Glyptodontidae

Glyptodonts Temporal range: Early Miocene–Pleistocene PreЄ Є O S D C P T J K Pg N
Fossil Glyptodon in the Naturhistorisches Museum, Vienna
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Superorder:	Xenarthra
Order:	Cingulata
Family:	†Glyptodontidae Burmeister 1879
Genera
†Pseudoneothoracophorus †Neothoracophorus †Glyptatelus †Clypeotherium †Neoglyptatelus †Pachyarmatherium †Hoplophorus †Palaehoplophorus †Propalaehoplophorus †Parapropalaehoplophorus †Metopotoxus †Eucinepeltus †Asterostemma †Eleutherocercus †Prodaedicurus †Lomaphorus †Castellanosia †Xiphuroides †Doedicurus †Daedicuroides †Plaxhaplous †Glyptotherium †Glyptodontidium †Paraglyptodon †Glyptodon †Stromatherium †Chlamydotherium †Glyptostracon †Heteroglyptodon †Panochthus †Sclerocalyptus †Neosclerocalyptus †Neuryurus

Glyptodonts are an extinct family of large, heavily armored cingulate xenarthrans related to the extinct pampatheres and modern armadillos.

Evolution

Doedicurus and Glyptodon by Robert Bruce Horsfall

Glyptodonts first evolved during the Miocene in South America, which remained their center of species diversity. For example, an Early Miocene glyptodont with many primitive features, when compared to other species, Parapropalaehoplophorus septentrionalis, was discovered at a now-elevated site in Chile and described in 2007.^[1] When the Isthmus of Panama formed about three million years ago, several species, such as Glyptotherium texanum, spread north as part of the Great American Interchange, as did pampatheres and armadillos.

Description

Glyptodonts possessed a tortoise-like body armour, made of bony deposits in their skin called osteoderms or scutes. Each species of glyptodont had a unique osteoderm pattern and shell type. With this protection, they were armored like turtles; glyptodonts could not withdraw their heads, but their armoured skin provided a bony cap on the top of their skull. Even the tail of glyptodonts had a ring of bones for protection. Doedicurus possessed a large mace-like spiked tail that it would have used to defend itself from predators and, possibly, other Doedicurus. Glyptodonts also had size on their side; many such as the type genus, Glyptodon, were the size of modern automobiles. The presence of such heavy defenses suggests the presence of a large, effective predator. At the time the glyptodontids evolved, the apex predators in the island continent of South America were phorusrhacids, a family of giant flightless carnivorous birds.

In physical appearance glyptodonts superficially resembled the much earlier dinosaurian ankylosaurs and, to a lesser degree, the recently extinct giant meiolaniid turtles of Australia, examples of the convergent evolution of unrelated lineages into similar forms.

Behaviour

Detail of Propalaehoplophorus scutes, early Miocene, in the permanent collection of The Children’s Museum of Indianapolis

The glyptodonts were grazing herbivores. Like many other xenarthrans, they had no incisor or canine teeth, but had a number of cheek teeth that would have been able to grind up tough vegetation, such as grasses. They also had distinctively deep jaws, with large downward bony projections that would have anchored powerful chewing muscles in life.^[2]

Extinction

Glyptodonts became extinct at the end of the last ice age along with a large number of other megafaunal species, including pampatheres, the giant ground sloths and the Macrauchenia. Their much smaller, more lightly armored and flexible relatives, the armadillos, survived. Extinction of the glyptodonts coincided with the arrival of early humans in the Americas and evidence suggests that these humans made use of the animal's armored shells.^[3]

Notes

External links

Wikispecies has information related to: Glyptodontidae

• Glyptodont article at ScienceBlogs, with photos
•  Ernest Ingersoll (1920). "Glyptodont". Encyclopedia Americana.