New Zealand kaka

New Zealand kākā
A pair
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Psittaciformes
Superfamily: Strigopoidea
Family: Nestoridae
Genus: Nestor
Species: N. meridionalis
Binomial name
Nestor meridionalis
(Gmelin, 1788)
Range in green

The New Zealand kaka, also known as kākā, (Nestor meridionalis) is a large species of parrot of the superfamily Strigopoidea found in native forests of New Zealand. Two subspecies are recognised. It is endangered and has disappeared from much of its former range.

Taxonomy and naming

The New Zealand kaka was described by German naturalist Johann Friedrich Gmelin in 1788. There are two subspecies, the North Island kākā, Nestor meridionalis septentrionalis, and the South Island kākā, N. m. meridionalis. The name kākā is a Māori language word meaning "parrot", possibly related to kā, 'to screech'.[2]

The genus Nestor contains four species: the New Zealand kākā (Nestor meridionalis), the kea (N. notabilis), the extinct Norfolk kaka (N. productus), and the extinct Chatham kaka (N. sp.). All four are thought to stem from a "proto-kākā", dwelling in the forests of New Zealand five million years ago.[3][4] Their closest relative is the kakapo (Strigops habroptila).[3][4][5][6] Together, they form the parrot family Strigopidae, an ancient group that split off from all other Psittacidae before their radiation.[3][4][6][7]

Description

The New Zealand kaka, like many parrots, uses its feet to hold its food

The New Zealand kākā is a medium-sized parrot, measuring 45 cm (18 in) in length and weighing from 390 to 560 g (14 to 20 oz), with an average of 452 g (0.996 lb).[8] It is closely related to the kea, but has darker plumage and is more arboreal. The forehead and crown are greyish-white and the nape is greyish-brown. The neck and abdomen are more reddish, while the wings are more brownish. Both sub-species have a strongly patterned brown/green/grey plumage with orange and scarlet flashes under the wings; color variants which show red to yellow coloration especially on the breast are sometimes found.

This group of parrots is unusual, retaining more primitive features lost in most other parrots, because it split off from the rest around 100 million years ago.[9]

The calls include a harsh ka-aa and a whistling u-wiia.[10]

Distribution and habitat

The New Zealand kākā lives in lowland and mid-altitude native forest. Its strongholds are currently the offshore reserves of Kapiti Island, Codfish Island and Little Barrier Island. It is breeding rapidly in the mainland island sanctuary at Zealandia (Karori Wildlife Sanctuary), with over 600 birds banded since their reintroduction in 2002.[11] From their reintroduction in 2002 the North Island Kākā continue to re-colonise Wellington and a recent report shows a significant increase in their numbers over the last 12 years.[12]

Behaviour

Kākā are mainly arboreal and occupy mid-to-high canopy. Often seen flying across valleys or calling from the top of emergent trees. They are very gregarious and move in large flocks often containing kea where present.

Diet

North Island kākā in flight, showing red plumage on underside of wing. At Pukaha Mt Bruce National Wildlife Centre.
A North Island kaka at Karori Wildlife Sanctuary, Wellington, New Zealand

The New Zealand kākā eats fruits, berries, seeds, flowers, buds, nectar, sap, plants and invertebrates. It uses its strong beak to shred the cones of the kauri tree to obtain the seeds.[13] It has a brush tongue with which it feeds on nectar, and it uses its strong beak to dig out the grubs of the huhu beetle and to remove bark to feed on sap.[14]

Nesting

New Zealand kākā make their nests in hollow trees, laying clutches of 2 to 4 eggs in late winter. Both parents assist in feeding the chicks. In a good fruiting year pairs can double clutch often utilizing the same nest hole for the second clutch. It is unusual for a pair to raise more than three chicks in a clutch.

Conservation status

The New Zealand kākā is considered vulnerable (CITES II). It has greatly declined across its traditional range as a result of habitat loss; predation by introduced predators like rats, possums and stoats; and competition from wasps and bees for the honeydew excreted by scale insects. A closely related species, Nestor productus, the Norfolk kaka, became extinct in 1851 for similar reasons.

Predation

Predatory mammals are responsible for the loss of an estimated 26 million native birds and their eggs each year in New Zealand.[15]

As cavity nesters with a long incubation period that requires the mother to stay on the nest for at least 90 days, kākā are particularly vulnerable to predation. Stoats were the main cause of death of nesting adult females, nestlings and fledglings, but possums were also important predators of adult females, eggs and nestlings.[16] There is strong evidence that predation of chicks and females has led to a serious age and sex imbalance, even amongst ostensibly healthy populations.[17]

In parts of the country, the Department of Conservation and local conservation groups have attempted to control predators of kākā through the use of traps, ground baiting and the aerial deployment of sodium fluoroacetate (1080). Where pest control has been carried out, there has been significant recovery of kākā populations. For example, in Pureora Forest Park 20 kākā were radio-tracked in an area to be treated with aerial 1080 in 2001. In nearby Waimanoa Forest, which was not to be treated with 1080, nine kākā were radio-tracked. In the area where 1080 was used, all 20 birds survived that season. Of the nine birds tagged in the untreated area, five were killed by predators that same season.[18]

Competition

Research has shown that honeydew is very important for breeding birds, especially those breeding in southern beech forests. The difficult nature of controlling the wasps makes the New Zealand kākā's future very uncertain.

References

  1. ↑ BirdLife International (2013). "Nestor meridionalis". IUCN Red List of Threatened Species. Version 2013.2. International Union for Conservation of Nature. Retrieved 26 November 2013.
  2. ↑ "Entry for kā on yourdictionary.com".
  3. 1 2 3 Wright, T.F.; Schirtzinger E. E.; Matsumoto T.; Eberhard J. R.; Graves G. R.; Sanchez J. J.; Capelli S.; Muller H.; Scharpegge J.; Chambers G. K.; Fleischer R. C. (2008). "A Multilocus Molecular Phylogeny of the Parrots (Psittaciformes): Support for a Gondwanan Origin during the Cretaceous". Mol Biol Evol 25 (10): 2141–2156. doi:10.1093/molbev/msn160. PMC 2727385. PMID 18653733.
  4. 1 2 3 Grant-Mackie, E.J.; J.A. Grant-Mackie; W.M. Boon; G.K. Chambers (2003). "Evolution of New Zealand Parrots". NZ Science Teacher 103.
  5. ↑ Juniper, T., Parr, M. (1998) Parrots: A guide to parrots of the world. New Haven, CT: Yale University Press (ISBN 0-300-07453-0)
  6. 1 2 De Kloet, Rolf S.; De Kloet, Siwo R. (September 2005). "The evolution of the spindlin gene in birds: sequence analysis of an intron of the spindlin W and Z gene reveals four major divisions of the Psittaciformes". Mol. Phylogenet. Evol. 36 (3): 706–21. doi:10.1016/j.ympev.2005.03.013. PMID 16099384.
  7. ↑ Schweizer, M.; Seehausen O; Güntert M; Hertwig ST (2009). "The evolutionary diversification of parrots supports a taxon pulse model with multiple trans-oceanic dispersal events and local radiations". Molecular Phylogenetics and Evolution 54 (3): 984–94. doi:10.1016/j.ympev.2009.08.021. PMID 19699808.
  8. ↑ CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), ISBN 978-0-8493-4258-5.
  9. ↑ "Click4Biology". Click4biology.info. Retrieved 27 December 2012.
  10. ↑ Falla RA, Sibson RB & Turbot EG (1966) A Field guide to the birds of New Zealand. Collins, London (ISBN 0-00-212022-4)
  11. ↑ "600th Kākā Banded at ZEALANDIA". Retrieved 9 November 2015.
  12. ↑ McArthur, Nikki; Harvey, Annette; Flux, Ian (October, 2015). State and trends in the diversity, abundance and distribution of birds in Wellington City. (PDF). Wellington: Greater Wellington Regional Council. Retrieved 18 January 2016. Check date values in: |date= (help)
  13. ↑ "Agathis australis, Kauri". Bushmans Friend. Retrieved 27 August 2007.
  14. ↑ Charles, K. E. 2012. Tree damage in Wellington as a result of foraging for sap and bark-dwelling invertebrates by the North Island kaka (Nestor meridionalis septentrionalis). Notornis 59:180-184
  15. ↑ "Landcare Research scientist John Innes talks about the extent of predation by introduced mammalian predators" (video interview). Retrieved 11 August 2011.
  16. ↑ Taylor, G.; et al. (2009). "Effect of controlling introduced predators on kaka (Nestor meridionalis) in the Rotoiti Nature Recovery Project".
  17. ↑ "New Zealand Journal of Ecology 22(1)". 1998.
  18. ↑ "The use of 1080 for pest control - Outcomes for bird populations". Department of Conservation. Retrieved 11 August 2011.

Further reading

External links

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