2016 in paleontology
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Paleontology, palaeontology or [palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2016.
Plants
Flowering plants
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Celastrus caducidentatus[2] |
Sp. nov |
In press |
Liang & Zhou in Liang et al. |
Miocene |
A species of Celastrus. |
|||
Cretacaeiporites aegyptiaca[3] |
Sp. nov |
In press |
Ibrahim et al. |
Late Cretaceous (Cenomanian) |
||||
Ettingshausenia vetviensis[4] |
Sp. nov |
In press |
Herman in Herman & Sokolova |
Late Cretaceous (Turonian–Coniacian) |
Vetvinskaya Member (Chalbugchan Group) |
A member of Platanofolia, a species of Ettingshausenia. |
||
Euanthus[5] |
Gen. et sp. nov |
In press |
Liu & Wang |
Jurassic (probably Callovian or Oxfordian) |
An early flowering plant. The type species is Euanthus panii. |
|||
Hedycaryoxylon burmeisteri[6] |
Sp. nov |
Valid |
Egerton, Williams & Lacovara |
Late Cretaceous (late Campanian to early Maastrichtian) |
Cerro Fortaleza Formation |
A member of Monimiaceae, a species of Hedycaryoxylon. |
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Juraherba[7] |
Gen. et sp. nov |
Valid |
Han & Wang in Han et al. |
Middle Jurassic |
An early flowering plant, resembling members of Hydatellaceae in general morphology and habit. The type species is Juraherba bodae. |
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Jusinghipollis ticoensis[8] |
Sp. nov |
Valid |
Llorens & Perez Loinaze |
Early Cretaceous (Aptian) |
Anfiteatro de Ticó Formation |
A flowering plant known from pollen grains, a species of Jusinghipollis. |
||
Lonchocarpus miocenicus[9] |
Sp. nov |
Valid |
Hernández-Damián et al. |
Miocene |
A species of Lonchocarpus. |
|||
Mahonia mioasiatica[10] |
Sp. nov |
In press |
Huang et al. |
Late Miocene |
Xiaolongtan Formation |
A species of Mahonia. |
||
Malloranga dentata[11] |
Sp. nov |
In press |
Conran, Lee & Reichgelt |
Miocene |
A member of Euphorbiaceae belonging to the subfamily Acalyphoideae, a species of Malloranga. |
|||
Parvileguminophyllum penzhinense[4] |
Sp. nov |
In press |
Herman in Herman & Sokolova |
Late Cretaceous (Turonian–Coniacian) |
Vetvinskaya Member (Chalbugchan Group) |
A member of Legumifolia, a species of Parvileguminophyllum. |
||
Quercus tibetensis[12] |
Sp. nov |
In press |
Xu, Su & Zhou in Xu et al. |
Late Miocene |
Lawula Formation |
An oak. |
||
Rosa fortuita[13] |
Sp. nov |
In press |
Su & Zhou in Su et al. |
Late Miocene |
A species of rose. |
|||
Zanthoxylum trachyspermum[14] |
Sp. nov |
In press |
Zhu & Zhou in Zhu et al. |
Late Miocene |
A member of Rutaceae, a species of Zanthoxylum. |
|||
Other seed plants
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Baiera aquilonia[15] |
Sp. nov |
Valid |
Pott, van der Burgh & van Konijnenburg-van Cittert |
Helvetiafjellet Formation |
A member of Ginkgoales, a species of Baiera. |
|||
Baiera baishanensis[16] |
Sp. nov |
In press |
Zhao et al. |
Early Cretaceous |
Yingzuilazi Formation |
A member of Ginkgoales, a species of Baiera. |
||
Cunninghamia beardii[17] |
Sp. nov |
Valid |
Buczkowski et al. |
Eocene |
A species of Cunninghamia. |
|||
Elatides sandaolingensis[18] |
Sp. nov |
In press |
Wang & Sun in Wang et al. |
Middle Jurassic |
Xishanyao Formation |
A conifer belonging or related to the family Cupressaceae; a species of Elatides. |
||
Galeacornea guayaguensis[19] |
Sp. nov |
Valid |
Prámparo, Narváez & Mego |
Early Cretaceous (probably Albian) |
A plant of uncertain phylogenetic placement, possibly a member or a relative of Gnetophyta; a species of Galeacornea. |
|||
Nidpuria falcatum[20] |
Sp. nov |
Valid |
Parveen & Bhowmik |
Triassic |
A member of Coniferopsida, possibly a member of Voltziales; a species of Nidpuria. |
|||
Phoenicopsis (Windwardia) daohugouensis[21] |
Sp. nov |
In press |
Huang et al. |
Middle Jurassic |
A member of Czekanowskiales, a species of Phoenicopsis. |
|||
Protocedroxylon shengjinbeigouense[22] |
Sp. nov |
In press |
Ding et al. |
Early Cretaceous |
A conifer, a species of Protocedroxylon. |
|||
Prototianshanoxylon[23] |
Gen. et 2 sp. nov |
In press |
Wei & Zhang in Wei et al. |
Permian |
A member of Coniferopsida. The type species is Prototianshanoxylon erdaogouense; genus also contains Prototianshanoxylon hamiense. |
|||
Pseudoephedra[24] |
Gen. et sp. nov |
In press |
Liu & Wang |
Early Cretaceous (Barremian–Aptian) |
An Ephedra-like seed plant of uncertain phylogenetic placement. The type species is Pseudoephedra paradoxa. |
|||
Sequoiadendron tchucoticum[25] |
Sp. nov |
Valid |
Sokolova in Sokolova & Moiseeva |
A species of Sequoiadendron. |
||||
Taxodioxylon heichengziense[22] |
Sp. nov |
In press |
Ding et al. |
Early Cretaceous |
A conifer, a species of Taxodioxylon. |
|||
Thujoxylon beipiaoense[22] |
Sp. nov |
In press |
Ding et al. |
Early Cretaceous |
A conifer, a species of Thujoxylon. |
|||
Umkomasia mongolica[26] |
Sp. nov |
In press |
Shi et al. |
Early Cretaceous |
A member of Umkomasiales (also known as Corystospermales), a species of Umkomasia. |
|||
Xenoxylon guangyuanensis[27] |
Sp. nov |
In press |
Tian et al. |
Late Triassic |
Xujiahe Formation |
|||
Xenoxylon junggarensis[28] |
Sp. nov |
Valid |
Wan et al. |
Late Triassic (Norian) |
Huangshanjie Formation |
A gymnosperm, a species of Xenoxylon. |
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Other plants
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Adinosporus[29] |
Gen. et 3 sp. nov |
In press |
Strother |
Cambrian |
A green alga belonging to the group Charophyta. The type species is Adinosporus voluminosus; genus also contains Adinosporus bullatus and Adinosporus geminus. |
|||
Brabantophyton[30] |
Gen. et sp. nov |
In press |
Momont, Gerrienne & Prestianni |
A member of Stenokoleales (a group of plants of uncertain phylogenetic placement, possibly related to the seed plants). The type species is Brabantophyton runcariense. |
||||
Cyclosorus scutum[31] |
Sp. nov |
Valid |
Naugolnykh et al. |
Eocene |
Changchang Formation |
A member of Thelypteridaceae, a species of Cyclosorus. |
||
Krameropteris[32] |
Gen. et sp. nov |
Valid |
Schneider, Schmidt & Heinrichs |
Cretaceous (Albian-Cenomanian) |
A member of Polypodiales belonging to the group Dennstaedtiaceae. The type species is Krameropteris resinatus. |
|||
Melvillipteris[33] |
Gen. et sp. nov |
In press |
Xue & Basinger |
Devonian (Famennian) |
A member of Rhacophytales. The type species is Melvillipteris quadriseriata. |
|||
Sigillariostrobus barkeri[34] |
Sp. nov |
Valid |
Thomas & Seyfullah |
Carboniferous (Bolsovian) |
A lycophyte belonging to the group Sigillariostrobaceae, a species of Sigillariostrobus. |
|||
Sigillariostrobus saltwellensis[34] |
Sp. nov |
Valid |
Thomas & Seyfullah |
Carboniferous (Langsettian–Duckmantian) |
A lycophyte belonging to the group Sigillariostrobaceae, a species of Sigillariostrobus. |
|||
Spissuspora[29] |
Gen. et sp. nov |
In press |
Strother |
Cambrian |
A green alga belonging to the group Charophyta. The type species is Spissuspora laevigata. |
|||
Tecaropteris[35] |
Gen. et sp. nov |
Valid |
Rozefelds, Dettmann & Clifford in Rozefelds et al. |
Paleogene |
Redbank Plains Formation |
A member of Pteridaceae belonging to the subfamily Ceratopteridoideae. The type species is Tecaropteris aquaincola |
||
Vidalgea[29] |
Gen. et sp. nov |
In press |
Strother |
Cambrian |
A green alga belonging to the group Charophyta. The type species is Vidalgea maculata. |
|||
Cnidarians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Bsokolovia[36] |
Gen. et sp. nov |
Valid |
Tsyganko |
Late Devonian |
A tabulate coral. The type species is Bsokolovia pershinae. |
|||
Caninophyllum charli[37] |
Sp. nov |
In press |
Denayer |
Carboniferous (Tournaisian) |
Yılanlı Formation |
A rugose coral belonging to the family Bothrophyllidae; a species of Caninophyllum. |
||
Eosiderastrea[38] |
Gen. et comb. nov |
In press |
Löser |
Cretaceous (late Valanginian to Santonian) |
A stony coral belonging to the family Siderastreidae; a new genus for the Cretaceous species formerly assigned to the extant genus Diploastrea. The type species is "Siderastrea" cuyleri Wells (1932) |
|||
Fungiaphyllia[39] |
Gen. et sp. nov |
In press |
Melnikova & Roniewicz |
Early Jurassic |
A stony coral. The type species is Fungiaphyllia communis. |
|||
Guembelastraea dronovi[39] |
Sp. nov |
In press |
Melnikova & Roniewicz |
Early Jurassic |
A stony coral, a species of Guembelastraea. |
|||
Oppelismilia spectabilis[39] |
Sp. nov |
In press |
Melnikova & Roniewicz |
Early Jurassic |
A stony coral, a species of Oppelismilia. |
|||
Palaeosiderofungia[38] |
Gen. et comb. nov |
In press |
Löser |
A stony coral belonging to the family Siderastreidae; a new genus for "Thamnasteria" exigua Reuss (1854). |
||||
Parepismilia dolichostoma[39] |
Sp. nov |
In press |
Melnikova & Roniewicz |
Early Jurassic |
A stony coral, a species of Parepismilia. |
|||
Parepismilia dronovi[39] |
Sp. nov |
In press |
Melnikova & Roniewicz |
Early Jurassic |
A stony coral, a species of Parepismilia. |
|||
Potyphyllum[40] |
Gen. et comb. nov |
Valid |
Coen-Aubert |
Devonian (late Frasnian) |
Belgium |
A rugose coral belonging to the family Phillipsastreidae. The type species is "Cyathophyllum" ananas Goldfuss (1826); genus also includes "Pseudoacervularia" dybowskii Różkowska (1953), "Pseudoacervularia" intercellulosa (Phillips, 1841) sensu Pickett (1967), "Schlüteria" lyskovensis Ermakova (1957), "Acervularia" macrommata Roemer (1855), "Phillipsastrea" plantana Różkowska (1979), "Cyathophyllum" profundum Michelin (1845), "Phillipsastrea" rozkowskae Scrutton (1968), "Pseudoacervularia" cf. smithi (Różkowska, 1953) sensu Pickett (1967), "Phillipsastrea" ananas veserensis Coen-Aubert (1974) and "Phillipsastrea" zerda Galle, 1992 in Hladil et al. (1992). |
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Talfania[41] |
Gen. et sp. nov |
In press |
Peel & McDermott |
Ordovician (Katian) |
Sholeshook Limestone Formation |
A solitary coral. The type species is Talfania calicula. |
||
Arthropods
Brachiopods
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Harperoides[42] |
Gen. et sp. nov |
Valid |
Baranov & Blodgett |
Devonian (Pragian) |
Soda Creek Limestone |
A member of Strophomenida belonging to the subfamily Mesodouvillininae. The type species is Harperoides alaskensis. |
||
Kjaerina (Kjaerina) gondwanensis[43] |
Sp. nov |
In press |
Colmenar |
Gabian Formation |
A rafinesquinid strophomenid brachiopod, a species of Kjaerina. |
|||
Kjaerina (Villasina)[43] |
Subgen. et 3 sp. et comb. nov |
In press |
Colmenar |
Ordovician (Katian) |
Cavá Formation |
A rafinesquinid strophomenid brachiopod, a subgenus of Kjaerina. The type species of the subgenus is Kjaerina (Villasina) pedronaensis; the subgenus also contains "Hedstroemina" almadenensis Villas (1995), as well as new species Kjaerina (Villasina) meloui and Kjaerina (Villasina) pyrenaica. |
||
Mesoleptostrophia belli[44] |
Sp. nov |
In press |
Earp |
Early Devonian |
Montys Hut Formation |
|||
Nasakia[45] |
Gen. et sp. nov |
Valid |
Streng et al. |
Cambrian |
A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Nasakia thulensis. |
|||
Pedderia[42] |
Gen. et sp. nov |
Valid |
Baranov & Blodgett |
Devonian (Pragian) |
Soda Creek Limestone |
A member of Rhynchonellida belonging to the family Pygmaellidae. The type species is Pedderia fragosa. |
||
Rafinesquina (Mesogeina)[43] |
Subgen. et comb. et 2 sp. nov |
In press |
Colmenar |
Ordovician (Katian) |
Bohdalec Formation |
A rafinesquinid strophomenid brachiopod, a subgenus of Rafinesquina. The type species of the subgenus is "Leptaena" pseudoloricata Barrande (1848); the subgenus also includes Rafinesquina pomoides Havlíček (1971), as well as new species Rafinesquina (Mesogeina) gabianensis and Rafinesquina (Mesogeina) loredensis. |
||
Spinobolus[46] |
Gen. et sp. nov |
In press |
Zhang et al. |
Cambrian |
Shuijingtuo Formation |
A linguloid brachiopod. The type species is Spinobolus popovi. |
||
Tapongaspirifer[44] |
Gen. et sp. nov |
In press |
Earp |
Early Devonian |
Montys Hut Formation |
A member of Spiriferida belonging to the family Filispiriferidae. The type species is Tapongaspirifer melodiae. |
||
Taungurungetes[44] |
Gen. et comb. nov |
In press |
Earp |
Early Devonian |
Montys Hut Formation |
A possible member of Devonochonetinae. The type species is "Chonetes" taggertyensis Gill (1945) from the Montys Hut Formation; genus also contains second, unnamed species from the Norton Gully Sandstone. |
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Thulatrypa[47] |
Gen. et sp. et comb. nov |
Valid |
Huang et al. |
Silurian |
An atrypoid brachiopod. The type species is Thulatrypa gregaria; genus also contains "Meifodia" orientalis Rong, Xu & Yang (1974). |
|||
Tomteluva[45] |
Gen. et sp. nov |
Valid |
Streng et al. |
Cambrian |
A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Tomteluva perturbata. |
|||
Molluscs
Echinoderms
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Apsidocrinus doreckae[48] |
Sp. nov |
Valid |
Konieczyński, Pisera & Fózy |
Early Cretaceous |
A cyrtocrinid crinoid, a species of Apsidocrinus. |
|||
Arabicodiadema[49] |
Gen. et comb. et sp. nov |
Valid |
Abdelhamid, El Qot & Abdelghany |
Cretaceous (Albian to Cenomanian) |
A heterodiadematid sea urchin. The type species is "Trochodiadema" dhofarense Roman (1991); genus also includes new species Arabicodiadema alii. |
|||
Ateleocystites? lansae[50] |
Sp. nov |
In press |
McDermott & Paul |
Ordovician (Katian) |
Slade and Redhill Beds |
A mitrate belonging to the family Anomalocystitidae, possibly a species of Ateleocystites. |
||
Loriolidiadema[49] |
Gen. et comb. nov |
Valid |
Abdelhamid, El Qot & Abdelghany |
Cretaceous (Aptian-Cenomanian) |
A heterodiadematid sea urchin. The type species is "Pseudodiadema" libanoticum de Loriol (1887); genus also includes Loriolidiadema sculptile (de Loriol, 1887). |
|||
Mariania comaschicariae[51] |
Sp. nov |
In press |
Stara, Borghi & Kroh |
Miocene (Aquitanian to early Burdigalian) |
Nurallao Formation |
A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania. |
||
Mariania stefaninii[51] |
Sp. nov |
In press |
Stara, Borghi & Kroh |
Miocene (late Burdigalian to early Langhian) |
Pantano Formation |
A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania. |
||
Phyllocrinus bersekensis[48] |
Sp. nov |
Valid |
Konieczyński, Pisera & Fózy |
Early Cretaceous |
A cyrtocrinid crinoid, a species of Phyllocrinus. |
|||
Conodonts
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Acodus zeballus[52] |
Sp. nov |
In press |
Voldman & Albanesi in Voldman et al. |
Early Ordovician |
||||
Gothodus vetus[52] |
Sp. nov |
In press |
Voldman & Albanesi in Voldman et al. |
Early Ordovician |
||||
Icriodus praealternatus ferus[53] |
Subsp. nov |
In press |
Wang et al. |
Late Devonian (Frasnian/Famennian boundary) |
Hongguleleng Formation |
A subspecies of Icriodus praealternatus. |
||
Icriodus plurinodosus[53] |
Sp. nov |
In press |
Wang et al. |
Late Devonian (early Famennian) |
Hongguleleng Formation |
A species of Icriodus. |
||
Icriodus stenoancylus junggarensis[53] |
Subsp. nov |
In press |
Wang et al. |
Late Devonian (early Famennian) |
Hongguleleng Formation |
A subspecies of Icriodus stenoancylus. |
||
Neopolygnathus huijunae[53] |
Sp. nov |
In press |
Wang et al. |
Late Devonian (early Famennian) |
Hongguleleng Formation |
A species of Neopolygnathus. |
||
Neospathodus planus[54] |
Sp. nov |
In press |
Chen et al. |
Early Triassic |
||||
Platyvillosus corniger[54] |
Sp. nov |
In press |
Chen et al. |
Early Triassic |
||||
“Polygnathus” pseudocommunis[53] |
Sp. nov |
In press |
Wang et al. |
Late Devonian (early Famennian) |
Hongguleleng Formation |
Possibly a species of Polygnathus. |
||
Zentagnathus[52] |
Gen. et comb. nov |
In press |
Voldman & Albanesi in Voldman et al. |
Early Ordovician |
A new genus for "Trapezognathus" primitivus Voldman, Albanesi & Zeballo in Voldman et al. (2013); genus also includes "Trapezognathus" argentinensis Rao et al. (1994) |
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Fishes
Amphibians
Temnospondyls
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Dahoumane et al. |
Early-Middle Triassic |
Zarzaïtine Series, Illizi Basin |
A mastodonsauroid temnospondyl, a species of Stanocephalosaurus. |
|||
Lissamphibians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Hyogobatrachus[56] |
Gen. et sp. nov |
In press |
Ikeda, Ota & Matsui |
Early Cretaceous |
Sasayama Group |
A frog. The type species is Hyogobatrachus wadai. |
||
Phosphotriton[57] |
Gen. et sp. nov |
In press |
Tissier, Rage, Boistel, Fernandez, Pollet, Garcia and Laurin |
Eocene |
A salamander. The type species is Phosphotriton sigei. |
|||
Prospea[58] |
Gen. et sp. nov |
Valid |
Chen et al. |
Paleocene |
Naranbulak Formation |
A member of Scaphiopodidae. The type species is Prospea holoserisca. |
||
Tambabatrachus[56] |
Gen. et sp. nov |
In press |
Ikeda, Ota & Matsui |
Early Cretaceous |
Sasayama Group |
A frog. The type species is Tambabatrachus kawazu. |
||
Lepidosaurs
Lizards
Research
- New anatomical data on the Late Cretaceous lizard Slavoia darevskii is published by Tałanda (2016), who interprets it as a stem-amphisbaenian.[59]
New taxa
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Janosikia[60] |
Gen. et comb. nov |
In press |
Čerňanský, Klembara & Smith |
Early Miocene |
A member of Lacertidae; a new genus for "Ophisaurus" ulmensis Gerhardt (1903). |
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Snakes
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Rieppelophis[61] |
Gen. et comb. nov |
Valid |
Scanferla, Smith & Schaal |
Eocene |
A member of Boidae. A new genus for "Messelophis" ermannorum Schaal & Baszio (2004). |
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Sauropterygians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Alexandronectes[62] |
Gen. et sp. nov |
Valid |
Otero et al. |
Late Cretaceous (early Maastrichtian) |
An aristonectine elasmosaurid plesiosaur. The type species is Alexandronectes zealandiensis. |
|||
Kawanectes[63] |
Gen. et comb. nov |
In press |
O'Gorman |
Late Cretaceous (late Campanian–early Maastrichtian) |
An elasmosaurid plesiosaur. The type species is "Trinacromerum" lafquenianum Gasparini & Goñi (1985). |
|||
Turtles
Research
- Fossils of Plesiochelys etalloni and Tropidemys langii, otherwise known from the Kimmeridgian of the Swiss and French Jura Mountains, are described from the British Kimmeridge Clay by Anquetin & Chapman (2016).[64]
Archosauriformes
Pseudosuchians
Research
- New fossil material from the Triassic (Ladinian or earliest Carnian) Pinheiros-Chiniquá Sequence of the Santa Maria Supersequence in Brazil attributed to the rauisuchian species Prestosuchus chiniquensis is described by Lacerda et al. (2016).[65]
- Description of postcranial skeletons of three specimens of the sphagesaurid Caipirasuchus (representing Caipirasuchus montealtensis, Caipirasuchus paulistanus and Caipirasuchus sp.) is published by Iori, Carvalho & Marinho (2016).[66]
- Fossils of the dyrosaurid crocodylomorph Hyposaurus are described from the Late Cretaceous Shendi Formation of Sudan by Salih et al. (2016).[67]
- The first unambiguous Middle Jurassic atoposaurid fossil (a dentary from the Isle of Skye, Scotland, United Kingdom, referred to Theriosuchus sp.) is described by Young et al. (2016).[68]
- New fossil material of Allodaposuchus precedens is described from the Late Cretaceous of France by Martin et al. (2016).[69]
- Fossil mekosuchine vertebrae, tentatively assigned to Mekosuchus whitehunterensis, are described from Riversleigh (Australia) by Stein, Archer & Hand (2016), who interpret them as confirming that even adult specimens of this species were smaller in snout-vent length than adults of extant small crocodilian species belonging to the genera Paleosuchus and Osteolaemus, and indicating that this species employed feeding behaviours that were unusual for crocodilians.[70]
New taxa
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
In press |
Barrios, Paulina-Carabajal & Bona |
Late Cretaceous |
Cerro Lisandro Formation |
A peirosaurid crocodyliform. The type species is Bayomesasuchus hernandezi. |
|||
Llanosuchus[72] |
Gen. et sp. nov |
In press |
Fiorelli et al. |
Late Cretaceous (Campanian?) |
Los Llanos Formation |
A notosuchian crocodyliform. The type species is Llanosuchus tamaensis. |
||
Sp. nov |
In press |
Fanti et al. |
Early Cretaceous |
A teleosaurid crocodylomorph, a species of Machimosaurus. |
||||
Gen. et sp. nov |
In press |
Lio et al. |
Late Cretaceous (Turonian–Coniacian) |
A peirosaurid crocodylomorph. The type species is Patagosuchus anielensis. |
||||
Scutarx[75] |
Gen. et sp. nov |
Valid |
Parker |
Late Triassic (middle Norian) |
An aetosaur. The type species is Scutarx deltatylus. |
|||
Basal dinosauromorphs
Research
- Marsicano et al. (2016) date the Chañares Formation, containing fossils of non-dinosaurian dinosauromorphs Lagerpeton, Lewisuchus, Marasuchus and Pseudolagosuchus, to early Carnian (236–234 Ma), 5–10 million years younger than previously thought. On this basis the authors postulate that the origin of dinosaurs was a relatively rapid event, as the transition from vertebrate communities containing only non-dinosaurian dinosauromorphs to communities containing the first dinosaurs occurred in less than a 5-million year interval.[76]
New taxa
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Martínez et al. |
Late Triassic (Norian) |
A lagerpetid dinosauromorph, a species of Dromomeron. |
||||
Non-avian dinosaurs
Research
- Six isolated spinosaurid quadrates, most likely coming from the Kem Kem Beds, are described by Hendrickx, Mateus & Buffetaut (2016), who interpret the differences in their anatomy as confirming the presence of two spinosaurine taxa in the Cenomanian of North Africa, rather than only one (Spinosaurus aegyptiacus).[78]
- A study of the phylogenetic relationships of tyrannosauroid theropods is published by Brusatte and Carr (2016).[79]
- New specimens of Elmisaurus rarus are described from the Late Cretaceous of Mongolia by Currie, Funston & Osmólska (2016).[80]
- New specimens of Leptorhynchos elegans and Leptorhynchos sp. are described from the Late Cretaceous of Canada by Funston, Currie & Burns (2016).[81]
- Isolated teeth of large-bodied iguanodontians are described from the Early Cretaceous (Albian) of Tunisia by Fanti et al. (2016).[82]
- A study of the frill bones of Protoceratops andrewsi, indicating that its frill increased in length and width during the ontogeny of the animal and that the growth of the frill was greater than the overall growth of the animal, is published by Hone, Wood & Knell (2016), who interpret these findings as indicating that Protoceratops most likely used its frill for sexual and social dominance signaling.[83]
- A revision of the species assigned to the genus Chasmosaurus is published by Campbell et al. (2016).[84]
New taxa
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et comb. nov |
Valid |
Late Jurassic |
A stegosaur; a new genus for "Stegosaurus" longispinus Gilmore (1914). This species was previously made the type species of the new genus Natronasaurus by Ulansky (2014); however, Galton & Carpenter (2016) claim it did not meet the requirements of the International Code of Zoological Nomenclature.[85] |
|||||
Gen. et sp. nov |
In press |
Xu et al. |
Late Cretaceous |
A non-hadrosaurid hadrosauroid ornithopod. The type species is Datonglong tianzhenensis. |
||||
Gen. et sp. nov |
Valid |
Martill et al. |
Early Jurassic (Hettangian) |
Blue Lias Formation |
A basal member of Neotheropoda. The type species is Dracoraptor hanigani. |
|||
Gen. et sp. nov |
Valid |
Prieto-Marquez, Erickson & Ebersole |
Late Cretaceous (latest Santonian) |
A hadrosaurid ornithopod. The type species is Eotrachodon orientalis. |
||||
Gen. et comb. nov |
Valid |
Peyre de Fabrègues & Allain |
Late Triassic |
Lower Elliot Formation |
A non-sauropod sauropodomorph. The type species is "Melanorosaurus" thabanensis Gauffre (1993). |
|||
Gen. et sp. nov |
Valid |
Rozadilla et al. |
Late Cretaceous (Maastrichtian) |
An iguanodontian ornithopod. The type species is Morrosaurus antarcticus. |
||||
Gen. et sp. nov |
Valid |
González Riga et al. |
Late Cretaceous (late Coniacian–early Santonian) |
A titanosaur sauropod. The type species is Notocolossus gonzalezparejasi. |
||||
Gen. et sp. nov |
In press |
Serrano-Brañas et al. |
Late Cretaceous |
Packard Shale Formation |
An ornithomimid theropod. The type species is Tototlmimus packardensis. |
|||
Viavenator[93] |
Gen. et sp. nov |
In press |
Filippi et al. |
Late Cretaceous (Santonian) |
A brachyrostran abelisaurid theropod. The type species is Viavenator exxoni. |
|||
Gen. et sp. nov |
In press |
Wang et al. |
Late Cretaceous (Cenomanian) |
Zhumapu Formation |
A basal member of Hadrosauroidea. The type species is Zuoyunlong huangi. |
|||
Birds
Research
- A new specimen of the Early Cretaceous species Archaeorhynchus spathula is described by Wang and Zhou (2016).[95]
- Zelenkov, Boev & Lazaridis (2016) reinterpret Otis hellenica from the Miocene of Greece, originally thought to be a bustard, as a member of Gruiformes belonging to the family Eogruidae and the subfamily Ergilornithinae; the authors classify it as a possible member of the genus Amphipelargus of uncertain specific assignment ("?Amphipelargus sp.").[96]
- Zelenkov, Volkova and Gorobets (2016) describe buttonquail fossils from the late Miocene of Hungary, southern Ukraine and northern Kazakhstan, and transfer the species Calidris janossyi Kessler (2009) to the genus Ortyxelos.[97]
- Agnolin, Tomassini and Contreras (2016) describe a distal end of tarsometatarsus from the late Miocene levels of the Loma de Las Tapias Formation (San Juan Province, Argentina), identified as the oldest seedsnipe fossil discovered so far.[98]
- Body mass estimates for 25 extinct pan-alcids and a study of body mass evolution in Pan-Alcidae are published by Smith (2016).[99]
- A study of eggshell fragments from the Pleistocene of Australia putatively referred to Genyornis newtoni is published by Grellet-Tinner, Spooner & Worthy (2016), who argue that these fossils are more likely to be remains of eggs laid by megapodes. Based on the similarities in the structure of eggshells of megapodes and dromornithids, the authors also hypothezise that dromornithids might be a sister group to galliforms rather than to or within anseriforms.[100]
- A study of burnt putative Genyornis eggshell fragments from the Pleistocene of Australia is published by Miller et al. (2016), who interpret them as confirming that eggs of Genyornis newtoni were harvested by humans.[101]
- Gastornithid and presbyornithid fossils are described from the early Eocene of Ellesmere Island (Canada) by Stidham & Eberle (2016).[102]
- A new tracksite with bird footprints (attributed to the ichnospecies Uvaichnites riojana), preserved in the early Miocene Lerín Formation (Bardenas Reales de Navarra Natural Park, Navarre, Spain), is described by Díaz-Martínez et al. (2016).[103]
New taxa
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Antarctoboenus [104] |
Gen. et sp. nov. |
In press |
Cenizo, Noriega & Reguero |
Early Eocene |
A stem-falconid. The type species is Antarctoboenus carlinii. |
|||
Bellulia [105] |
Gen. et sp. nov |
Junior homonym |
Wang, Zhou & Zhou |
Early Cretaceous |
A basal member of Ornithuromorpha. The type species is Bellulia rectusunguis. The generic name is preoccupied by Bellulia Fibiger (2008). |
|||
Chiappeavis [106] |
Gen. et sp. nov. |
Valid |
O’Connor et al. |
Early Cretaceous |
A member of Enantiornithes, probably belonging to the family Pengornithidae. The type species is Chiappeavis magnapremaxillo. |
|||
Chongmingia [107] |
Gen. et sp. nov. |
Valid |
Wang et al. |
Early Cretaceous (Aptian) |
A member of Avialae of uncertain phylogenetic placement; it might be a member of Avebrevicauda that didn't belong to Omnivoropterygidae, Confuciusornithidae or Ornithothoraces, or one of the basalmost known avialans, more closely related to Jeholornis and Euavialae than Archaeopteryx was. The type species is Chongmingia zhengi. |
|||
Daphoenositta trevorworthyi [108] |
Sp. nov |
Valid |
Nguyen |
Miocene |
Riversleigh World Heritage Area |
A sittella (a member of the family Neosittidae), a species of Daphoenositta. |
||
Dingavis [109] |
Gen. et sp. nov |
Valid[110] |
O'Connor, Wang & Hu |
Early Cretaceous |
A basal member of Ornithuromorpha. The type species is Dingavis longimaxilla. |
|||
Galligeranoides [111] |
Gen. et sp. nov. |
In press |
Bourdon, Mourer-Chauviré, & Laurent |
Eocene (middle Ypresian) |
A member of the family Geranoididae. The type species is G. boriensis. |
|||
Linyiornis [112] |
Gen. et sp. nov. |
Valid |
Wang et al. |
Early Cretaceous |
A member of Enantiornithes. The type species is Linyiornis amoena. |
|||
Pterosaurs
Research
- New information on the braincase anatomy of Pterodaustro guinazui is published by Codorniú, Paulina-Carabajal & Gianechini (2016).[113]
Other reptiles
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Langeronyx[114] |
Gen. et comb. nov |
Valid |
Ezcurra, Montefeltro & Butler |
Middle Triassic (Anisian) |
A rhynchosaur; a new genus for "Rhynchosaurus" brodiei Benton (1990). |
|||
Synapsids
Non-mammalian synapsids
Research
- A revision of the systematics of the gorgonopsian subfamily Rubidgeinae is published by Kammerer (2016).[115]
Non-eutherian mammals
Research
- A partial mandible of the amphitheriid Palaeoxonodon ooliticus, previously known only from isolated teeth, is described from the Middle Jurassic (late Bathonian) Kilmaluag Formation (Isle of Skye, Scotland, United Kingdom) by Close et al. (2016).[116]
New taxa
Metatherians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Kutjamarcoot[117] |
Gen. et sp. nov |
In press |
Chamberlain et al. |
Early Miocene |
Wipajiri Formation |
A bandicoot. The type species is Kutjamarcoot brevirostrum. |
||
Minusculodelphis modicum[118] |
Sp. nov |
Valid |
Oliveira, Zimicz & Goin |
Itaboraí Quarry |
A member of Jaskhadelphyidae, a species of Minusculodelphis. |
|||
Eutherian mammals
Research
- A study of the phylogenetic relationships of the Paleocene placental mammals is published by Halliday, Upchurch and Goswami (2016).[119]
- New fossil material of the Pleistocene wildebeest-like bovid Rusingoryx atopocranion is described from the Rusinga Island (Kenya) by O’Brien et al. (2016), who note the presence of large, hollow, bony nasal crests in this mammal, similar to crests present in hadrosaurid dinosaurs.[120]
- A revision of the systematics of the North American members of Nimravidae is published by Barrett (2016).[121]
- Virtual cranial endocasts of the Eocene rodents Paramys copei and Paramys delicatus are described by Bertrand, Amador-Mughal and Silcox (2016).[122]
- Fossils of the probable relative of the gorillas, Chororapithecus abyssinicus, are dated to ~8.0 Myr by Katoh et al. (2016).[123]
New taxa
Afrotherians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Balaguer & Alba |
Eocene |
A member of Dugongidae, a species of Prototherium. |
||||
Sinomammut[125] |
Gen. et sp. nov |
Valid |
Mothé et al. |
Late Miocene |
A member of Mammutidae. The type species is Sinomammut tobieni. |
|||
Sp. nov |
In press |
Wang et al. |
Late Miocene |
A gomphothere, a species of Sinomastodon. |
||||
Odd-toed ungulates
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Koufos & Vlachou |
Miocene (Turolian) |
A member of Equidae, a species of Hipparion. |
||||
Sp. nov |
In press |
Koufos & Vlachou |
Miocene (Turolian) |
A member of Equidae, a species of Hipparion. |
||||
Even-toed ungulates
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Diplobunops kardoula[128] |
Sp. nov |
Valid |
Emery, Davis & Hopkins |
Eocene (Duchesnean) |
An agriochoerid oreodont, a species of Diplobunops. |
|||
Megaloceros stavropolensis[129] |
Sp. nov |
Valid |
Titov & Shvyreva |
Early Pleistocene (late Villafranchian) |
A species of Megaloceros. |
|||
Sp. nov |
In press |
Danowitz, Domalski & Solounias |
Early Miocene |
A member of Ruminantia, a species of Prolibytherium. |
||||
Cetaceans
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Mesoplodon posti[131] |
Sp. nov |
Valid |
Lambert & Louwye |
Pliocene (Zanclean) |
||||
Sp. nov |
Valid |
Marx, Bosselaers & Louwye |
Miocene (late Tortonian) |
A member of Cetotheriidae, a species of Metopocetus. |
||||
Carnivorans
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Actiocyon parverratis[133] |
Sp. nov |
In press |
Smith, Czaplewski & Cifelli |
Miocene (Barstovian) |
A simocyonine ailurid (a relative of the red panda), a species of Actiocyon. |
|||
Alagtsavbaatar[134] |
Gen. et comb. nov |
Valid |
Egi et al. |
Late Eocene |
Ergilin Dzo Formation |
A member of Feliformia. The type species is "Stenoplesictis" indigenus Dashzeveg (1996). |
||
Brevimalictis[133] |
Gen. et sp. nov |
In press |
Smith, Czaplewski & Cifelli |
Miocene (Barstovian) |
A member of Mustelidae of uncertain phylogenetic placement. The type species is Brevimalictis chikasha. |
|||
Lutraeximia[135] |
Gen. et sp. nov |
Valid |
Cherin et al. |
Early Pleistocene (late Villafranchian) |
An otter. The type species is Lutraeximia umbra. |
|||
Sp. nov |
In press |
Zhu et al. |
Early Pleistocene |
A machairodontine felid, a species of Megantereon. |
||||
Negodiaetictis[133] |
Gen. et sp. nov |
In press |
Smith, Czaplewski & Cifelli |
Miocene (Barstovian) |
A member of Mustelidae of uncertain phylogenetic placement. The type species is Negodiaetictis rugatrulleum. |
|||
Rodents
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Brachyscirtetes tomidai[137] |
Sp. nov |
Valid |
Li |
Late Miocene |
A member of Dipodidae, a species of Brachyscirtetes. |
|||
Gliruloides[138] |
Gen. et sp. et comb. nov |
In press |
Wu et al. |
Late Oligocene to early Miocene |
Junggar Basin |
A dormouse related to the forest dormouse. The type species is Gliruloides zhoui; genus also includes "Vasseuromys" duplex Ünay (1994). |
||
Megacricetodon hellenicus[139] |
Sp. nov |
In press |
Oliver & Peláez-Campomanes |
Early Miocene |
A cricetid rodent, a species of Megacricetodon. |
|||
Mimomys chandolensis[140] |
Sp. nov |
In press |
Tiunov, Golenishchev & Voyta |
Late Pleistocene |
An arvicoline cricetid, a species of Mimomys. |
|||
Minocastor[141] |
Gen. et sp. nov |
Valid |
Mörs, Tomida & Kalthoff |
Early Miocene |
Nakamura Formation |
A member of Castoridae. The type species is Minocastor godai. |
||
Myocricetodon tomidai[142] |
Sp. nov |
Valid |
Lindsay & Flynn |
Early Miocene |
Chitarwata Formation |
A member of Cricetidae, a species of Myocricetodon. |
||
Paciculus walshi[143] |
Sp. nov |
Valid |
Lindsay et al. |
Oligocene |
A member of Cricetidae, a species of Paciculus. |
|||
Potamarchus adamiae[144] |
Sp. nov |
In press |
Kerber et al. |
Late Miocene |
Solimões Formation |
A potamarchine dinomyid, a species of Potamarchus. |
||
Potwarmus mahmoodi[142] |
Sp. nov |
Valid |
Lindsay & Flynn |
Early Miocene |
Vihowa Formation |
A member of Cricetidae, a species of Potwarmus. |
||
Primus cheemai[142] |
Sp. nov |
Valid |
Lindsay & Flynn |
Late Oligocene to Early Miocene |
Chitarwata Formation |
A member of Cricetidae, a species of Primus. |
||
Pseudopotamarchus[144] |
Gen. et sp. nov |
In press |
Kerber et al. |
Late Miocene |
Solimões Formation |
A potamarchine dinomyid. The type species is Pseudopotamarchus villanuevai. |
||
Primates
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Beard et al. |
Early Oligocene |
A species of Apidium. |
||||
Semnopithecus gwebinensis[146] |
Sp. nov |
Valid |
Takai et al. |
Late Pliocene |
A gray langur. |
|||
Other eutherians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Amphilagus tomidai[147] |
Sp. nov |
Valid |
Erbajeva, Angelone & Alexeeva |
Miocene |
A lagomorph, a species of Amphilagus. |
|||
Hipposideros (Pseudorhinolophus) amenhotepos[148] |
Sp. nov |
Valid |
Gunnell et al. |
Miocene |
||||
Ounalashkastylus[149] |
Gen. et sp. nov |
Valid |
Chiba et al. |
Miocene |
A desmostylian. The type species is Ounalashkastylus tomidai. |
|||
Ptilocercus kylin[150] |
Sp. nov |
Valid |
Li & Ni |
Earliest Oligocene |
Lijiawa Mammalian Fossil locality |
A treeshrew related to the pen-tailed treeshrew. |
||
Rzebikia[151] |
Gen. et comb. nov |
In press |
Sansalone, Kotsakis & Piras |
Pliocene to Pleistocene (Villanyian) |
A relative of the American shrew mole. A new genus for "Neurotrichus" polonicus Skoczeń (1980) and "Neurotrichus" skoczeni Zijlstra (2010). |
|||
Sanshuilophus[152] |
Gen. et sp. nov |
In press |
Mao et al. |
Early Eocene |
Huayong Formation |
A member of Phenacolophidae (a group of archaic ungulate mammals of uncertain phylogenetic placement). The type species is Sanshuilophus zhaoi. |
||
Other animals
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Annulitubus[153] |
Gen. et sp. nov |
In press |
Vinn et al. |
Devonian (late Emsian) |
Ponta Grossa Formation |
An annelid, possibly a polychaete. The type species is Annulitubus mutveii. |
||
Bolidium bertii[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Bolidium. |
|||
Camerospongia tuberculata[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Camerospongia. |
|||
Camerospongia visentinae[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Camerospongia. |
|||
Cavispongia scarpai[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Cavispongia. |
|||
Corallistes multiosculata[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Corallistes. |
|||
Coronispongia[154] |
Gen. et sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge. The type species is Coronispongia confossa. |
|||
Hexactinella clampensis[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Hexactinella. |
|||
Sp. nov |
In press |
Yang, Zhao & Zhang |
Cambrian |
A stem-group priapulid, a species of Ottoia. |
||||
Rigonia[154] |
Gen. et sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge. The type species is Rigonia plicata. |
|||
Sarmentofascis zamparelliae[157] |
Sp. nov |
Valid |
Schlagintweit, Frijia & Parente |
Late Cretaceous (early Campanian) |
A sponge, a species of Sarmentofascis. |
|||
Stauractinella eocenica[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Stauractinella. |
|||
Toulminia italica[154] |
Sp. nov |
Valid[155] |
Frisone, Pisera & Preto |
Eocene (early Lutetian) |
A sponge, a species of Toulminia. |
|||
Sp. nov |
In press |
Peng et al. |
Cambrian |
A palaeoscolecid worm, a species of Wronascolex. |
||||
Other organisms
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Bianchina[159] |
Gen. et sp. nov |
In press |
Schiøler |
Cretaceous (late Aptian–middle Cenomanian) |
East Coast Basin |
A dinoflagellate. The type species is Bianchina hieroglyphica. |
||
Eohalothece[160] |
Gen. et sp. nov |
Valid |
Strother & Wellman |
Precambrian |
Cailleach Head Formation |
A member of Cyanobacteria belonging to the family Chroococcaceae. The type species is Eohalothece lacustrina. |
||
Perissothallus dekovensis[161] |
Sp. nov |
In press |
Pšenička & Krings |
Carboniferous (Gzhelian) |
Slaný Formation |
An alga of uncertain phylogenetic placement, a species of Perissothallus. |
||
Trewinomyces[162] |
Gen. et sp. nov |
In press |
Krings, Taylor & Martin |
Early Devonian |
A fungus of uncertain phylogenetic placement, showing similarities to members of the genera Macrochytrium (Chytridiomycota) and Blastocladiella (Blastocladiomycota). The type species is Trewinomyces annulifer. |
|||
References
- ↑ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
- ↑ Xiao-Qing Liang, David K. Ferguson, Frédéric M.B. Jacques, Tao Su, Li Wang and Zhe-Kun Zhou (2016). "A new Celastrus species from the middle Miocene of Yunnan, China and its palaeoclimatic and palaeobiogeographic implications". Review of Palaeobotany and Palynology. in press. doi:10.1016/j.revpalbo.2015.11.005.
- ↑ Mohamed I.A. Ibrahim, Mohamed K. Zobaa, Zainab M. El–Noamani and Sameh S. Tahoun (2016). "A review of the angiosperm pollen genus Cretacaeiporites Herngreen, with one new species from the Upper Cretaceous of Egypt". Palynology. in press. doi:10.1080/01916122.2015.1093561.
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