagus grandifolia (American beech or North American beech) is the species of beech tree native to the eastern United States and Canada. The genus name Fagus is Latin for "beech", and the species name grandifolia comes from grandis "large" and folium "leaf".

Distribution and varieties

The American beech is native to eastern North America, from Nova Scotia west to southern Ontario in southeastern Canada, west to Wisconsin and south to eastern Texas and northern Florida in the United States.

Trees in the southern half of the range are sometimes distinguished as a variety, F. grandifolia var. caroliniana, but this is not considered distinct in the Flora of North America. The Mexican beech (Fagus mexicana), native to the mountains of central Mexico, is closely related, and is sometimes treated as a subspecies of American beech, but more often as a distinct species. The only Fagus species found in the Western Hemisphere (assuming F. mexicana is treated as a subspecies), F. grandifolia is believed to have spanned the width of the North American continent all the way to the Pacific coast before the last ice age.

Description

It is a deciduous tree growing to 20–35 m (66–115 ft) tall, with smooth, silver-gray bark. The leaves are dark green, simple and sparsely-toothed with small teeth that terminate each vein, 6–12 cm (2.4–4.7 in) long (rarely 15 centimetres (5.9 in)), with a short petiole. The winter twigs are distinctive among North American trees, being long and slender (15–20 mm (0.59–0.79 in) by 2–3 mm (0.079–0.118 in)) with two rows of overlapping scales on the buds. Beech buds are distinctly thin and long, resembling cigars; this characteristic makes beech trees relatively easy to identify. The tree is monoecious, with flowers of both sexes on the same tree. The fruit is a small, sharply-angled nut, borne in pairs in a soft-spined, four-lobed husk. It has two ways of reproducing: one is through the usual dispersal of seedlings, and the other is through root sprouts. This is where the tree will have smaller trees growing out of its roots in different locations.^[1]

The American beech is a shade-tolerant species, favoring shade more than other trees, commonly found in forests in the final stage of succession. Ecological succession is essentially the process of forests changing their composition through time; it is a pattern of events often observed on disturbed sites.^[2] Although sometimes found in pure stands, it is more often associated with sugar maple (forming the beech-maple climax community), yellow birch, and Eastern hemlock, typically on moist well drained slopes and rich bottomlands. Near its southern limit, it often shares canopy dominance with Southern Magnolia. Although it has a reputation for slow growth (sometimes only 13 feet in 20 years), rich soil and ample moisture will greatly speed the process up. Much like sugar maple, American beech is picky about growing conditions, favoring a well-watered, but also well-drained spot and is intolerant of urban pollution, salt, and soil compaction. It also casts dense shade and has an extensive network of shallow, fibrous surface roots that make gardening underneath it almost impossible. Because American beech needs plenty of moisture and rich soil to thrive, it naturally occurs in bottomland forests. Early settlers often looked for beeches as a sign of a good potential place to clear the forest for farming.

Gallery

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  The long and thin winter bud of a beech

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  Spring leaves

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  Beech leaf and nut

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  In the winter, some fall leaves stay on beech trees (marcescence)

Disease

Beech bark disease has become a major killer of beech trees in the Northeastern United States. This disease occurs when the beech scale insect, Cryptococcus fagisuga, attacks the bark, creating a wound that is then infected by one of two different species of fungi in the genus Nectria. This causes a canker to develop and the tree is eventually killed.

Beech blight aphids colonize branches of the tree, but without serious harm to otherwise healthy trees. Below these colonies, deposits of sooty mold develop caused by the fungus Scorias spongiosa growing saprophytically on the honeydew the insects exude. This is also harmless to the trees.^[3]

Uses

Autumn foliage

American beech is an important tree in forestry. The wood is heavy, hard, tough and strong, and until the advent of power tools in the 20th century, lumbering beech trees were often left uncut to grow. As a result, many areas today still have extensive groves of old beeches that would not otherwise occur. Today, the wood is harvested for uses such as flooring, containers, furniture, handles and woodenware.

Like European beech bark, the American beech bark is smooth and uniform, making it an attraction for vandals to carve names, dates, gang symbols, and other material into its surface.^[4] One such beech tree in Louisville, Kentucky, in what is now the southern part of Iroquois Park, bore the legend "D. Boone kilt a bar" and the year in the late 18th century. This carving was first reported in the 19th century, but is believed to have not been produced by Boone since he was an educated man who would not have written something so semi-literate. The beech finally fell over in 1916 during a storm; its age was estimated at around 325 years. Its trunk is now on display at The Filson Historical Society.

It is sometimes planted as an ornamental tree, but even within its native area, it is planted much less often than the European beech. Although American beech can handle hotter climates,^[5] its European variant is faster-growing and more pollution-tolerant, in addition to being easier to propagate.

American beech can take up to 40 years to begin producing seeds. Large crops are produced by 60 years and the tree's total lifespan may be up to 300 years. The fruit is a triangle-shaped shell containing 2-3 nuts inside, but many of them do not fill in, especially on solitary trees. Beech nuts are edible to humans, although too small to be commercially valuable.

The mast (crop of nuts) from American beech provides food for numerous species of animals. Among vertebrates alone, these include ruffed grouse, wild turkeys, raccoons, red/gray foxes, white-tailed deer, rabbits, squirrels, opossums, pheasants, black bears, porcupines, and humans. For lepidopteran caterpillars feeding on American beech, see list of Lepidoptera that feed on beeches. Beech nuts were one of the primary foods of the now-extinct passenger pigeon; the clearing of beech and oak forests are pointed to as one of the major factors that may have contributed to the bird's extinction.^[6]

References

• R.C. Hosie, 1969. Native Trees of Canada. Canadian Forestry Service, Ottawa.

External links

Wikimedia Commons has media related to Fagus grandifolia.

• USDA Plants Profile for Fagus grandifolia (American beech)
• efloras.org: Flora of North America — Fagus grandifolia
• efloras.org: range map
• Interactive distribution map for Fagus grandifolia
• Bioimages.vanderbilt.edu — Fagus grandifolia photo gallery