2015 in paleontology

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Paleontology, palaeontology or [palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2015.

Plants

Main article: 2015 in paleobotany

Cnidarians

Name Novelty Status Authors Age Unit Location Notes Images

Acropora darrellae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora elenae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora emanuelae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora hasibuani[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora laurae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Late Oligocene

 Malaysia

A staghorn coral, a species of Acropora.

Acropora renemai[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Actinaraea michoacanensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (early Aptian)

Cumburindio Formation

 Mexico

A stony coral belonging to the family Actinacididae, a species of Actinaraea.

Actinastrea chumbitaroensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Actinastraeidae, a species of Actinastrea.

Australastraea[4]

Gen. et comb. et 2 sp. nov

Valid

Denayer & Webb

Carboniferous (Mississippian)

 Australia

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae. A new genus for "Lithostrotion" columnare Etheridge (1900); genus also contains "Orionastraea" columellaris Pickett (1966), "Lithostrotion" parvicolumnare Pickett (1966), "Lithostrotion" wilkinsoni Pickett (1966) and "Orionastraea" flemingi Webb (1990), as well as new species Australastraea arcifera and Australastraea carinata.

Axisvacuus tenerus[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Antiphyllidae, a species of Axisvacuus.

Birkenmajerites[5]

Gen. et sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Ostravaiaidae. The type species is Birkenmajerites primus.

Cairnsipsammia[6]

Gen. nov

Valid

Baron-Szabo

Early Cretaceous (late Barremian–early Aptian)

Schrattenkalk Formation

 Austria

A stony coral belonging to the family Dendrophylliidae.

Chelmia[5]

Gen. et sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida. The type species is Chelmia radiata.

Cionodendron smithi[4]

Sp. nov

Valid

Denayer & Webb

Carboniferous (early Viséan)

 Australia

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae; a species of Cionodendron.

Craspedophyllia japonica[7]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Reimaniphylliidae, a species of Craspedophyllia.

Crateroseris stefani[8]

Sp. nov

Valid

Eliášová

Late Jurassic (Tithonian)

Štramberk Limestone

 Czech Republic

A stony coral belonging to the family Synastraeidae, a species of Crateroseris.

Cryptosepta[9]

Gen. et sp. nov

Valid

Gretz, Lathuilière & Martini

Early Jurassic (Hettangian)

 France

A zardinophyllid coral. The type species is Cryptosepta nuda.

Echinophyllia sassellensis[10]

Sp. nov

Valid[11]

Budd & Bosellini

Oligocene

 Italy

Eocolumastrea[12]

Gen. et comb. nov

Valid

Löser & Zell

Cretaceous

 Romania

A stony coral belonging to the group Faviina and the family Columastraeidae. A new genus for "Columnocoenia" ksiazkiewiczi bucovinensis Morycowa (1971) (raised to the rank of a separate species Eocolumastrea bucovinensis)

Goniophyllum osloense[13]

Sp. nov

Valid

Johannessen in Wright

Silurian (late Telychian)

Vik Formation

 Norway

An operculate coral, a species of Goniophyllum.

Guembelastraea kanmerae[7]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Guembelastraeidae, a species of Guembelastraea.

Halucinophyllia[8]

Gen. et sp. nov

Valid

Eliášová

Late Jurassic (Tithonian)

Štramberk Limestone

 Czech Republic

A stony coral belonging to the family Latomeandridae. The type species is Halucinophyllia subridens.

Heteropistophyllum carchensis[14]

Sp. nov

Valid

Löser, Arias & Vilas

Early Cretaceous (Aptian and Albian)

 Mexico
 Spain

A stony coral belonging to the family Elasmocoeniidae, a species of Heteropistophyllum.

Isopora matahari[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Isopora.

Latiphyllia mexicana[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Montlivaltiidae, a species of Latiphyllia.

Margarosmilia mizukamia[7]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Margarophylliidae, a species of Margarosmilia.

Mycetophyllopsis azteca[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Montlivaltiidae, a species of Mycetophyllopsis.

Munusculum[8]

Gen. et sp. nov

Valid

Eliášová

Late Jurassic (Tithonian)

Štramberk Limestone

 Czech Republic

A stony coral belonging to the suborder Microsolenina. The type species is Munusculum martinaseki.

Nervophyllum lukoviensis[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Aulophyllidae, a species of Nervophyllum.

Nudacolumastrea[12]

Gen. et sp. nov

Valid

Löser & Zell

Cretaceous

 Spain

A stony coral belonging to the group Faviina and the family Columastraeidae. The type species is Nudacolumastrea stefani.

Occulogermen[5]

Gen. et sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae. The type species is Occulogermen luciae.

Orbignygyra? incognita[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Dendrogyridae, possibly a species of Orbignygyra.

Ovalastrea malpaso[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Latomeandridae, a species of Ovalastrea.

Paraclausastrea vorarlbergensis[15]

Sp. nov

Valid

Baron-Szabo

Early Cretaceous (late Barremian-early Aptian)

Schrattenkalk Formation

 Austria

A montlivaltiid stony coral, a species of Paraclausastrea.

Paracycloseris effrenatus[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Cunnolitidae, a species of Paracycloseris.

Paraleptoria[10]

Gen. et comb. nov

Valid[11]

Budd & Bosellini

Eocene (Lutetian) to Oligocene (Rupelian)

 Italy

A new genus for "Meandrina" polygonalis Catullo (1856); genus also includes "Diploria" flexuosissima d'Achiardi (1868).

Parascolymia bracherti[16]

Sp. nov

Valid

Reuter in Reuter, Wiedl & Piller

Miocene (Langhian‒early Serravallian)

Leitha Limestone

 Austria

A stony coral belonging to the family Lobophylliidae, a species of Parascolymia.

Pickettodendron[4]

Gen. et comb. et sp. nov

Valid

Denayer & Webb

Carboniferous (Tournaisian)

 Australia

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae. A new genus for "Lithostrotion" williamsi Pickett (1966); genus also contains "Schoenophyllum" dalmaensis Webb (1990), as well as new species Pickettodendron nudum.

Preverastraea coatlicueae[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Rhipidogyridae, a species of Preverastraea.

Preverastraea tociae[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Rhipidogyridae, a species of Preverastraea.

Retiophyllia tosaensis[7]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Reimaniphylliidae, a species of Retiophyllia.

Rotiphyllum plumeum[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Antiphyllidae, a species of Rotiphyllum.

Saltocyathus cumburindioensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (early Aptian)

Cumburindio Formation

 Mexico

A stony coral belonging to the family Rhipidogyridae, a species of Saltocyathus.

Sclerangia[17]

Gen. et sp. nov

Valid

Baron-Szabo & Cairns

Miocene

Chipola Formation

 United States

A rhizangiid stony coral. The type species is Sclerangia floridana.

?Sochkineophyllum symmetricum[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Polycoeliidae, possibly a species of Sochkineophyllum.

Sylviella[8]

Gen. et comb. et 4 sp. nov

Valid

Eliášová

Late Jurassic (Tithonian)

Štramberk Limestone

 Czech Republic

A stony coral belonging to the suborder Caryophylliina. The type species is "Haplaraea" columnaris Ogilvie (1897); genus also contains new species Sylviella exquisita, Sylviella multisepta, Sylviella noveni and Sylviella benjamin.

Sylviellopsis[8]

Gen. et sp. nov

Valid

Eliášová

Late Jurassic (Tithonian)

Štramberk Limestone

 Czech Republic

A stony coral belonging to the suborder Caryophylliina. The type species is Sylviellopsis erici.

Tegocoenia[8]

Gen. et comb. nov

Valid

Eliášová

Late Jurassic (Tithonian)

Štramberk Limestone

 Czech Republic

A stony coral belonging to the family Placophylliidae. The type species is "Diplocoenia" jasenica Frajová (1960).

Thalamocaeniopsis mexicanensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Thamnasteriidae, a species of Thalamocaeniopsis.

Thamnarea hornosensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (early Aptian)

Cumburindio Formation

 Mexico

A stony coral belonging to the family Actinacididae, a species of Thamnarea.

Thamnasteria tonantzinae[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Thamnasteriidae, a species of Thamnasteria.

Thamnasteriamorpha okudai[7]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Tropiastraeidae, a species of Thamnasteriamorpha.

Thecosmilia guerreroensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Montlivaltiidae, a species of Thecosmilia.

Variabilifavia ausuganensis[10]

Sp. nov

Valid[11]

Budd & Bosellini

Oligocene (late Rupelian–early Chattian)

 Italy

Zaphrentites rotiphylloides[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Stereophrentidae, a species of Zaphrentites.

Zaphriphyllum daleki[18]

Sp. nov

Valid

Denayer

Carboniferous (late Tournaisian)

Yılanlı Formation

 Turkey

A rugose coral belonging to the family Ekvasophyllidae, a species of Zaphriphyllum.

Zaphrufimia anceps[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Stereophrentidae, a species of Zaphrufimia.

Arthropods

Bryozoans

Name Novelty Status Authors Age Unit Location Notes Images

Acantholunaria[19]

Gen. et sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A cystoporate bryozoan. The type species is Acantholunaria expansa.

Adeonellopsis incompta[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An adeonid cheilostome bryozoan, a species of Adeonellopsis.

Amphiblestrum paleogenicum[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A calloporid cheilostome bryozoan, a species of Amphiblestrum.

Amplexopora crassiparietum[22]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

An amplexoporid trepostome bryozoan, a species of Amplexopora.

Amplexopora evae[22]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

An amplexoporid trepostome bryozoan, a species of Amplexopora.

Andreella dubia[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A microporid cheilostome bryozoan, a species of Andreella.

Anaphragma undulata[23]

Sp. nov

Valid

Jiménez-Sánchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Anaphragma.

Aostipora elongata[24]

Sp. nov

Valid

Jiménez-Sánchez in Jiménez-Sánchez, Villas & Vennin

Ordovician (late Katian)

Khabt-el-Hajar Formation

 Morocco

A trepostomate bryozoan, a species of Aostipora.

Arachnopusia dimorpha[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An arachnopusiid cheilostome bryozoan, a species of Arachnopusia.

Arthropoma renipora[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A lacernid schizoporelloid bryozoan, a species of Arthropoma.

Aspidostoma clava[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Aspidostoma.

Aspidostoma gelasinus[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Aspidostoma.

Aspidostoma twinn[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Aspidostoma.

Atactoporella moroccoensis[23]

Sp. nov

Valid

Jiménez-Sánchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Atactoporella.

Baculopora redondensis[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A member of Fenestrata belonging to the family Acanthocladiidae; a species of Baculopora.

Bryopesanser bragai[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian to Serravallian)

 Indonesia

An escharinid schizoporelloid bryozoan, a species of Bryopesanser.

Bryopesanser sanfilippoae[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Burdigalian-Langhian boundary to Serravallian)

 Indonesia

An escharinid schizoporelloid bryozoan, a species of Bryopesanser.

Buffonellaria sagittaria[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A celleporid celleporoid bryozoan, a species of Buffonellaria.

Caberoides gordoni[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A catenicellid bryozoan, a species of Caberoides.

Celiopsis[27]

Gen. et sp. et comb. nov

Valid

Zágoršek, Gordon & Vávra

Eocene to Oligocene

 Austria
 Hungary
 Italy
 Poland
 Slovakia
 Spain

A chlidoniopsid cheilostome bryozoan. The type species is Celiopsis vici; genus also contains "Crisidia" vindobonensis Reuss (1847) and "Chlidoniopsis" vavrai Zágoršek (2003).

Cellaria bicuspidata[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Cellaria gigas[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Cellaria inarticulata[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Cellaria palatum[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Chaperiopsis cookae[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A chaperiid cheilostome bryozoan, a species of Chaperiopsis.

Chataimulosia revelator[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lepraliellid cheilostome bryozoan, a species of Chataimulosia.

Cladobryopastor[20]

Gen. et sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A bryopastorid cheilostome bryozoan. The type species is Cladobryopastor philipi.

Conopeum stamenocelloides[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An electrid cheilostome bryozoan, a species of Conopeum.

Curvipora[19]

Gen. et sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A cystoporate bryozoan. The type species is Curvipora monostylata.

Cyphotrypa regularis[24]

Sp. nov

Valid

Jiménez-Sánchez in Jiménez-Sánchez, Villas & Vennin

Ordovician (late Katian)

Khabt-el-Hajar Formation

 Morocco

A trepostomate bryozoan, a species of Cyphotrypa.

Cystodictya gallensis[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Cystodictyonidae; a species of Cystodictya.

Dactylostega spiralis[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A foveolariid cheilostome bryozoan, a species of Dactylostega.

Diedropora[20]

Gen. et sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An adeonoid cheilostome bryozoan. The type species is Diedropora gravabilis.

Dybowskiella piriforme[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Dybowskiella.

Dybowskiella rotunda[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Dybowskiella.

Dyscritella ornata[28]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Dyscritella perforata[29]

Sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A dyscritellid trepostome bryozoan, a species of Dyscritella.

Eridopora suarezi[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Eridopora.

Exechonella chathamensis[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An exechonellid cheilostome bryozoan, a species of Exechonella.

Exochella abigailae[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Exochella linearis[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Exochella reidae[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Exochella woodae[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Fabifenestella macrofenestrata[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A member of Fenestrata belonging to the family Fenestellidae; a species of Fabifenestella.

?Filaguria kalimantanensis[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

A cribrilinid bryozoan, possibly a species of Filaguria.

Fistulipora tolokonnikovae[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Fistulipora.

Floridina elegans[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An onychocellid cheilostome bryozoan, a species of Floridina.

Gigantopora milenae[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

A gigantoporid schizoporelloid bryozoan, a species of Gigantopora.

Gigantopora grandis[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A gigantoporid cheilostome bryozoan, a species of Gigantopora.

Gigantopora modesta[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A gigantoporid cheilostome bryozoan, a species of Gigantopora.

Gorjunopora[26]

Gen. et sp. et comb. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France
 Russia

A member of Fenestrata belonging to the family Acanthocladiidae. The type species is Gorjunopora gallica; genus also includes Gorjunopora triangulata (Schulga-Nesterenko, 1955).

Hemicyclopora dissidens[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Hemicyclopora.

Hemicyclopora ventricosa[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Hemicyclopora.

Heminematopora gaetula[30]

Sp. nov

Valid

Ernst, Jiménez-Sánchez & Baidder

Ordovician (Katian)

 Morocco

A species of Heminematopora.

Heteropora scholzi[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cerioporid cyclostome bryozoan, a species of Heteropora.

Heterotrypa ringerikense[22]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

A heterotrypid trepostome bryozoan, a species of Heterotrypa.

?Hippomenella devatasae[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A romancheinid lepralielloid bryozoan, possibly a species of Hippomenella.

?Hippomenella uniserialis[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A romancheinid lepralielloid bryozoan, possibly a species of Hippomenella.

Hippopleurifera barbosae[31]

Sp. nov

Valid

Ramalho et al.

Miocene

Pirabas Formation

 Brazil

A cheilostome bryozoan belonging to the superfamily Lepralielloidea and the family Romancheneidae, a species of Hippopleurifera.

Hippopleurifera confusa[31]

Sp. nov

Valid

Ramalho et al.

Miocene

Pirabas Formation

 Brazil

A cheilostome bryozoan belonging to the superfamily Lepralielloidea and the family Romancheneidae, a species of Hippopleurifera.

Hippopodina indicata[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian to Langhian)

 Indonesia

A hippopodinid schizoporelloid bryozoan, a species of Hippopodina.

Hyporosopora nana[32]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

A plagioeciid cyclostome bryozoan, a species of Hyporosopora.

Hyporosopora negevensis[32]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

A plagioeciid cyclostome bryozoan, a species of Hyporosopora.

Idmonea snehi[32]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

A multisparsid cyclostome bryozoan, a species of Idmonea.

Illusiopora[20]

Gen. et 2 sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A schizoporelloid cheilostome bryozoan. The type species is Illusiopora bifax; genus also contains Illusiopora recta.

Isostylus veserensis[33]

Sp. nov

Valid

Ernst, Tolokonnikova & Denayer

Devonian (late Frasnian)

 Belgium

A species of Isostylus.

Lacerna ordinaria[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lacernid cheilostome bryozoan, a species of Lacerna.

Lagenipora sciutoi[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A celleporid celleporoid bryozoan, a species of Lagenipora.

Lenapora gurievensis[34]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Lenapora.

Leptotrypa enodis[19]

Sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A trepostome bryozoan, a species of Leptotrypa.

Leptotrypa hexagona[28]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Leptotrypa perforata[19]

Sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A trepostome bryozoan, a species of Leptotrypa.

Macrocamera obesa[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An eminooeciid cheilostome bryozoan, a species of Macrocamera.

Margaretta amitabhae[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Burdigalian-Langhian boundary to Serravallian)

 Indonesia

A margarettid schizoporelloid bryozoan, a species of Margaretta.

Marssonopora connexa[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A calloporid cheilostome bryozoan, a species of Marssonopora.

Mediaporina kiaeri[22]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

An arthrostylid cryptostome bryozoan, a species of Mediaporina.

Megacanthopora enodata[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A member of Cryptostomata belonging to the family Rhomboporidae; a species of Megacanthopora.

Melychocella bilamellata[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Melychocella.

Melychocella obliqua[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Melychocella.

Mesenteripora jamesi[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A plagioeciid cyclostome bryozoan, a species of Mesenteripora.

Microeciella yoavi[32]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

An oncousoeciid cyclostome bryozoan, a species of Microeciella.

Micropora chathamica[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A microporid cheilostome bryozoan, a species of Micropora.

Monticulipora globulata[23]

Sp. nov

Valid

Jiménez-Sánchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Monticulipora.

Monticulipora irregularis[23]

Sp. nov

Valid

Jiménez-Sánchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Monticulipora.

Moyanopora[20]

Gen. et sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An arachnopusiid cheilostome bryozoan. The type species is Moyanopora hugoi.

Multescharellina pisiformis[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lepraliellid cheilostome bryozoan, a species of Multescharellina.

Myriapora sciutoi[35]

Sp. nov

Valid

Rosso & Di Martino

Miocene (Tortonian and Messinian) to Pleistocene (Gelasian)

 Italy
 Morocco

A member of Neocheilostomina belonging to the family Myriaporidae; a species of Myriapora.

Nematopora rara[34]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Nematopora.

Nikiforovastylus[29]

Gen. et sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A nikiforovellid cryptostome bryozoan. The type species is Nikiforovastylus ourthensis.

Nikiforovella dinantia[29]

Sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A nikiforovellid cryptostome bryozoan, a species of Nikiforovella.

Nikiforovella tobolensis[36]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Russia

Onychocella exilis[37]

Sp. nov

Valid

Koromyslova & Shcherbinina

Late Cretaceous (Maastrichtian)

 Uzbekistan

An onychocellid cheilostome bryozoan, a species of Onychocella.

Onychocella subtriangulata[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An onychocellid cheilostome bryozoan, a species of Onychocella.

Orthopora worsleyi[22]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

A rhabdomesid cryptostome bryozoan, a species of Orthopora.

Osthimosia aurora[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A celleporid cheilostome bryozoan, a species of Osthimosia.

Osthimosia curiosa[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A celleporid cheilostome bryozoan, a species of Osthimosia.

Oviexechonella[25]

Gen. et sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

An exechonellid arachnopusioid bryozoan. The type species is Oviexechonella digeronimoi.

Paracuneatopora egregia[34]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Paracuneatopora.

Paracuneatopora kuvaschensis[34]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Paracuneatopora.

Paranicklesopora ornaisa[28]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Parvohallopora cystata[24]

Sp. nov

Valid

Jiménez-Sánchez in Jiménez-Sánchez, Villas & Vennin

Ordovician (late Katian)

Khabt-el-Hajar Formation

 Morocco

A trepostomate bryozoan, a species of Parvohallopora.

Perkhurovella[38]

Gen. et sp. nov

Valid

Gorjunova

Carboniferous

 Russia

A diploporariine acanthocladiid bryozoan. The type species is Perkhurovella clara.

Phonicosia sinuosa[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lacernid cheilostome bryozoan, a species of Phonicosia.

Plagioecia zatoni[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A plagioeciid cyclostome bryozoan, a species of Plagioecia.

Platonea dilatata[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A tubuliporid cyclostome bryozoan, a species of Platonea.

Porella tiorioriensis[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A bryocryptellid cheilostome bryozoan, a species of Porella.

Porina turrita[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A porinid cheilostome bryozoan, a species of Porina.

Primorella variata[36]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Russia

Prophyllodictya simplex[39]

Sp. nov

Valid[40]

Ma et al.

Ordovician (early Tremadocian)

Nantzinkuan Formation

 China

A cryptostome bryozoan, a species of Prophyllodictya.

Pseudobatostomella parva[29]

Sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A dyscritellid trepostome bryozoan, a species of Pseudobatostomella.

Pseudothyracella campbelli[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A bryopastorid cheilostome bryozoan, a species of Pseudothyracella.

Puellina bontangensis[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A cribrilinid bryozoan, a species of Puellina.

Pyrisinella primazelandiae[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A pyrisinellid cheilostome bryozoan, a species of Pyrisinella.

Quasitrilaminopora[20]

Gen. et sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An arachnopusiid cheilostome bryozoan. The type species is Quasitrilaminopora curiosa.

Radiotrypa alnifensis[30]

Sp. nov

Valid

Ernst, Jiménez-Sánchez & Baidder

Ordovician (Katian)

 Morocco

A species of Radiotrypa.

Reptadeonella curvabilis[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

An adeonid bryozoan, a species of Reptadeonella.

Reptadeonella toddi[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian or early Langhian to Serravallian)

 Indonesia

An adeonid bryozoan, a species of Reptadeonella.

Reteporella mediocris[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A phidoloporid cheilostome bryozoan, a species of Reteporella.

Revalotrypa ramosa[30]

Sp. nov

Valid

Ernst, Jiménez-Sánchez & Baidder

Ordovician (Katian)

 Morocco

A species of Revalotrypa.

Saevitella renemai[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A hippopodinid schizoporelloid bryozoan, a species of Saevitella.

Saffordotaxis spinigerus[28]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Salairella[34]

Gen. et 5 sp. nov

Junior homonym

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan. Genus contains five species: S. petaloida, S. variabilis, S. maculata, S. baskuskanensis and S. recta. The generic name is a junior homonym of Salairella Khalfina (1961) and Salairella Severgina (1984).

Schizoretepora tamagawensis[41]

Sp. nov

Valid

Zágoršek, Takashima & Hirose

Miocene

Tanosawa Formation

 Japan

A member of Phidoloporidae, a species of Schizoretepora.

Sendinopora[25]

Gen. et sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

A tessaradomid lepralielloid bryozoan. The type species is Sendinopora prima.

Siphonicytara litotes[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A siphonicytarid cheilostome bryozoan, a species of Siphonicytara.

Siphonicytara primitiva[20]

Sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A siphonicytarid cheilostome bryozoan, a species of Siphonicytara.

Spinofenestella nodosa[28]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Stellatotrypa[19]

Gen. et sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A trepostome bryozoan. The type species is Stellatotrypa hirsuta.

Sulcoretepora regularis[36]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Russia

Terepennipora[38]

Gen. et comb. nov

Valid

Gorjunova

Carboniferous

 Russia

A diploporariine acanthocladiid bryozoan. A new genus for "Pinnatopora" longicellata Morozova (1955); genus also contains "Pinnatopora" volgensis Shishova (1959), "Pinnatopora" inconstans Shishova (1959), "Pinnatopora" subpunctata Shishova (1959) and "Pinnatopora" distincta Shishova (1950).

Trematopora maculata[22]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

A trematoporid trepostome bryozoan, a species of Trematopora.

Triznotrypa potii[33]

Sp. nov

Valid

Ernst, Tolokonnikova & Denayer

Devonian (late Frasnian)

 Belgium

A species of Triznotrypa.

Trypostega hasibuani[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A trypostegid hippothooid bryozoan, a species of Trypostega.

Tubiporella magnipora[25]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A didymosellid didymoselloid bryozoan, a species of Tubiporella.

Tumaiella[20]

Gen. et sp. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A calloporid cheilostome bryozoan. The type species is Tumaiella dieffenbachi.

Volgia deftera[26]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Hexagonellidae; a species of Volgia.

Zagorsekia[20]

Gen. et comb. nov

Valid[21]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 Australia
 New Zealand

A tubuliporid cyclostome bryozoan; a new genus for "Heteropora" nodulosa MacGillivray (1895).

Brachiopods

Name Novelty Status Authors Age Unit Location Notes Images

Alaskothyris[42]

Gen. et sp. nov

Valid

Blodgett, Baranov & Santucci

Devonian (Givetian)

 United States

A stringocephalid terebratulid brachiopod. The type species is Alaskothyris frosti.

Albasphe[43]

Gen. et sp. nov

In press

Halamski et al.

Middle Triassic (Ladinian)

 Croatia

A member of Terebratulida. The type species is Albasphe albertimagni.

Alexenia? delepinei[44]

Sp. nov

Valid

Chacón & Winkler Prins

Carboniferous (Pennsylvanian)

 Spain

A member of Productidae, possibly a species of Alexenia.

Amictocracens? brocki[45]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, possibly a species of Amictocracens.

Aphelotreta khemangarensis[45]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, a species of Aphelotreta.

Bifida rara[46]

Sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae, a species of Bifida.

Bockelia canalis[47]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Strophomenida belonging to the superfamily Plectambonitoidea and the family Taffidae; a species of Bockelia.

Chilcorthis[47]

Gen. et sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Orthida belonging to the superfamily Plectorthoidea. The type species is Chilcorthis huarpe.

Chonetinetes elongatus[48]

Sp. nov

Valid

Afanasjeva, Jun-ichi & Tomohiko

Permian

 Japan

A member of Chonetida belonging to the family Rugosochonetidae, a species of Chonetinetes.

Copperella[46]

Gen. et sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae. The type species is Copperella kaskevichi.

Cyrtonotreta vasculata[47]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Acrotretida belonging to the family Acrotretidae; a species of Cyrtonotreta.

Dagyssia[49]

Gen. et comb. nov

Valid

Gaetani & Mantovani

Triassic (Anisian)

 Croatia
 Hungary
 Italy
 China?

A mentzeliine mentzeliid brachiopod. The type species is "Spiriferina" palaeotypus Loretz (1875); genus might also contain "Mentzelia" multicostata Yang & Xu (1966).

Duartea latoresensis[44]

Sp. nov

Valid

Chacón & Winkler Prins

Carboniferous (Pennsylvanian)

 Spain

A member of Productidae, possibly a species of Duartea.

Eodmitria sezgini[50]

Sp. nov

Valid

Gourvennec & Hoşgör

Devonian (latest Givetian to early Frasnian)

Gümüșali Formation

 Turkey

A member of Spiriferida belonging to the family Cyrtospiriferidae, a species of Eodmitria.

Eohadrotreta haydeni[45]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, a species of Eohadrotreta.

Evanidisinurostrum saouraense[51]

Sp. nov

Valid

Mottequin et al.

Devonian (Famennian)

 Algeria

A rhynchonellid brachiopod, a species of Evanidisinurostrum.

Hadrotreta timchristiorum[45]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, a species of Hadrotreta.

Hibernodonta lakhensis[52]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Katian)

Chahgonbad Formation

 Iran

A rafinesquinid strophomenid brachiopod; a species of Hibernodonta.

Hindella prima[52]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Katian)

Chahgonbad Formation

 Iran

A meristellid athyridid brachiopod; a species of Hindella.

Inflatia ovetensis[44]

Sp. nov

Valid

Chacón & Winkler Prins

Carboniferous (Pennsylvanian)

 Spain

A member of Productidae, possibly a species of Inflatia.

Jafarirhynchus[53]

Gen. et sp. nov

Valid

Popov et al.

Silurian (Telychian)

Niur Formation

 Iran

A rhynchotrematoid rhynchonellid brachiopod. The type species is Jafarirhynchus alatus.

Kerberellus[54]

Gen. et sp. nov

Valid

Devaere, Holmer & Clausen in Devaere et al.

Cambrian (Terreneuvian)

Marcou Formation

 France

A mickwitziid-like stem-group brachiopod. The type species is Kerberellus marcouensis.

Kirkirhynchus[55]

Gen. et comb. nov

Valid

Baranov & Blodgett

Devonian (early Pragian)

 United States

A trigonirhynchiid rhynchonellid brachiopod; a new genus for "Camarotoechia" reesidei Kirk & Amsden (1952).

Ktenopotamorhynchus[56]

Gen. et comb. nov

Valid

Sartenaer

Devonian (Lochkovian)

 Czech Republic
 Ukraine

A member of Rhynchonellida. The type species is "Rhynchonella" borealiformis Siemiradzki (1906); genus also includes "Plethorhyncha" altera (Barrande, 1879).

Kutorgina songshanensis[57]

Sp. nov

Valid

Liu et al.

Cambrian

 China

A species of Kutorgina.

Lingularia salymica[58]

Sp. nov

Valid

Smirnova in Smirnova et al.

Late Jurassic

 Russia

A member of Lingulida, a species of Lingularia.

Lomatorthis? multilamellosa[52]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Darriwilian)

Chahgonbad Formation

 Iran

A glyptorthid orthid brachiopod; possibly a species of Lomatorthis.

Misensia[46]

Gen. et sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae. The type species is Misensia gracilis.

Ogilviella gerensis[46]

Sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae, a species of Ogilviella.

Paillettia menae[50]

Sp. nov

Valid

Gourvennec & Hoşgör

Devonian (latest Givetian to early Frasnian)

Gümüșali Formation

 Turkey

A member of Spiriferida belonging to the family Adolfiidae, a species of Paillettia.

Palaeoglossa minima[47]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Obolidae; a species of Palaeoglossa.

Pelmanotreta[59]

Nom. nov

Valid

Skovsted et al.

Early Cambrian

 Russia

A replacement name for Cryptotreta Pelman (1977) (preoccupied).

Philhedra pauciradiata[47]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Craniidae; a species of Philhedra.

Portrania[60]

Gen. et sp. nov

Valid[61]

Cocks

Ordovician (late Katian)

Portrane Limestone

 Ireland

A chilidiopsoidean brachiopod. The type species is Portrania wrighti Cocks & Harper in Cocks (2015).

Prototreta? sumnaensis[45]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, possibly a species of Prototreta.

Ptychomentzelia[49]

Gen. et comb. et sp. nov

Valid

Gaetani & Mantovani

Middle Triassic

 Austria
 Bosnia and Herzegovina
 Hungary
 Italy
 Romania
 Turkey

A tethyspirine mentzeliid brachiopod. The type species is "Spiriferina" propontica Toula (1896); genus also contains "Spiriferina" ptychitiphila Bittner (1890), as well as the new species Ptychomentzelia ritensis.

Raisiya[46]

Gen. et sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae. The type species is Raisiya solonchanensis.

Rugaria semicircularis[48]

Sp. nov

Valid

Afanasjeva, Jun-ichi & Tomohiko

Permian

 Japan

A member of Chonetida belonging to the family Rugosochonetidae, a species of Rugaria.

Savagerhynchus[55]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (early Pragian)

 United States

A trigonirhynchiid rhynchonellid brachiopod. The type species is Savagerhynchus hecetaensis.

Scalenomena[60]

Gen. et comb. nov

Valid[61]

Cocks

Ordovician (early Katian

Craighead Limestone

 United Kingdom

A chilidiopsoidean brachiopod. A new genus for "Fardenia" scalena Williams (1962).

Sillakkudirhynchia[62]

Gen. et comb. nov

Valid

Radulović et al.

Late Cretaceous (Coniacian to early Maastrichtian)

Sillakkudi Formation

 India

A member of Rhynchonellida belonging to the family Tetrarhynchiidae. A new genus for "Rhynchonella" plicatiloides Stoliczka (1872); genus also contains "Cretirhynchia" sahnii Radulović & Ramamoorthy (1992).

Sivorthis precordillerana[47]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation
San Juan Formation

 Argentina

A member of Orthida belonging to the family Orthidae; a species of Sivorthis.

Talexirhynchia[63]

Gen. et sp. nov

Valid

Feldman et al.

Middle Jurassic (Callovian)

Mughanniyya Formation

 Jordan

A rhynchonellid brachiopod. The type species is Talexirhynchia kadishi.

Thecocyrtella dagysii[43]

Sp. nov

In press

Halamski et al.

Middle Triassic (Ladinian)

 Croatia

A member of Spiriferida; a species of Thecocyrtella.

Theodossia kagarmanovi[64]

Sp. nov

Valid

Bezgodova

Devonian (Frasnian)

 Russia

A member of Spiriferida belonging to the family Theodossiidae, a species of Theodossia.

Theodossia novosemelica[64]

Sp. nov

Valid

Bezgodova

Devonian (Frasnian)

 Russia

A member of Spiriferida belonging to the family Theodossiidae, a species of Theodossia.

Theodossia tcherkesovae[64]

Sp. nov

Valid

Bezgodova

Devonian (Frasnian)

 Russia

A member of Spiriferida belonging to the family Theodossiidae, a species of Theodossia.

Yangtzeella chupananica[52]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Darriwilian)

Chahgonbad Formation

 Iran

A clarkellid pentamerid brachiopod; a species of Yangtzeella.

Molluscs

Echinoderms

Name Novelty Status Authors Age Unit Location Notes Images

Aegyptiaris[65]

Gen. et sp. et comb. nov

Valid

Abdelhamid & Abdelghany

Cretaceous (Aptian to Cenomanian)

 Egypt

An emiratiid sea urchin. The type species is Aegyptiaris halalensis;[65] genus also contains "Trochodiadema" isidis Fourtau (1921).[66]

Aganaster jagiellonicus[67]

Sp. nov

Valid

Thuy, Kutscher & Płachno

Carboniferous (late Tournaisian)

Mazurowe Doły Formation

 Poland

A brittle star belonging to the suborder Ophiurina, a species of Aganaster.

Archaeocidaris marmorcataractensis[68]

Sp. nov

Valid

Thompson et al.

Carboniferous (early Pennsylvanian)

Marble Falls Formation

 United States

A sea urchin, a species of Archaeocidaris.

Baficrinus[69]

Gen. et sp. nov

Valid

Prokop & Nohejlová

Devonian (late Emsian)

Daleje-Třebotov Formation

 Czech Republic

A crinoid belonging to the group Disparida and the family Zophocrinidae. The type species is Baficrinus vigilis.

Baudicrinus[70]

Gen. et sp. nov

Valid

Oji & Twitchett

Early Triassic

AI Jil Formation

 Oman

A dadocrinid articulate crinoid, a relative of encrinids. The type species is Baudicrinus krystyni.

Bizarroglobus[71]

Gen. et sp. nov

Valid

Sumrall & Sprinkle

Ordovician

Kanosh Shale

 United States

An edrioasteroid. The type species is Bizarroglobus medusae.

Browerocrinus[72]

Gen. et sp. nov

Valid

Ausich, Peter & Ettensohn

Silurian (Rhuddanian)

Brassfield Formation

 United States

A crinoid. The type species is Browerocrinus arthrikos.

Callocystites fresti[73]

Sp. nov

Valid

Paul

Silurian

Massie Formation

 United States

A callocystitid cystoid, a species of Callocystites.

‘Cidaris’ labergensis[74]

Sp. nov

Valid[75]

Zonneveld, Furlong & Sanders

Late Triassic

Aksala Formation

 Canada

A sea urchin. Zonneveld, Furlong & Sanders (2015) assign it to the form genus ‘Cidaris’, stressing that it does not imply affinity with the extant genus Cidaris.[74]

Conollia sporranoides[76]

Sp. nov

Valid

Rahman, Stewart & Zamora

Ordovician (Sandbian)

Balclatchie Formation

 United Kingdom

A member of Ctenocystoidea, a species of Conollia.

Ekmelocrinus[77]

Gen. et comb. nov

Valid

Webster & Donovan

Permian

 Indonesia

A crinoid. Genus includes Ekmelocrinus amplior (Wanner, 1924), Ekmelocrinus subamplior (Wanner, 1949), Ekmelocrinus ovoides (Wanner, 1949), Ekmelocrinus verbeeki (Wanner, 1916) and Ekmelocrinus vermistriatus (Wanner, 1916).

Enakomusium[78]

Gen. et comb. nov

Valid

Thuy

Jurassic (Toarcian to Oxfordian)

 France
 Germany
  Switzerland
 United Kingdom

A brittle star. A new genus for "Ophioderma" weymouthiense Damon (1880); genus also contains "Ophiomusium" ferrugineum Boehm (1889), "Ophiolepis" leckenbyi Wright (1880), "Ophiomusium" geisingense Kutscher (1992), "Ophiomusium" mammillatum Hess (1966) and "Ophiura" gagnebini Thurmann (1851).

Eotiaris guadalupensis[79]

Sp. nov

Valid

Thompson in Thompson et al.

Permian (Roadian-Capitanian)

Bell Canyon Formation
Road Canyon Formation
Word Formation

 United States

A cidaroid sea urchin, a species of Eotiaris.

Exaesiodiscus derbyi[80]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Exaesiodiscus.

Frasericrinus[81]

Gen. et sp. nov

Valid

Ausich et al.

Early Cretaceous (probably early late Albian)

Peace River Formation

 Canada

An isocrinid crinoid. The type species is Frasericrinus mauricensis.

Globoeocrinus zhaoyuanlongensis[82]

Sp. nov

Valid

Yang et al.

Cambrian

Balang Formation

 China

An eocrinoid, a species of Globoeocrinus.

Guangxicrinus[83]

Gen. et sp. nov

Valid

Webster & Chen

Devonian (Eifelian)

Yingtang Formation

 China

A marsupiocrinid crinoid. The type species is Guangxicrinus xiangzhouensis.

Hemidiadema batalleri[84]

Sp. nov

Valid

Clément

Early Cretaceous (Aptian)

 Spain

A sea urchin belonging to the family Glyphocyphidae, a species of Hemidiadema.

Hemidiadema forcallensis[84]

Sp. nov

Valid

Clément

Early Cretaceous (Aptian)

 Spain

A sea urchin belonging to the family Glyphocyphidae, a species of Hemidiadema.

Hemidiadema malladai[84]

Sp. nov

Valid

Clément

Early Cretaceous (Aptian)

 Spain

A sea urchin belonging to the family Glyphocyphidae, a species of Hemidiadema.

Heviacrinus tromelini[85]

Sp. nov

Valid

Lefebvre et al.

Ordovician (late Darriwilian)

 France

A crinoid, a species of Heviacrinus.

Iocrinus africanus[86]

Sp. nov

Valid

Zamora, Rahman & Ausich

Ordovician (Darriwilian)

Taddrist Formation

 Morocco

An iocrinid crinoid, a species of Iocrinus.

Lanxocolumnus (col.) chaleurensis[87]

Sp. nov

Valid

Donovan & Keighley

Silurian (Telychian)

 Canada

A crinoid, a species of Lanxocolumnus.

Lapidaster hellersi[88]

Sp. nov

Valid

Numberger-Thuy & Thuy

Middle Jurassic (earliest Bajocian)

 Luxembourg

A brittle star belonging to the family Ophiacanthidae, a species of Lapidaster.

Laudonomphalus ferreirai[80]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Laudonomphalus.

Llanocystis[89]

Gen. et sp. nov

Valid

Zamora, Sumrall & Sprinkle

Cambrian (Furongian)

Wilberns Formation

 United States

An eocrinoid of uncertain phylogenetic placement. The type species is Llanocystis wilbernsensis.

Lovenicystis kopfi[90]

Sp. nov

Valid

Paul

Silurian

 United States

A callocystitid cystoid, a species of Lovenicystis.

Marettocrinus hartti[80]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Marettocrinus.

Marettocrinus katzeri[80]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Marettocrinus.

Marginix[91]

Nom. nov

Valid

Martínez & del Río

Devonian

Talacasto Formation

 Argentina

A brittle star belonging to the order Oegophiurida and the family Encrinasteridae; a replacement name for Marginaster Haude (1995) and Marginura Haude (1999) (both preoccupied).

Notatudiadema[65]

Gen. et sp. nov

Valid

Abdelhamid & Abdelghany

Late Cretaceous (Cenomanian)

 Egypt

A diadematoid sea urchin. The type species is Notatudiadema rekeibensis.

Ophiocantabria[92]

Gen. et sp. nov

Valid

Blake, Zamora & García-Alcalde

Devonian (Lochkovian)

Furada Formation

 Spain

A brittle star belonging to the order Oegophiurida, suborder Lysophiurina and the family Encrinasteridae. The type species is Ophiocantabria elegans.

Ophiodoris reconciliator[78]

Sp. nov

Valid

Thuy

Middle Jurassic (early Bajocian)

Longwy Formation

 France

A brittle star belonging to the family Ophionereididae, a species of Ophiodoris.

Ophiotitanos aschmannicor[78]

Sp. nov

Valid

Thuy

Middle Jurassic (early Bajocian)

Longwy Formation

 France

A brittle star belonging to the family Ophiodermatidae, a species of Ophiotitanos.

Paracidaris (Anisocidaris) eluveitie[93]

Sp. nov

Valid

Hostettler & Menkveld-Gfeller

Late Jurassic (Oxfordian)

Wildegg Formation

  Switzerland

A member of Cidaroida, a species of Paracidaris.

Pentaridica mendesi[80]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Pentaridica.

Polycidaris gygii[93]

Sp. nov

Valid

Hostettler & Menkveld-Gfeller

Late Jurassic (Oxfordian)

Lochen Formation
Wildegg Formation

 Germany
  Switzerland

A member of Cidaroida, a species of Polycidaris.

Porosublastus[94]

Gen. et sp. nov

Valid

Sprinkle & Sumrall

Early Ordovician

Ninemile Shale

 United States

An edrioasteroid. The type species is Porosublastus inexpectus.

Pseudedriophus[94]

Gen. et sp. nov

Valid

Sprinkle & Sumrall

Early Ordovician

Ninemile Shale

 United States

An edrioasteroid. The type species is Pseudedriophus guensburgi.

Quechuacrinus[95]

Gen. et sp. nov

In press

Guensburg & Waisfeld

Ordovician (Floian)

 Argentina

A crinoid. The type species is Quechuacrinus ticsa.

Ramseyocrinus argentinus[95]

Sp. nov

In press

Guensburg & Waisfeld

Ordovician (Floian)

 Argentina

A crinoid, a species of Ramseyocrinus.

Rautscholdticrinus[77]

Gen. et comb. nov

Valid

Webster & Donovan

Permian

 Indonesia

A crinoid. Genus includes Rautscholdticrinus indicus (Wanner, 1916) and Rautscholdticrinus weidnerii (Wanner, 1937).

Rhabdocidaris deubelbeissi[93]

Sp. nov

Valid

Hostettler & Menkveld-Gfeller

Late Jurassic (Oxfordian)

Wildegg Formation

  Switzerland

A member of Cidaroida, a species of Rhabdocidaris.

Ristnacrinus oepiki[96]

Sp. nov

Valid

Ausich, Rozhnov & Kammer

Ordovician (Katian)

 Estonia

A crinoid related to Iocrinus, a species of Ristnacrinus.

Salamonicrinus[97]

Gen. et sp. nov

Valid

Trzęsiok

Late Jurassic (Tithonian)

 Poland

A cyrtocrinid crinoid. The type species is Salamonicrinus prodigiosum.

Salirocystis[90]

Gen. et comb. nov

Valid

Paul

Silurian

 United States

A callocystitid cystoid; a new genus for "Apiocystites" anna Safford (1869).

Temnocrinus americanus[72]

Sp. nov

Valid

Ausich, Peter & Ettensohn

Silurian (Rhuddanian)

Brassfield Formation

 United States

A crinoid, a species of Temnocrinus.

Velocrinus[98]

Gen. et sp. nov

Valid

Ausich, Wilson & Vinn

Late Silurian

Kaugatuma Formation

 Estonia

A crotalocrinitid crinoid. The type species is Velocrinus coniculus.

Conodonts

Name Novelty Status Authors Age Unit Location Notes Images

Arianagnathus[99]

Gen. et sp. nov

Valid

Männik, Miller & Hairapetian

Silurian

Niur Formation

 Iran

A member of Prioniodontida, possibly a balognathid. The type species is Arianagnathus jafariani.

Eolinguipolygnathus radula[100]

Sp. nov

Valid

Aboussalam & Becker in Aboussalam, Becker & Bultynck

Devonian (early Emsian)

 Morocco

Gapparodus gapparites[101]

Sp. nov

Valid

Yang, Steiner & Keupp

Early Cambrian

Shuijingtuo Formation

 China

A member of Furnishinidae; a species of Gapparodus.

Hayashiella[102]

Gen. et comb. nov

Valid

Kiliç et al.

Late Triassic

 Canada
 Italy
 Japan
 Slovakia

A member of Gondolellidae. A new genus for "Metapolygnathus" nodosus Hayashi (1968); genus also includes "Gondolella" carpathica Mock (1979), H. lindae (Orchard) and H. permica (Hayashi).

Icriodus ovalis[100]

Sp. nov

Valid

Bultynck in Aboussalam, Becker & Bultynck

Devonian (late Emsian)

Mdâouer-el-Kbîr Formation

 Morocco

Icriodus praerectirostratus[100]

Sp. nov

Valid

Bultynck in Aboussalam, Becker & Bultynck

Devonian (late Emsian)

Timrhanrhart Formation

 Morocco

Mazzaella[102]

Gen. et comb. nov

Valid

Kiliç et al.

Late Triassic

 Germany
 Hungary
 Italy
 Turkey

A member of Gondolellidae. A new genus for "Epigondolella" carnica Krystyn (1975); genus also includes "Metapolygnathus" baloghi Kovacs (1977).

Moskalenkodus[103]

Gen. et comb. nov

Valid

Dzik

Late Ordovician

 Russia

A new genus for "Amorphognathus" quinquiradiatus Moskalenko in Kanygin et al. (1977). Genus also includes Moskalenkodus cruciformis (Moskalenko, 1970).

Neostreptognathodus costatus[104]

Sp. nov

Valid

Wang & Zhang

Early Permian

Amushan Formation

 China

Ozarkodina derenjalensis[99]

Sp. nov

Valid

Männik, Miller & Hairapetian

Silurian

Niur Formation

 Iran

A member of Ozarkodinida, a spathognathodontid; a species of Ozarkodina.

Polygnathus bicristatus[105]

Sp. nov

Valid

Mossoni et al.

Late Devonian or Early Carboniferous

 Italy

A member of Ozarkodinida belonging to the family Polygnathidae, a species of Polygnathus.

Polygnathus nuragicus[105]

Sp. nov

Valid

Mossoni et al.

Late Devonian or Early Carboniferous

 Italy

A member of Ozarkodinida belonging to the family Polygnathidae, a species of Polygnathus.

Pseudopolygnathus granulobatus[105]

Sp. nov

Valid

Mossoni et al.

Late Devonian or Early Carboniferous

 Italy

A member of Ozarkodinida belonging to the family Polygnathidae, a species of Pseudopolygnathus.

Spathogondolella[106]

Gen. et sp. nov

Valid

Jiang in Chen et al.

Early Triassic

 China

The type species is Spathogondolella jiarongensis.

Vjalovognathus nicolli[107]

Sp. nov

Valid[108]

Yuan, Shen & Henderson in Yuan et al.

Early Permian

Xiala Formation

 China

A species of Vjalovognathus.

Fishes

Amphibians

Research

New taxa

Temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Eocyclotosaurus appetolatus[112]

Sp. nov

Valid

Rinehart, Lucas & Schoch

Middle Triassic

Moenkopi Formation

 United States

A capitosaur, a species of Eocyclotosaurus.

Metoposaurus algarvensis[113]

Sp. nov

Valid

Brusatte et al.

Late Triassic

 Portugal

A species of Metoposaurus.

Parapytanga[114]

Gen. et sp. nov

Valid

Strapasson, Pinheiro & Soares

Middle or late Permian

Rio do Rasto Formation

 Brazil

A basal member of Stereospondylomorpha. The type species is Parapytanga catarinensis.

Procuhy[115]

Gen. et sp. nov

Valid

Cisneros et al.

Permian (Cisuralian)

Lower Pedra de Fogo Formation

 Brazil

A member of Trimerorhachidae. The type species is Procuhy nazariensis.

Timonya[115]

Gen. et sp. nov

Valid

Cisneros et al.

Permian (Cisuralian)

Lower Pedra de Fogo Formation

 Brazil

A member of Dvinosauria. The type species is Timonya anneae.

Lepospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Aletrimyti[116]

Gen. et sp. nov

Valid

Szostakiwskyj, Pardo & Anderson

Permian (Kungurian)

Fairmont Shale

 United States

A member of Recumbirostra. The type species is Aletrimyti gaskillae.

Batropetes appelensis[117]

Sp. nov

Valid

Glienke

Early Permian

Saar–Nahe Basin

 Germany

A species of Batropetes.

Batropetes palatinus[117]

Sp. nov

Valid

Glienke

Early Permian

Saar–Nahe Basin

 Germany

A species of Batropetes.

Dvellecanus[116]

Gen. et sp. nov

Valid

Szostakiwskyj, Pardo & Anderson

Permian (Kungurian)

Fairmont Shale

 United States

A member of Recumbirostra. The type species is Dvellecanus carrolli.

Lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Bufo mirus[118]

Sp. nov

Valid

Syromyatnikova

Late Early – early Middle Miocene

Khalagay Formation

 Russia

A true toad of uncertain phylogenetic placement, most similar and possibly related to the Mongolian toad.

Hensonbatrachus[119]

Gen. et sp. nov

Valid

Gardner & Brinkman

Late Cretaceous (Campanian)

Dinosaur Park Formation
Oldman Formation

 Canada

A frog. The type species is Hensonbatrachus kermiti.

Kiyatriton krasnolutskii[120]

Sp. nov

Valid

Skutschas

Middle Jurassic (Bathonian)

Itat Formation

 Russia

A salamander, a species of Kiyatriton.

Lepidobatrachus australis[121]

Sp. nov

Valid

Nicoli

Late Miocene to Early Pliocene

Monte Hermoso Formation

 Argentina

A frog belonging to the family Ceratophryidae, a species of Lepidobatrachus.

Palaeoplethodon[122]

Gen. et sp. nov

Valid

Poinar & Wake

Eocene or Miocene

 Dominican Republic

A lungless salamander found in Dominican amber. The type species is Palaeoplethodon hispaniolae.

Tyrrellbatrachus[123]

Gen. et sp. nov

Valid

Gardner

Late Cretaceous (Campanian)

Dinosaur Park Formation

 Canada

A frog of uncertain phylogenetic placement. The type species is Tyrrellbatrachus brinkmani.

Varibatrachus[124]

Gen. et sp. nov

Valid

Parmley, Chandler & Chandler

Miocene (late Clarendonian)

Ash Hollow Formation

 United States

A frog of uncertain phylogenetic placement. The type species is Varibatrachus abraczinskasae.

Diadectomorphs

Name Novelty Status Authors Age Unit Location Notes Images

Alveusdectes[125]

Gen. et sp. nov

Valid

Liu & Bever

Late Permian (Wuchiapingian, roughly 256 Ma)

Shangshihezi Formation

 China

A diadectid. The type species is Alveusdectes fenestralis.

Ichthyosauromorphs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Cartorhynchus[128]

Gen. et sp. nov

Valid

Motani et al.

Early Triassic (Olenekian)

Nanlinghu Formation

 China

A relative of ichthyopterygians. The type species is Cartorhynchus lenticarpus.

Dearcmhara[129]

Gen. et sp. nov

Valid

Brusatte et al.

Jurassic (Toarcian–Bajocian)

 United Kingdom

A basal member of Neoichthyosauria. The type species is Dearcmhara shawcrossi.

Eretmorhipis[130]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic (late Spathian)

Jialingjiang Formation

 China

A hupehsuchian. The type species is Eretmorhipis carrolldongi.

Ichthyosaurus anningae[131]

Sp. nov

Valid

Lomax & Massare

Early Jurassic (Hettangian/Sinemurian–Pliensbachian)

 United Kingdom

A species of Ichthyosaurus.

Muiscasaurus[132]

Gen. et sp. nov

Valid[133]

Maxwell et al.

Early Cretaceous (Barremian–Aptian)

Paja Formation

 Colombia

A member of Ophthalmosauridae. The type species is Muiscasaurus catheti.

Sauropterygians

Name Novelty Status Authors Age Unit Location Notes Images

Anguanax[134]

Gen. et sp. nov

Valid[135]

Cau & Fanti

Late Jurassic (Oxfordian)

Rosso Ammonitico Veronese Formation

 Italy

A pliosaurid. The type species is Anguanax zignoi.

Atychodracon[136]

Gen. et comb. nov

Valid

Smith

Early Jurassic

 United Kingdom

A rhomaleosaurid plesiosaur; a new genus for "Plesiosaurus" megacephalus Stutchbury (1846).

Cardiocorax[137]

Gen. et sp. nov

Valid

Araújo et al.

Late Cretaceous (early Maastrichtian)

Mocuio Formation

 Angola

An elasmosaurid plesiosaur. The type species is Cardiocorax mukulu.

Dianmeisaurus[138]

Gen. et sp. nov

Valid

Shang & Li

Middle Triassic (Anisian)

Guangling Formation

 China

A member of Eosauropterygia of uncertain phylogenetic placement, more closely related to nothosaurs than to plesiosaurs. The type species is Dianmeisaurus gracilis.

Makhaira[139]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (Hauterivian)

 Russia

A pliosaurid. The type species is Makhaira rossica.

Vegasaurus[140]

Gen. et sp. nov

Valid

O’Gorman et al.

Late Cretaceous (early Maastrichtian)

Snow Hill Island Formation

Antarctica (Vega Island)

An elasmosaurid plesiosaur. The type species is Vegasaurus molyi.

Wangosaurus[141]

Gen. et sp. nov

Valid

Ma et al.

Middle Triassic (Ladinian)

Falang Formation

 China

A pistosauroid. The type species is Wangosaurus brevirostris.

Lepidosaurs

Research

New taxa

Rhynchocephalians

Name Novelty Status Authors Age Unit Location Notes Images

Clevosaurus sectumsemper[144]

Sp. nov

Valid

Klein et al.

Late Triassic (Rhaetian)

 United Kingdom

A clevosaurid sphenodontian, a species of Clevosaurus.

Lizards

Name Novelty Status Authors Age Unit Location Notes Images

Archaerhineura[145]

Gen. et sp. nov

Valid

Longrich et al.

Late Palaeocene

Polecat Bench Formation

 United States

A rhineurid amphisbaenian. The type species is Archaerhineura mephitis.

Asagaolacerta[146]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

Kuwajima Formation

 Japan

A lizard of uncertain phylogenetic placement. The type species is Asagaolacerta tricuspidens.

Asprosaurus[147]

Gen. et sp. nov

Valid

Park, Evans & Huh

Late Cretaceous

 South Korea

A member of Anguimorpha, probably a member of Monstersauria. The type species is Asprosaurus bibongriensis.

Babibasiliscus[148]

Gen. et sp. nov

Valid

Conrad

Eocene (approximately 48 Ma)

Bridger Formation

 United States

A member of Corytophanidae. The type species is Babibasiliscus alxi.

Barbatteius[149]

Gen. et sp. nov.

Valid[150]

Venczel & Codrea

Late Cretaceous (early Maastrichtian)

Haţeg Basin

 Romania

A member of Teiidae. The type species is Barbatteius vremiri.

Cadurcogekko verus[151]

Sp. nov

Valid

Bolet et al.

Eocene

 France

A gekkotan lizard, a species of Cadurcogekko.

Chromatogenys[152]

Gen. et sp. nov

Valid

Makádi & Nydam

Late Cretaceous (Santonian)

 Hungary

A member of Scincomorpha of uncertain phylogenetic placement. The type species is Chromatogenys tiliquoides.

Chthonophis[145]

Gen. et sp. nov

Valid

Longrich et al.

Probably early Palaeocene

Bug Creek Anthills, Fort Union Formation

 United States

An amphisbaenian, the only member of the new family Chthonophidae. The type species is Chthonophis subterraneus.

Cuvieribaena[153]

Gen. et sp. nov

Valid

Čerňanský, Augéc & Rage

Eocene (Bartonian)

 France

A blanid amphisbaenian. The type species is Cuvieribaena carlgansi.

Dryadissector[154]

Gen. et sp. nov

Valid

Wick, Lehman & Brink

Late Cretaceous (early Campanian)

Aguja Formation

 United States

A member of Varanoidea. The type species is Dryadissector shilleri.

Gekkomimus[151]

Gen. et comb. nov

Valid

Bolet et al.

Eocene

 France

A skink; a new genus for "Cadurcogekko" rugosus Augé (2005).

Gueragama[155]

Gen. et sp. nov

Valid

Simões et al.

Late Cretaceous (Turonian-Campanian)

Goio-Erê Formation

 Brazil

The first member of Acrodonta (a relative of agamids and chameleons) described from South America. The type species is Gueragama sulamericana.

Hakuseps[146]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

 Japan

A squamate of uncertain phylogenetic placement. The type species is Hakuseps imberis.

Kuroyuriella[146]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

Kuwajima Formation

 Japan

A lizard of uncertain phylogenetic placement. The type species is Kuroyuriella mikikoi.

Norellius[156]

Gen. et sp. nov

Valid

Conrad & Daza

Early Cretaceous (probably approximately 130 Mya)

Öösh Basin

 Mongolia

A member of Gekkonomorpha of uncertain phylogenetic placement. The type species is Norellius nyctisaurops.

Ophisaurus holeci[157]

Sp. nov

Valid

Klembara

Early Miocene

 Czech Republic

A glass lizard.

Pachygenys adachii[158]

Sp. nov

Valid

Ikeda, Ota & Saegusa

Early Cretaceous

Sasayama Group

 Japan

A squamate of uncertain phylogenetic placement, a species of Pachygenys.

Phosphorosaurus ponpetelegans[159]

Sp. nov

Valid[160]

Konishi et al.

Late Cretaceous (Maastrichtian)

 Japan

A halisaurine mosasaur, a species of Phosphorosaurus.

Plioplatecarpus peckensis[161]

Sp. nov

Valid

Cuthbertson & Holmes

Late Cretaceous (Campanian)

Bearpaw Formation

 United States

A mosasaur, a species of Plioplatecarpus.

Pseudopus rugosus[157]

Sp. nov

Valid

Klembara

Early Miocene

 Czech Republic

A relative of the sheltopusik.

Snakes

Name Novelty Status Authors Age Unit Location Notes Images

Diablophis[162]

Gen. et comb. nov

Valid

Caldwell et al.

Late Jurassic

Morrison Formation

 United States

A basal snake; a new genus for "Parviraptor" gilmorei Evans (1996).

Eophis[162]

Gen. et sp. nov

Valid

Caldwell et al.

Middle Jurassic (Bathonian)

Forest Marble

 United Kingdom

A basal snake. The type species is Eophis underwoodi.

Portugalophis[162]

Gen. et sp. nov

Valid

Caldwell et al.

Late Jurassic (Kimmeridgian)

Camadas de Guimarota

 Portugal

A basal snake. The type species is Portugalophis lignites.

Renenutet[163]

Gen. et sp. nov

Valid

Mccartney & Seiffert

Eocene (Priabonian)

 Egypt

A member of Colubroidea. The type species is Renenutet enmerwer.

Tetrapodophis[164]

Gen. et sp. nov

Valid

Martill, Tischlinger & Longrich

Early Cretaceous (Aptian)

Crato Formation

 Brazil

An early, four-legged snake. The type species is Tetrapodophis amplectus.

Turtles

Name Novelty Status Authors Age Unit Location Notes Images

Adocus sengokuensis[165]

Sp. nov

Valid

Sonoda et al.

Early Cretaceous

Sengoku Formation

 Japan

A species of Adocus.

Arvinachelys[166]

Gen. et sp. nov

Valid

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

 United States

A member of Baenidae. The type species is Arvinachelys goldeni.

Bairdemys thalassica[167]

Sp. nov

Valid

Ferreira et al.

Miocene

Capadare Formation

 Venezuela

A podocnemidid belonging to the clade Stereogenyina, a species of Bairdemys.

Corsochelys bentleyi[168]

Sp. nov

Valid

Schwimmer et al.

Late Cretaceous

 United States

A member of Dermochelyidae, a species of Corsochelys.

Desmatochelys padillai[169]

Sp. nov

Valid

Cadena & Parham

Early Cretaceous (late Barremian-early Aptian)

Paja Formation

 Colombia

A member of Protostegidae, species of Desmatochelys.

Gaffneylania[170]

Gen. et sp. nov

Valid

Sterli, de la Fuente & Krause

Middle Eocene

Sarmiento Formation

 Argentina

A member of Meiolaniidae. The type species is Gaffneylania auricularis.

Gobiapalone palaeocenica[171]

Sp. nov

Valid

Danilov et al.

Paleocene

Bugin Tsav Basin

 Mongolia

A member of Trionychidae, a species of Gobiapalone.

Gomphochelys[172]

Gen. et sp. nov

Valid

Bourque et al.

Early Wasatchian

Willwood Formation

 United States

A member of Dermatemydidae. The type species is Gomphochelys nanus.

Judithemys russelli[173]

Sp. nov

Valid

Brinkman

Late Cretaceous

 Canada

A member of (likely non-monophyletic) Macrobaenidae, a species of Judithemys.

Jurassichelon[174]

Gen. et sp. nov

Valid

Pérez-García

Late Jurassic (early Tithonian)

 France

A basal member of Eucryptodira. The type species is Jurassichelon oleronensis.

Khunnuchelys lophorhothon[175]

Sp. nov

Valid

Danilov et al.

Late Cretaceous (Santonian or early Campanian)

Bostobe Formation

 Kazakhstan

A trionychine trionychid, a species of Khunnuchelys.

Kimurachelys[176]

Gen. et sp. nov

Valid

Brinkman et al.

Late Cretaceous (late Campanian)

Dinosaur Park Formation

 Canada

A chelonioid turtle. The type species is Kimurachelys slobodae.

Leyvachelys[177]

Gen. et sp. nov

Valid

Cadena

Early Cretaceous (Barremian-Albian)

Glen Rose Formation
Paja Formation

 Colombia
 United States

A member of Pan-Cryptodira belonging to the family Sandownidae. The type species is Leyvachelys cipadi.

Mauremys oshiroi[178]

Sp. nov.

Valid

Takahashi et al.

Late Pleistocene

Tomori Amaga Cave

 Japan

A pond turtle.

Neurankylus hutchisoni[179]

Sp. nov

Valid[180]

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

 United States

A member of Baenidae, a species of Neurankylus.

Neurankylus utahensis[179]

Sp. nov

Valid[180]

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

 United States

A member of Baenidae, a species of Neurankylus.

Perochelys[181]

Gen. et sp. nov.

Valid

Li, Joyce & Liu

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A soft-shelled turtle of uncertain phylogenetic placement; it might be a stem- or a crown-trionychid. The type species is Perochelys lamadongensis.

Phunoichelys[182]

Gen. et sp. nov.

Valid

Tong et al.

Late Jurassic

Phu Kradung Formation

 Thailand

A xinjiangchelyid eucryptodiran. The type species is Phunoichelys thirakhupti.

Portlandemys gracilis[183]

Sp. nov.

Valid

Anquetin, Püntener & Billon-Bruyat

Late Jurassic (late Kimmeridgian)

Reuchenette Formation

  Switzerland

A plesiochelyid eucryptodiran, a species of Portlandemys.

Sternotherus bonevalleyensis[184]

Sp. nov.

Valid

Bourque & Schubert

Late Hemphillian

 United States

A musk turtle.

Sternotherus palaeodorus[184]

Sp. nov.

Valid

Bourque & Schubert

Late Hemphillian

 United States

A musk turtle.

Taraschelon[185]

Gen. et comb. nov

Valid

Pérez-García

Early Oligocene

 France

A tortoise; a new genus for "Testudo" gigas Bravard (1844).

Testudo brevitesta[186]

Sp. nov

Valid[187]

Vlachos & Tsoukala

Late Pliocene (early Villafranchian)

 Greece

A tortoise, a species of Testudo.

Thalassemys bruntrutana[188]

Sp. nov

Valid

Püntener, Anquetin & Billon-Bruyat

Late Jurassic (late Kimmeridgian)

Reuchenette Formation

  Switzerland

A basal member of Pancryptodira, a species of Thalassemys.

Toremys[189]

Gen. et sp. nov.

Valid

Pérez-García et al.

Early Cretaceous (early Albian)

 Spain

A pleurosternid paracryptodiran. The type species is Toremys cassiopeia.

"Trionyx" jixiensis[190]

Sp. nov.

Valid

Li et al.

Early Cretaceous

Chengzihe Formation

 China

A member of Trionychinae of uncertain phylogenetic placement.

Xiaochelys[191]

Gen. et sp. nov

Valid

Zhou & Rabi

Early Cretaceous (Barremian-early Aptian)

Yixian Formation

 China

A member of Sinemydidae. The type species is Xiaochelys ningchengensis.

Archosauriformes

Pseudosuchians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Albertosuchus[197]

Gen. et sp. nov

Valid

Wu & Brinkman

Latest Cretaceous

Scollard Formation

 Canada

A crocodylian, probably a crocodyloid. The type species is Albertosuchus knudsenii.

Allodaposuchus hulki[198]

Sp. nov

Valid

Blanco et al.

Late Cretaceous (Maastrichtian)

Tremp Formation

 Spain

An eusuchian crocodylomorph, a species of Allodaposuchus.

Caiman wannlangstoni[199]

Sp. nov

Valid

Salas-Gismondi et al.

Late Middle Miocene

Pebas Formation

 Peru

A caiman, a species of Caiman.

Carnufex[200]

Gen. et sp. nov

Valid

Zanno et al.

Late Triassic (Carnian)

Pekin Formation

 United States

An early member of Crocodylomorpha. The type species is Carnufex carolinensis.

Dagasuchus[201]

Gen. et sp. nov

Valid

Lacerda, Schultz & Bertoni-Machado

Triassic (late Ladinian/early Carnian)

Santa Maria Formation

 Brazil

A non-crocodylomorph loricatan. The type species is Dagasuchus santacruzensis.

Gnatusuchus[199]

Gen. et sp. nov

Valid

Salas-Gismondi et al.

Late Middle Miocene

Pebas Formation

 Peru

A caiman. The type species is Gnatusuchus pebasensis.

Gorgetosuchus[202]

Gen. et sp. nov

Valid

Heckert et al.

Late Triassic

Pekin Formation

 United States

An aetosaur. The type species is Gorgetosuchus pekinensis.

Koumpiodontosuchus[203]

Gen. et sp. nov

Valid

Sweetman, Pedreira-Segade & Vidovic

Early Cretaceous (Barremian)

Wessex Formation

 United Kingdom

A crocodyliform related to Bernissartia. The type species is Koumpiodontosuchus aprosdokiti.

Kuttanacaiman[199]

Gen. et sp. nov

Valid

Salas-Gismondi et al.

Late Middle Miocene

Pebas Formation

 Peru

A caiman. The type species is Kuttanacaiman iquitosensis.

Lohuecosuchus[204]

Gen. et 2 sp. nov

Valid

Narváez et al.

Late Cretaceous (late Campanian-early Maastrichtian)

 France
 Spain

An eusuchian crocodyliform related to Allodaposuchus. The type species is Lohuecosuchus megadontos; genus also contains Lohuecosuchus mechinorum.

Zoneait[205]

Gen. et sp. nov

Valid

Wilberg

Middle Jurassic (Aalenian–Bajocian)

Snowshoe Formation

 United States

A thalattosuchian crocodylomorph. The type species is Zoneait nargorum.

Non-avian dinosaurs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Augustynolophus[211]

Gen. et comb. nov

Valid

Prieto-Márquez et al.

Late Cretaceous (late Maastrichtian)

Moreno Formation

 United States

A hadrosaurid; a new genus for "Saurolophus" morrisi Prieto-Márquez & Wagner (2013).

Boreonykus[212]

Gen. et sp. nov

Valid

Bell & Currie

Late Cretaceous (late Campanian)

Wapiti Formation

 Canada

A dromaeosaurid theropod. The type species is Boreonykus certekorum.

Chilesaurus[213]

Gen. et sp. nov.

Valid

Novas et al.

Late Jurassic (Tithonian)

Toqui Formation

 Chile

A theropod dinosaur, a basal member of Tetanurae. The type species is Chilesaurus diegosuarezi.

Crichtonpelta[214]

Gen. et comb. nov

Valid[215]

Arbour & Currie

Late Cretaceous (Cenomanian–Turonian)

Sunjiawan Formation

 China

An ankylosaurine ankylosaurid; a new genus for "Crichtonsaurus" benxiensis Lü, Ji, Gao & Li (2007).

Dakotaraptor[216]

Gen. et sp. nov.

Valid

DePalma et al.

Late Cretaceous (Maastrichtian)

Hell Creek Formation

 United States

A member of Dromaeosauridae. The type species is Dakotaraptor steini.

Eucnemesaurus entaxonis[217]

Sp. nov

Valid

McPhee et al.

Late Triassic

Lower Elliot Formation

 South Africa

A non-sauropod sauropodomorph, a species of Eucnemesaurus.

Galeamopus[206]

Gen. et comb. nov

Valid

Tschopp, Mateus & Benson

Late Jurassic (Kimmeridgian)

Morrison Formation

 United States

A diplodocid sauropod; a new genus for "Diplodocus" hayi Holland (1924).

Haestasaurus [218]

Gen. et comb. nov.

Valid

Upchurch, Mannion & Taylor

Early Cretaceous (late Berriasian—Valanginian)

Hastings Beds

 United Kingdom

A macronarian sauropod of uncertain phylogenetic placement; a new genus for "Pelorosaurus" becklesii Mantell (1852).

Horshamosaurus[219]

Gen. et comb. nov

Valid

Blows

Early Cretaceous

 United Kingdom

A new genus for "Polacanthus" rudgwickensis Blows (1996).

Hualianceratops [220]

Gen. et sp. nov.

Valid

Han et al.

Late Jurassic (Oxfordian)

Shishugou Formation

 China

A basal ceratopsian, possibly a member of Chaoyangsauridae. The type species is Hualianceratops wucaiwanensis.

Huanansaurus[221]

Gen. et sp. nov.

Valid

et al.

Late Cretaceous (Campanian-Maastrichtian)

Nanxiong Formation

 China

An oviraptorid theropod. The type species is Huanansaurus ganzhouensis.

Iguanodon galvensis[222]

Sp. nov

Valid

Verdú et al.

Early Cretaceous (early Barremian)

Galve, Teruel

 Spain

An ornithopod, a species of Iguanodon.

Ischioceratops[223]

Gen. et sp. nov.

Valid

He et al.

Late Cretaceous

Wangshi Group

 China

A leptoceratopsid ceratopsian. The type species is Ischioceratops zhuchengensis.

Koshisaurus[224]

Gen. et sp. nov

Valid

Shibata & Azuma

Early Cretaceous

Kitadani Formation

 Japan

A non-hadrosaurid hadrosauroid. The type species is Koshisaurus katsuyama.

Kunbarrasaurus[225]

Gen. et sp. nov.

Valid

Leahey et al.

Cretaceous (late Albian–(?)early Cenomanian)

Allaru Mudstone

 Australia

An ankylosaur. The type species is Kunbarrasaurus ieversi.

Lepidocheirosaurus[226]

Gen. et sp. nov.

Valid

Alifanov & Saveliev

Late Jurassic (Tithonian?)

Ukureyskaya Formation

 Russia

A dinosaur of uncertain phylogenetic placement; considered to be a theropod dinosaur related to Nqwebasaurus by the authors of its description,[226] but subsequently argued to be an ornithischian, possibly synonymous with Kulindadromeus.[227] The type species is Lepidocheirosaurus natatilis.

Lepidus[228]

Gen. et sp. nov.

Valid

Nesbitt & Ezcurra

Late Triassic

Dockum Group

 United States

A coelophysoid theropod. The type species is Lepidus praecisio.

Morelladon[229]

Gen. et sp. nov.

Valid

Gasulla et al.

Early Cretaceous (probably late Barremian)

Arcillas de Morella Formation

 Spain

An ornithopod dinosaur, a basal member of Styracosterna. The type species is Morelladon beltrani.

Mosaiceratops[230]

Gen. et sp. nov

Valid

Zheng, Jin & Xu

Late Cretaceous (early-middle Turonian—middle Campanian)

Xiaguan Formation

 China

A basal neoceratopsian. The type species is Mosaiceratops azumai.

Nebulasaurus[231]

Gen. et sp. nov

Valid

Xing et al.

Middle Jurassic (Aalenian or Bajocian)

Zhanghe Formation

 China

An eusauropod sauropod. The type species is Nebulasaurus taito.

Padillasaurus[232]

Gen. et sp. nov

Valid

Carballido et al.

Early Cretaceous (Barremian)

Paja Formation

 Colombia

A brachiosaurid sauropod. The type species is Padillasaurus leivaensis.

Probrachylophosaurus[233]

Gen. et sp. nov

Valid

Freedman Fowler & Horner

Late Cretaceous (Campanian)

Judith River Formation

 United States

A hadrosaurid. The type species is Probrachylophosaurus bergei.

Pulanesaura[234]

Gen. et sp. nov

Valid

McPhee et al.

Early Jurassic (probably late Hettangian to Sinemurian)

Upper Elliot Formation

 South Africa

A basal sauropod. The type species is Pulanesaura eocollum.

Qijianglong[235]

Gen. et sp. nov

Valid

Xing et al.

Late Jurassic

Suining Formation

 China

A mamenchisaurid sauropod. The type species is Qijianglong guokr.

Regaliceratops[236]

Gen. et sp. nov

Valid

Brown & Henderson

Late Cretaceous (probably middle Maastrichtian)

St. Mary River Formation

 Canada

A chasmosaurine ceratopsid. The type species is Regaliceratops peterhewsi.

Riabininohadros[237]

Gen. et comb. nov

Valid

Ulansky

Late Cretaceous (Maastrichtian)

Unnamed unit

 Ukraine

A hadrosaurid of uncertain placement; new genus for "Orthomerus" weberi Riabinin, 1945.

Saurornitholestes sullivani[238]

Sp. nov

Valid

Jasinski

Late Cretaceous (late Campanian)

Kirtland Formation

 United States

A member of Dromaeosauridae, a species of Saurornitholestes.

Sefapanosaurus[239]

Gen. et sp. nov

Valid

Otero et al.

Late Triassic or Early Jurassic

Elliot Formation

 South Africa

A non-sauropod sauropodomorph. The type species is Sefapanosaurus zastronensis.

Sirindhorna[240]

Gen. et sp. nov

Valid

Shibata et al.

Early Cretaceous (Aptian)

Khok Kruat Formation

 Thailand

A basal member of Hadrosauroidea. The type species is Sirindhorna khoratensis.

Ugrunaaluk[241]

Gen. et sp. nov

Valid

Mori, Druckenmiller & Erickson

Late Cretaceous (early Maastrichtian)

Prince Creek Formation

 United States

A saurolophine hadrosaurid. The type species is Ugrunaaluk kuukpikensis.

Wendiceratops [242]

Gen. et sp. nov.

Valid

Evans & Ryan

Late Cretaceous (Campanian)

Oldman Formation

 Canada

A centrosaurine ceratopsid. The type species is Wendiceratops pinhornensis.

Yi[243]

Gen. et sp. nov

Valid

Xu et al.

Jurassic (Callovian or Oxfordian)

Tiaojishan Formation

 China

A scansoriopterygid theropod. The type species is Yi qi.

Zhenyuanlong[244]

Gen. et sp. nov.

Valid

Lü & Brusatte

Early Cretaceous

Yixian Formation

 China

A dromaeosaurid theropod. The type species is Zhenyuanlong suni.

Birds

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Aegypius tugarinovi [253]

Sp. nov.

Valid

Manegold & Zelenkov

Early Pliocene, Ruscinian

 Moldova

An Old World vulture, Accipitridae, related to the cinereous vulture.

Alopecoenas norfolkensis [254]

Sp. nov

Valid

Forshaw

Holocene

 Australia

A new scientific name for the Norfolk ground dove. The previous name used to refer to this species, Columba norfolciensis Latham (1801), was suppressed by ICZN in 2010.[255]

Anas kurochkini [256]

Sp. nov.

Valid

Zelenkov & Panteleyev

Late Miocene

 Russia

An Anatidae.

Archaeornithura [257]

Gen. et sp. nov.

Valid

Wang et al.

Hauterivian

Huajiying Formation

 China

A Hongshanornithidae. Type species A. meemannae.

Athene inexpectata [258]

Sp. nov.

Valid

Pavia, Manegold, & Haarhoff

Early Pliocene

Varswater Formation

 South Africa

A Strigidae

Circaetus haemusensis [259]

Sp. nov.

Valid

Boev

Early Pleistocene

 Bulgaria

A snake eagle

Cratoavis [260]

Gen. et sp. nov.

Valid

Carvalho et al.

Aptian

Crato Member, Santana Formation

 Brazil

A member of Euenantiornithes of uncertain phylogenetic placement.[260][261]
The type species is C. cearensis.

Dunhuangia [262]

Gen. et sp. nov.

Valid

Wang et al.

Early Cretaceous

Xiagou Formation

 China

A member of Enantiornithes. The type species is D. cuii.

Eopachypteryx [263]

Gen. et sp. nov.

Valid

Mayr

Eocene

Messel Formation

 Germany

A bird of uncertain phylogenetic placement.
Type species E. praeterita.

Feitianius [264]

Gen. et sp. nov

Valid

O’Connor et al.

Early Cretaceous (Aptian)

Xiagou Formation

 China

A member of Enantiornithes. The type species is Feitianius paradisi.

Foshanornis [265]

Gen. et sp. nov.

Valid

Zhao et al.

Early Eocene

Buxin Formation

 China

A bird of uncertain phylogenetic placement, possibly a relative of trogons. Type species F. songi.

Fumicollis [266]

Gen. et sp. nov.

Valid

Bell & Chiappe

Late Cretaceous (late Coniacian—early Campanian)

The Smoky Hill Member of the Niobrara Chalk

 United States

A member of Hesperornithiformes. The type species is Fumicollis hoffmani.

Gallinago azovica [256]

Sp. nov.

Valid

Zelenkov & Panteleyev

Middle Turolian

 Russia

A snipe, Scolopacidae.

Hakawai [267]

Gen. et sp. nov.

Valid[268]

De Pietri et al.

Early Miocene (19–16 Ma)

Bannockburn Formation

 New Zealand

A member of Scolopaci of uncertain phylogenetic placement. The type species is Hakawai melvillei.

Holbotia [269]

Gen. et sp. nov.

Valid[270]

Zelenkov & Averianov

Early Cretaceous

 Mongolia

A member of Enantiornithes, probably a relative of Gobipteryx. Type species H. ponomarenkoi.

Houornis [271]

Gen. et comb. nov.

Valid[272]

Wang & Liu

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes of uncertain phylogenetic placement; a new genus for "Cathayornis" caudatus Hou (1997).

Hypsipetes cowlesi [273]

Sp. nov.

Valid[274]

Hume

Holocene

 Rodrigues

A bulbul, Pycnonotidae

Juehuaornis [275]

Gen. et sp. nov.

Valid

Wang, Wang & Hu

Early Cretaceous

Jiufotang Formation

 China

A basal member of Ornithuromorpha. Type species J. zhangi.

Kuntur [276]

Gen. et sp. nov.

Valid

Stucchi et al.

Late Miocene

Pisco Formation

 Peru

A condor. Type species K. cardenasi.

Lavadytis [277]

Gen. et sp. nov.

Valid[278]

Stidham & Hilton

Miocene (16.1–14.6 Ma)

 United States

A member of Anatidae; a member or a relative of members of the subfamily Oxyurinae. The type species is Lavadytis pyrenae.

Llallawavis [279]

Gen. et sp. nov

Valid

Degrange et al.

Late Pliocene

Playa Los Lobos Allo Formation

 Argentina

A mesembriornithine phorusrhacid. Type species L. scagliai.

Makahala [280]

Gen. et sp. nov.

Valid

Mayr

Latest Eocene or early Oligocene

Makah Formation

 United States

A member of Procellariiformes. Type species M. mirae.

Mangystania[281]

Gen. et sp. nov

Valid

Zvonok, Zelenkov & Danilov

Eocene (Bartonian)

Shorym Formation

 Kazakhstan

Possibly a member of Suliformes. The type species is Mangystania humilicristata.

Miobaptus huzhiricus [282]

Sp. nov

Valid

Zelenkov

Miocene

 Russia

A grebe

Mioporphyrula [283]

Gen. et comb. nov

Valid

Zelenkov in Zelenkov & Kurochkin

Late Miocene

 Moldova

A member of Rallidae. A new genus for "Tertiariaporphyrula" lungi Kurochkin & Ganea (1972).

Mwalau [284]

Gen. et sp. nov.

Valid

Worthy et al.

Holocene

 Vanuatu

A megapode. The type species is M. walterlinii.

Namapsitta [285]

Gen. et sp. nov.

Valid

Mourer-Chauviré, Pickford, & Senut

Middle Eocene

 Namibia

A Psittaciformes incertae sedis. Type species N. praeruptorum.

Nyctisoma [286]

Gen. et sp. nov.

Valid

Elżanowski & Zelenkov

Middle Miocene

Öoshin Formation

 Mongolia

An Ardeidae. Type species N. robusta.

Oligonomus [287]

Gen. et sp. nov.

Valid

De Pietri, Camens, & Worthy

Late Oligocene

 Australia

A member of Charadriiformes related to the plains-wanderer. Type species O. milleri.

Opisthodactylus horacioperezi [288]

Sp. nov.

Valid

Agnolin & Chafrat

Early Miocene

Chichinales Formation

 Argentina

A rheid ratite

Parapengornis [289]

Gen. et sp. nov.

Valid

Hu, O’Connor & Zhou

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes, a relative of Pengornis. Type species P. eurycaudatus.

Patagorhacos [288]

Gen. et sp. nov.

Valid

Agnolin & Chafrat

Early Miocene

Chichinales Formation

 Argentina

A member of Phorusrhacidae. Type species P. terrificus.

Petrosushkinia [283]

Nom. nov

Valid

Zelenkov in Zelenkov & Kurochkin

Late Miocene

 Kazakhstan

A member of Falconidae. A replacement name for Sushkinia Tugarinov (1935) (preoccupied).

Porzana botunensis [290]

Sp. nov.

Valid

Boev

Early Pleistocene

 Bulgaria

A member of Rallidae, a species of crake.

Pterodroma imberi [291]

Sp. nov.

Valid

Tennyson, Cooper & Shepherd

Holocene

 New Zealand

A member of Procellariidae, a species of gadfly petrel.

Pterygornis [292]

Gen. et sp. nov.

Valid [293]

Wang, Hu & Li

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes. Type species P. dapingfangensis.

Rallus adolfocaesaris [294]

Sp. nov.

Valid

Alcover et al.

Holocene

 Madeira (Porto Santo Island)

A member of Rallidae, a species of Rallus

Rallus carvaoensis [294]

Sp. nov.

Valid

Alcover et al.

Holocene

 Azores (São Miguel Island)

A member of Rallidae, a species of Rallus

Rallus lowei [294]

Sp. nov.

Valid

Alcover et al.

Holocene

 Madeira (Madeira Island)

A member of Rallidae, a species of Rallus

Rallus minutus [294]

Sp. nov.

Junior homonym

Alcover et al.

Holocene

 Azores (São Jorge Island)

A member of Rallidae, a species of Rallus. The specific name is a junior homonym of Rallus minutus Pallas (1776), Rallus minutus Gmelin (1789) and Rallus minutus Forster (1844).

Rallus montivagorum [294]

Sp. nov.

Valid

Alcover et al.

Holocene

 Azores (Pico Island)

A member of Rallidae, a species of Rallus

Ramphastosula aguirrei [295]

Sp. nov.

Valid

Stucchi, Varas-Malca & Urbina-Schmitt

Miocene

Pisco Formation

 Peru

A member of Sulidae, a species of Ramphastosula.

Scolopax brachycarpa [296]

Sp. nov.

Valid

Takano & Steadman

Late Holocene

 Haiti

A woodcock.

Scopelortyx [285]

Gen. nov. et Sp. nov.

Valid

Mourer-Chauviré, Pickford, & Senut

Middle Eocene

 Namibia

A Paraortygidae, Galliformes. Type species S. klinghardtensis.

Shiriyanetta [297]

Gen. et sp. nov.

Valid

Watanabe & Matsuoka

Pleistocene

 Japan

A duck. The type species is Shiriyanetta hasegawai.

Stemec [298]

Gen. et sp. nov.

Valid

Kaiser, Watanabe & Johns

Late Oligocene

Sooke Formation

 Canada

A member of Plotopteridae. The type species is Stemec suntokum.

Sula brandi [295]

Sp. nov.

Valid

Stucchi, Varas-Malca & Urbina-Schmitt

Miocene

Pisco Formation

 Peru

A booby.

Sula figueroae [295]

Sp. nov.

Valid

Stucchi, Varas-Malca & Urbina-Schmitt

Miocene

Pisco Formation

 Peru

A booby.

Tyto cravesae [299]

Sp. nov.

Valid

Suárez & Olson

Quaternary

 Cuba

A barn-owl, a species of Tyto.

Tyto richae [258]

Sp. nov

Valid

Pavia, Manegold, & Haarhoff

Early Pliocene

Varswater Formation

 South Africa

A Tytonidae barn-owl.

Yuanjiawaornis [300]

Gen. et sp. nov.

Valid

Hu et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes. Type species Yuanjiawaornis viriosus.

Pterosaurs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Arcticodactylus[303]

Gen. et comb. nov

Valid

Kellner

Late Triassic (?Norian-Rhaetian)

Fleming Fjord Formation

 Greenland

A non-pterodactyloid pterosaur of uncertain phylogenetic placement; a new genus for "Eudimorphodon" cromptonellus Jenkins et al. (2001).

Austriadraco[303]

Gen. et sp. nov

Valid

Kellner

Late Triassic (late Norian)

Seefeld Formation

 Austria

A non-pterodactyloid pterosaur of uncertain phylogenetic placement. The type species is Austriadraco dallavecchiai.

Banguela[304]

Gen. et sp. nov

Valid

Headden & Campos

Early Cretaceous

Romualdo Formation

 Brazil

A dsungaripterid. The type species is Banguela oberlii.

Bergamodactylus[303]

Gen. et sp. nov

Valid

Kellner

Late Triassic (Alaunian)

Calcari di Zorzino Formation

 Italy

A relative of Campylognathoides. The type species is Bergamodactylus wildi.

Cimoliopterus dunni[305]

Sp. nov

Valid

Myers

Late Cretaceous (late Cenomanian)

Britton Formation

 United States

A member of Pterodactyloidea, probably a pteranodontoid; a species of Cimoliopterus.

Daohugoupterus[306]

Gen. et sp. nov

Valid

Cheng et al.

Late Jurassic

Daohugou Beds

 China

A non-pterodactyloid pterosaur. The type species is Daohugoupterus delicatus.

Linlongopterus[307]

Gen. et sp. nov

Valid

Rodrigues et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Pteranodontoidea, possibly related to anhanguerians. The type species is Linlongopterus jennyae.

Orientognathus[308]

Gen. et sp. nov

Valid

et al.

Late Jurassic

Tuchengzi Formation

 China

A rhamphorhynchid. The type species is Orientognathus chaoyngensis.

Rhamphorhynchus etchesi[309]

Sp. nov

Valid

O'Sullivan & Martill

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A species of Rhamphorhynchus.

Other reptiles

Name Novelty Status Authors Age Unit Location Notes Images

Eohyosaurus[310]

Gen. et sp. nov

Valid

Butler et al.

Middle Triassic (early Anisian)

Burgersdorp Formation

 South Africa

A basal rhynchosaur. The type species is Eohyosaurus wolvaardti.

Erpetonyx[311]

Gen. et sp. nov

Valid

Modesto et al.

Carboniferous (Gzhelian)

Egmont Bay Formation

 Canada

A relative of bolosaurids. The type species is Erpetonyx arsenaultorum.

Glaurung[312]

Gen. et sp. nov

Valid

Bulanov & Sennikov

Late Permian

 Germany

A member of Weigeltisauridae. The type species is Glaurung schneideri.

Opisthodontosaurus[313]

Gen. et sp. nov

Valid

Reisz et al.

Early Permian

Garber Formation

 United States

A member of Captorhinidae. The type species is Opisthodontosaurus carrolli.

Pappochelys[314]

Gen. et sp. nov

Valid

Schoch & Sues

Middle Triassic (Ladinian)

Erfurt Formation

 Germany

A stem-turtle. The type species is Pappochelys rosinae.

"Thalassodromeus" sebesensis[315]

Sp. nov

Valid

Grellet-Tinner & Codrea

Late Cretaceous (Maastrichtian)

 Romania

A reptile of uncertain phylogenetic placement; initially classified as a pterosaur and a species of Thalassodromeus,[315] but subsequently argued to be a turtle and a junior synonym of Kallokibotion bajazidi.[316]

Synapsids

Newly named non-mammalian synapsids

Name Novelty Status Authors Age Unit Location Notes Images

Endothiodon tolani[317]

Sp. nov

Valid

Cox & Angielczyk

Permian

Ruhuhu Formation

 Tanzania

A dicynodont, a species of Endothiodon.

Ichibengops[318]

Gen. et sp. nov

Valid

Huttenlocker, Sidor & Angielczyk

Late Permian

Madumabisa Mudstone Formation

 Zambia

A therocephalian. The type species is Ichibengops munyamadziensis.

Lende[319]

Gen. et sp. nov

Valid

Kruger et al.

Late Permian

Chiweta Beds

 Malawi

A burnetiamorph biarmosuchian therapsid. The type species is Lende chiweta.

Nothogomphodon sanjiaoensis[320]

Sp. nov

Valid

Liu & Abdala

Middle Triassic

Ermaying Formation

 China

A therocephalian, a species of Nothogomphodon.

Ruberodon[321]

Gen. et sp. nov

Valid

Ray

Late Triassic

Tiki Formation

 India

A traversodontid cynodont. The type species is Ruberodon roychowdhurii.

Sinokannemeyeria baidaoyuensis[322]

Sp. nov

Valid

Liu

Middle Triassic

Tongchuan Formation

 China

A kannemeyeriiform dicynodont, a species of Sinokannemeyeria.

Mammals

Research

New taxa

Newly named non-eutherian mammals
Metatherians
Name Novelty Status Authors Age Unit Location Notes Images

Antawallathentes[325]

Gen. et 2 sp. nov

Valid

Rincón et al.

Oligocene (Deseadan)

 Bolivia

A palaeothentid marsupial (a relative of the shrew opossums). Genus contains two species: Antawallathentes illimani and Antawallathentes quimsacruza.

Archaeonothos[326]

Gen. et sp. nov

Valid

Beck

Early Eocene

Murgon fossil site

 Australia

A metatherian of uncertain phylogenetic placement. The type species is Archaeonothos henkgodthelpi.

Ganguroo robustiter[327]

Sp. nov

Valid

Cooke et al.

Miocene

Riversleigh World Heritage Area

 Australia

A basal member of Macropodidae, a species of Ganguroo.

Gaylordia mater[328]

Sp. nov

Valid

Oliveira & Goin

Itaboraian

Itaboraí Basin

 Brazil

A metatherian of uncertain phylogenetic placement, possibly a relative of Pucadelphys; a species of Gaylordia.

Lotheridium[329]

Gen. et sp. nov

Valid

Bi et al.

Late Cretaceous

Qiupa Formation

 China

A member of Deltatheridiidae. The type species is Lotheridium mengi.

Lycopsis padillai[330]

Sp. nov

Valid[331]

Suarez et al.

Miocene

Castilletes Formation

 Colombia

A member of Sparassodonta, a species of Lycopsis.

Madju[332]

Gen. et 2 sp. nov

Valid

Travouillon et al.

Late Oligocene to early late Miocene

 Australia

A bandicoot. Genus contains two species: Madju variae and Madju encorensis.

Nimbavombatus[333]

Gen. et sp. nov

Valid

Brewer et al.

Early Miocene

Riversleigh World Heritage Area

 Australia

A wombat. The type species is Nimbavombatus boodjamullensis.

Oklatheridium wiblei[334]

Sp. nov

Valid

Cifelli & Davis

Early Cretaceous (Albian)

Cloverly Formation

 United States

A member of Deltatheroida, a species of Oklatheridium.

Sedophascolomys[335]

Gen. et comb. nov

Valid

Louys

Pliocene to Pleistocene

 Australia

A wombat; a new genus for "Phascolomys" medius Owen (1872).

Sparassocynus maimarai[336]

Sp. nov

Valid

Abello et al.

Late Miocene

Maimará Formation

 Argentina

A sparassocynid didelphoid (a relative of the opossums), a species of Sparassocynus.

Tsagandelta[337]

Gen. et sp. nov

Valid

Rougier, Davis & Novacek

Late Cretaceous

Baynshiree Formation

 Mongolia

A member of Deltatheroida. The type species is Tsagandelta dashzevegi.

Wabularoo prideauxi[338]

Sp. nov

Valid

Travouillon, Archer & Hand

Early Miocene

Riversleigh World Heritage Area

 Australia

A member of Macropodoidea, a species of Wabularoo.

Others
Name Novelty Status Authors Age Unit Location Notes Images

Agilodocodon[339]

Gen. et sp. nov

Valid

Meng et al.

Middle Jurassic

"Daohugou site in Tiaojishan Formation (...), also known as Jiulongshan Formation"[339]

 China

A docodont. The type species is Agilodocodon scansorius.

Argaliatherium[334]

Gen. et sp. nov

Valid

Cifelli & Davis

Early Cretaceous (Albian)

Cloverly Formation

 United States

A basal member of Tribosphenida. The type species is Argaliatherium robustum.

Carinalestes[334]

Gen. et sp. nov

Valid

Cifelli & Davis

Early Cretaceous (Albian)

Cloverly Formation

 United States

A basal member of Tribosphenida. The type species is Carinalestes murensis.

Docodon apoxys[340]

Sp. nov

Valid

Rougier et al.

Late Jurassic

Morrison Formation

 United States

A docodont, a species of Docodon.

Docofossor[341]

Gen. et sp. nov

Valid

Luo et al.

Late Jurassic

Tiaojishan Formation

 China

A docodont. The type species is Docofossor brachydactylus.

Gobiconodon haizhouensis[342]

Sp. nov

Valid

Kusuhashi et al.

Early Cretaceous

Fuxin Formation

 China

A gobiconodontid, a species of Gobiconodon.

Gobiconodon tomidai[342]

Sp. nov

Valid

Kusuhashi et al.

Early Cretaceous

Shahai Formation

 China

A gobiconodontid, a species of Gobiconodon.

Kimbetopsalis[343]

Gen. et sp. nov

Valid

Williamson et al.

Early Paleocene (Puercan)

Nacimiento Formation

 United States

A taeniolabidid multituberculate. The type species is Kimbetopsalis simmonsae.

Spinolestes[344]

Gen. et sp. nov

Valid

Martin et al.

Early Cretaceous (latest Barremian)

Calizas de la Huérgina Formation

 Spain

A gobiconodontid. The type species is Spinolestes xenarthrosus.

Tirotherium[345]

Gen. et sp. nov

Valid

Montellano-Ballesteros & Fox

Late Cretaceous (late Santonian to early Campanian)

Milk River Formation

 Canada

An early member of Boreosphenida, probably a member of Picopsidae. The type species is Tirotherium aptum.

Valenopsalis[343]

Gen. et comb. nov

Valid

Williamson et al.

Early Paleocene (Puercan)

Bug Creek Anthills and Bug Creek West

 United States

A taeniolabidoid multituberculate; a new genus for "Catopsalis" joyneri Sloan & Van Valen (1965).

Yubaatar[346]

Gen. et sp. nov

Valid

Xu et al.

Late Cretaceous

Qiupa Formation

 China

A cimolodont multituberculate. The type species is Yubaatar zhongyuanensis.

Eutherians
Xenarthrans
Name Novelty Status Authors Age Unit Location Notes Images

Eionaletherium[347]

Gen. et sp. nov.

Valid

Rincón et al.

Late Miocene

Urumaco Formation

 Venezuela

A mylodontoid sloth. The type species is Eionaletherium tanycnemius.

Tonnicinctus[348]

Gen. et sp. nov.

Valid

Góis et al.

Pleistocene (Ensenadan to Lujanian)

 Argentina

A member of Pampatheriidae. The type species is Tonnicinctus mirus.

Afrotherians
Name Novelty Status Authors Age Unit Location Notes Images

Callistosiren[349]

Gen. et sp. nov.

Valid

Vélez-Juarbe & Domning

Late Oligocene

Lares Limestone

 United States ( Puerto Rico)

A dugong. The type species is Callistosiren boriquensis.

Palaeoloxodon xylophagou[350]

Sp. nov

Valid

Athanassiou et al.

Late middle Pleistocene

 Cyprus

A member of Elephantidae, a species of Palaeoloxodon.

Protanancus brevirostris[351]

Sp. nov.

Valid

Wang et al.

Early Miocene

Linxia Basin

 China

A gomphothere, a species of Protanancus.

Thyrohyrax libycus[352]

Sp. nov

Valid

Coster et al.

Early Oligocene

 Libya

A saghatheriid hyracoid, a species of Thyrohyrax.

Bats
Name Novelty Status Authors Age Unit Location Notes Images

Cardioderma leakeyi[353]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A member of Megadermatidae related to the heart-nosed bat.

Marnenycteris[354]

Gen. et sp. nov

Valid

Hand et al.

Eocene (Ypresian)

 France

A bat related to Onychonycteris. The type species is Marnenycteris michauxi.

Mystacina miocenalis[355]

Sp. nov.

Valid

Hand, Lee, Worthy & Archer in Hand et al.

Early Miocene (Altonian, 19–16 Ma)

 New Zealand

A New Zealand short-tailed bat.

Myzopoda africana[353]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A species of Myzopoda.

Nycticeinops serengetiensis[353]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A vesper bat related to the Schlieffen's bat.

Otonycteris rummeli[356]

Sp. nov

Valid

Rosina

Late Miocene

 Ukraine

A vesper bat related to the desert long-eared bat.

Scotoecus olduvensis[353]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A vesper bat, a species of Scotoecus.

Odd-toed ungulates
Name Novelty Status Authors Age Unit Location Notes Images

Boreohippidion[357]

Gen. et comb. nov.

Valid

Avilla, Bernardes & Mothé

Early Pliocene (late Hemphillian)

 United States

A member of Equidae; a new genus for "Onohippidium" galushai Macfadden & Skinner (1979).

Chasmotherium depereti[358]

Sp. nov

Valid

Remy

Eocene (Bartonian)

 France

A relative of Hyrachyus, a species of Chasmotherium.

Eolophiodon[359]

Gen. et sp. nov.

Valid

Robinet et al.

Early Eocene

 France

A relative of Lophiodon. The type species is Eolophiodon laboriense.

Pachynolophus gaytei[358]

Sp. nov

Valid

Remy

Eocene (Bartonian)

 France

A member of Palaeotheriidae, a species of Pachynolophus.

Persiatherium[360]

Gen. et sp. et comb. nov

Valid

Pandolfi

Late Miocene

 China
 Iran

A rhinoceros related to Aceratherium. Genus includes new species Persiatherium rodleri, as well as "Aceratherium" huadeensis.

Plesiohipparion shanxiense[361]

Sp. nov.

Valid

Bernor, Sun & Chen

Early Pleistocene

 China

A member of Equidae related to Hipparion; a species of Plesiohipparion.

Vastanolophus[362]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

Cambay Shale Formation

 India

A member of Tapiroidea, possibly a member of Helaletidae. The type species is Vastanolophus holbrooki.

Even-toed ungulates
Name Novelty Status Authors Age Unit Location Notes Images

Anthracokeryx naduongensis[363]

Sp. nov

Valid

Ducrocq et al.

Eocene

 Vietnam

An anthracothere, a species of Anthracokeryx.

Bothriogenys langsonensis[363]

Sp. nov

Valid

Ducrocq et al.

Eocene

 Vietnam

An anthracothere, a species of Bothriogenys.

Bothriogenys vietnamensis[363]

Sp. nov

Valid

Ducrocq et al.

Eocene

 Vietnam

An anthracothere, a species of Bothriogenys.

Eotragus lampangensis[364]

Sp. nov

Valid

Suraprasit et al.

Late Middle Miocene (13.4-13.2 Ma)

Nakhaem Formation

 Thailand

A bovid, a species of Eotragus.

Epirigenys[365]

Gen. et sp. nov

Valid

Lihoreau et al.

Oligocene

Lokone Sandstone Formation

 Kenya

A relative of hippopotamuses. The type species is Epirigenys lokonensis.

Eucladoceros montenegrensis[366]

Sp. nov

Valid

Van der Made & Dimitrijević

Early Pleistocene

 Montenegro

A deer, a species of Eucladoceros.

Euprox grandis[367]

Sp. nov

Valid

Hou

Late Miocene

Liushu Formation

 China

A member of Cervidae related to muntjacs, a species of Euprox.

Kolpochoerus phillipi[368]

Sp. nov

Valid

Souron, Boisserie & White

Late Pliocene or early Pleistocene (ca. 2.5 Ma)

Middle Awash

 Ethiopia

A suid, a species of Kolpochoerus.

Lucashyus[369]

Gen. et sp. nov

Valid

Prothero

Late early Arikareean

 United States

A peccary. The type species is Lucashyus coombsae.

Marshochoerus[369]

Gen. et comb. nov

Valid

Prothero

Late Arikareean to late Hemingfordian

 United States

A peccary; a new genus for "Thinohyus" socialis Marsh (1875).

Muntiacus ? huangi[370]

Sp. nov

Valid

Dong & Chen

Pleistocene

 China

A member of Cervidae, possibly a muntjac.

Nguruwe ? galaticum[371]

Sp. nov

Valid

Orliac et al.

Late early Miocene

Kumartaş Formation

 Turkey

A hyotheriine suid, possibly a species of Nguruwe.

Paratoceras coatesi[372]

Sp. nov

Valid

Rincon et al.

Miocene (early Hemingfordian)

Cucaracha Formation

 Panama

A member of Protoceratidae, a species of Paratoceras.

Paratoceras orarius[372]

Sp. nov

Valid

Rincon et al.

Early Miocene

Culebra Formation

 Panama

A member of Protoceratidae, a species of Paratoceras.

Sivacobus sankaliai[373]

Sp. nov

Valid

Vrba, Bibi & Costa

Late Pleistocene

 India

A member of Bovidae belonging to the subfamily Reduncinae, a species of Sivacobus.

Stuckyhyus[369]

Gen. et comb. nov

Valid

Prothero

Late Arikareean

 United States

A peccary; a new genus for "Thinohyus" siouxensis Peterson (1905).

Wrightohyus[369]

Gen. et sp. nov

Valid

Prothero

Late Arikareean to early Hemingfordian

 United States

A peccary. The type species is Wrightohyus yatkolai.

Xenokeryx[374]

Gen. et sp. nov

Valid

Sánchez et al.

Miocene (middle Aragonian)

 Spain

A member of Palaeomerycidae. The type species is Xenokeryx amidalae.

Cetaceans
Name Novelty Status Authors Age Unit Location Notes Images

Albicetus[375]

Gen. et comb. nov

Valid

Boersma & Pyenson

Miocene (Langhian)

Monterey Formation

 United States

A basal member of Physeteroidea. The type species is "Ontocetus" oxymycterus Kellogg (1925).

Ashleycetus[376]

Gen. et sp. nov

Valid

Sanders & Stepleton

Oligocene (late Rupelian)

Ashley Formation

 United States

A basal odontocete. The type species is Ashleycetus planicapitis.

Brabocetus[377]

Gen. et sp. nov

Valid

Colpaert, Bosselaers & Lambert

Pliocene (Zanclean)

Kattendijk Formation

 Belgium

A porpoise. The type species is Brabocetus gigasei.

Chilcacetus[378]

Gen. et sp. nov

Valid

Lambert, De Muizon & Bianucci

Early Miocene

Chilcatay Formation

 Peru

A toothed whale of uncertain phylogenetic placement, possibly related to Argyrocetus and Macrodelphinus. The type species is Chilcacetus cavirhinus.

Chrysocetus fouadassii[379]

Sp. nov

Valid

Gingerich & Zouhri

Eocene (Bartonian)

Aridal Formation

Western Sahara

A basilosaurid, a species of Chrysocetus.

Fucaia[380]

Gen. et sp. et comb. nov

Valid

Marx, Tsai & Fordyce

Oligocene

Makah Formation
Pysht Formation

 United States

A member of Aetiocetidae. The type species is Fucaia buelli; genus also includes "Chonecetus" goedertorum Barnes & Furusawa in Barnes et al. (1994).

Horopeta[381]

Gen. et sp. nov

Valid[382]

Tsai & Fordyce

Oligocene (early Chattian)

Kokoamu Greensand

 New Zealand

An early baleen whale. The type species is Horopeta umarere.

Isthminia[383]

Gen. et sp. nov

Valid

Pyenson et al.

Miocene (Messinian)

Chagres Formation

 Panama

An inioid river dolphin. The type species is Isthminia panamensis.

Nanokogia[384]

Gen. et sp. nov

Valid

Velez-Juarbe et al.

Late Miocene

Chagres Formation

 Panama

A member of Kogiidae. The type species is Nanokogia isthmia.

Otekaikea huata[385]

Sp. nov

Valid

Tanaka & Fordyce

Oligo/Miocene boundary (Waitakian)

Otekaike Limestone

 New Zealand

A relative of the South Asian river dolphin, a species of Otekaikea.

Platyosphys aithai[379]

Sp. nov

Valid

Gingerich & Zouhri

Eocene (Bartonian)

Aridal Formation

Western Sahara

A basilosaurid, a species of Platyosphys.

Tokarahia[386]

Gen. et sp. et comb. nov

Valid[387]

Boessenecker & Fordyce

Late Oligocene

Kokoamu Greensand
Otekaike Limestone

 New Zealand

An eomysticetid baleen whale. The type species is Tokarahia kauaeroa; genus also contains "Mauicetus" lophocephalus Marples (1956).

Tranatocetus[388]

Gen. et comb. nov

Valid

Gol’din & Steeman

Miocene (Tortonian)

Gram Formation

 Denmark

A baleen whale related to rorquals and gray whales; a new genus for "Mesocetus" argillarius Roth (1978).

Holotype of Tranatocetus argillarius

Waharoa[389]

Gen. et sp. nov

Valid

Boessenecker & Fordyce

Late Oligocene

Otekaike Limestone

 New Zealand

An eomysticetid baleen whale. The type species is Waharoa ruwhenua.

Carnivorans
Name Novelty Status Authors Age Unit Location Notes Images

Archaeodobenus[390]

Gen. et sp. nov

Valid

Tanaka & Kohno

Late Miocene

Ichibangawa Formation

 Japan

A member of Odobenidae (a relative of the walrus). The type species is Archaeodobenus akamatsui.

Australophoca[391]

Gen. et sp. nov

Valid[392]

Valenzuela-Toro et al.

Late Miocene

Bahía Inglesa Formation
Pisco Formation

 Chile
 Peru

An earless seal belonging to the subfamily Monachinae. The type species is Australophoca changorum.

Cynelos malasi[393]

Sp. nov

Valid

Hunt & Stepleton

Early Miocene

 United States

An amphicyonid, a species of Cynelos.

Cynotherium malatestai[394]

Sp. nov

Valid

Madurell-Malapeira, Palombo & Sotnikova

Early middle Pleistocene

 Italy

A relative of the Sardinian dhole.

Devinophoca emryi[395]

Sp. nov

Valid

Koretsky & Rahmat

Early Middle Miocene (early Badenian)

 Slovakia

A devinophocine earless seal, a species of Devinophoca.

Eotaria[396]

Gen. et sp. nov

Valid

Boessenecker & Churchill

Miocene (Burdigalian–Langhian)

 United States

A fur seal. The type species is Eotaria crypta.

Lutra hearsti[397]

Sp. nov

Valid

Geraads et al.

Pliocene

Hadar Formation

 Ethiopia

An otter, a species of Lutra.

Praepusa boeska[398]

Sp. nov

Valid

Koretsky, Peters & Rahmat

Late Miocene—Pliocene

 Belgium
 Netherlands

An earless seal, a species of Praepusa.

Vulpes mathisoni[399]

Sp. nov

Valid

Geraads et al.

Pliocene

Mursi Formation

 Ethiopia

A fox, a species of Vulpes.

Yoshi[400]

Gen. et sp. et comb. nov

Valid

Spassov & Geraads

Late Miocene

 China
 Greece
 Macedonia

A felid. The type species is Yoshi garevskii; genus also contains "Metailurus" parvulus (Hensel, 1862) (nomen dubium) and "Metailurus" minor Zdansky, 1924.

Rodents
Name Novelty Status Authors Age Unit Location Notes Images

Acarechimys leucotheae[401]

Sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

An octodontoid caviomorph of uncertain phylogenetic placement, a species of Acarechimys.

Allotypomys[402]

Gen. et sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Eutypomyidae. The type species is Allotypomys pictus.

Apeomys whistleri[402]

Sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Eomyidae, a species of Apeomys.

Bursagnathus[402]

Gen. et sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Heteromyidae. The type species is Bursagnathus aterosseus.

Cephalomys ceciae[401]

Sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

A cephalomyid, a species of Cephalomys.

Chenomys[403]

Gen. et sp. nov

Valid

Li & Meng

Earliest Eocene

Nomogen Formation

 China

A ctenodactyloid rodent. The type species is Chenomys orientalis.

Doryperimys[404]

Gen. et sp. nov

Valid

Kramarz, Bond & Arnal

Early Miocene

Cerro Bandera Formation

 Argentina

A member of Neoepiblemidae. The type species is Doryperimys olsacheri.

Eliomys yevesi[405]

Sp. nov

Valid

Mansino et al.

Late Miocene (late Turolian)

Villatoya-Venta del Moro Formation

 Spain

A dormouse, a species of Eliomys.

Ethelomys[401]

Gen. et comb. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

An octodontoid caviomorph of uncertain phylogenetic placement; a new genus for "Deseadomys" loomisi Wood & Patterson (1959).

Eumysops marplatensis[406]

Sp. nov

Valid

Olivares & Verzi

Late Pliocene to early Pleistocene

San Andrés Formation
Vorohué Formation

 Argentina

A member of Echimyidae, a species of Eumysops.

Galileomys baios[401]

Sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

An acaremyid octodontoid caviomorph, a species of Galileomys.

Heterotamias[407]

Gen. et comb. nov

Valid

Qiu

Early Miocene

Xiacaowan Formation

 China

A chipmunk; a new genus for "Eutamias" sihongensis Qiu & Lin (1986).

Incamys menniorum[401]

Sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

A chinchilloid caviomorph of uncertain phylogenetic placement, a species of Incamys.

Kabirmys prius[408]

Sp. nov

Valid

Coster et al.

Eocene

 Libya

A nementchamyid anomaluromorph, a species of Kabirmys.

Lamugaulus[409]

Gen. et sp. nov

Valid

Tesakov & Lopatin

Early Miocene

Khalagay Formation

 Russia

A promylagauline mylagaulid. The type species is Lamugaulus olkhonensis.

Leggadina gregoriensis[410]

Sp. nov

Valid

Klinkhamer & Godthelp

Early Pleistocene

Riversleigh World Heritage Area

 Australia

A murine murid, a species of Leggadina.

Leggadina macrodonta[410]

Sp. nov

Valid

Klinkhamer & Godthelp

Pliocene or Pleistocene

 Australia

A murine murid, a species of Leggadina.

Leucokephalos[401]

Gen. et 2 sp. nov

Valid

Vucetich et al.

Late Oligocene to early Miocene

 Argentina

A caviomorph of uncertain phylogenetic placement. The type species is Leucokephalos zeffiae Vucetich et al. (2015);[401] genus also contains Leucokephalos maior Kramarz, Bond & Arnal (2015).[404]

Llitun[401]

Gen. et sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

A caviomorph of uncertain phylogenetic placement. The type species is Llitun notuca.

Loncolicu[401]

Gen. et sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

A chinchilloid caviomorph of uncertain phylogenetic placement. The type species is Loncolicu tretos.

Metaphiomys zallahensis[352]

Sp. nov

Valid

Coster et al.

Early Oligocene

 Libya

A hystricognathous rodent, a species of Metaphiomys.

Microtheriomys[402]

Gen. et sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

An anchitheriomyine castorid. The type species is Microtheriomys brevirhinus.

Miosciurus covensis[402]

Sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A sciurine sciurid, a species of Miosciurus.

Mus denizliensis[411]

Sp. nov

Valid

Erten, Sen & Sagular

Early Pleistocene

Denizli Basin

 Turkey

A mouse.

Neoadjidaumo arctozophus[402]

Sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Eomyidae, a species of Neoadjidaumo.

Neophiomys dawsonae[352]

Sp. nov

Valid

Coster et al.

Early Oligocene

 Libya

A relative of the cane rats, a species of Neophiomys.

Omboomys[412]

Gen. et sp. nov

Valid

Maridet et al.

Late Miocene

 Mongolia

An eomyid rodent. The type species is Omboomys builstynensis.

Palaeosciurus jiangi[407]

Sp. nov

Valid

Qiu

Early Miocene

Xiacaowan Formation

 China

A ground squirrel, a species of Palaeosciurus.

Plesiosminthus fremdi[402]

Sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A dipodid related to the birch mice, a species of Plesiosminthus.

Proafricanomys[413]

Gen. et sp. nov

Valid

López-Antoñanzas et al.

Miocene (Tortonian)

 Lebanon

A gundi. The type species is Proafricanomys libanensis.

Proapeomys[402]

Gen. et sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Eomyidae. The type species is Proapeomys condoni; genus might also contain the species "Florentiamys" lulli Wood (1936).

Proheteromys latidens[402]

Sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Heteromyidae, a species of Proheteromys.

Promimomys enginae[414]

Sp. nov

Valid

Suata-Alpaslan

Early Pliocene

 Turkey

A member of Arvicolidae, a species of Promimomys.

Protacaremys? adilos[401]

Sp. nov

Valid

Vucetich et al.

Late Oligocene

 Argentina

An octodontoid caviomorph of uncertain phylogenetic placement, possibly a species of Protacaremys.

Prozenkerella[408]

Gen. et sp. nov

Valid

Coster et al.

Early Oligocene

 Libya

An anomalure. The type species is Prozenkerella saharaensis.

Pseudoacaremys[415]

Gen. et sp. nov

Valid

Arnal & Vucetich

Miocene

 Argentina

An acaremyid octodontoid caviomorph rodent. The type species is Pseudoacaremys kramarzi.

Simplicimys[403]

Gen. et sp. nov

Valid

Li & Meng

Early-middle Eocene

Arshanto Formation
Irdin Manha Formation

 China

A ctenodactyloid rodent. The type species is Simplicimys bellus.

Subathumys[416]

Gen. et 2 sp. nov

Valid

Gupta & Kumar

Early Eocene (Ypresian)

Subathu Formation

 India

A relative of gundis. The type species is Subathumys solanorius; genus also contains Subathumys globulus.

Tachyoryctoides bayarmae[417]

Sp. nov

Valid

Daxner-Höck, Badamgarav & Maridet

Late Oligocene

 Mongolia

A subterranean rodent belonging to the subfamily Tachyorytoidinae, a species of Tachyoryctoides.

Tachyoryctoides radnai[417]

Sp. nov

Valid

Daxner-Höck, Badamgarav & Maridet

Late Oligocene

 Mongolia

A subterranean rodent belonging to the subfamily Tachyorytoidinae, a species of Tachyoryctoides.

Tamquammys fractus[403]

Sp. nov

Valid

Li & Meng

Middle Eocene

Irdin Manha Formation

 China

A ctenodactyloid rodent, a species of Tamquammys.

Tamquammys longus[403]

Sp. nov

Valid

Li & Meng

Early Eocene

Arshanto Formation

 China

A ctenodactyloid rodent, a species of Tamquammys.

Tamquammys robustus[403]

Sp. nov

Valid

Li & Meng

Early Eocene

Arshanto Formation
Nomogen Formation

 China

A ctenodactyloid rodent, a species of Tamquammys.

Trogomys oregonensis[402]

Sp. nov

Valid

Korth & Samuels

Arikareean

John Day Formation

 United States

A member of Heteromyidae, a species of Trogomys.

Yongshengomys[403]

Gen. et sp. nov

Valid

Li & Meng

Middle Eocene

Irdin Manha Formation

 China

A ctenodactyloid rodent. The type species is Yongshengomys extensus.

Yuomys huheboerhensis[403]

Sp. nov

Valid

Li & Meng

Middle Eocene

Irdin Manha Formation

 China

A ctenodactyloid rodent, a species of Yuomys.

Primates
Name Novelty Status Authors Age Unit Location Notes Images

Australopithecus deyiremeda[418]

Sp. nov

Valid

Haile-Selassie et al.

Pliocene

 Ethiopia

A species of Australopithecus.

Ekembo[419]

Gen. et comb. nov

Valid

McNulty et al.

Early Miocene

Hiwegi Formation

 Kenya

An ape of uncertain phylogenetic placement. A new genus for "Proconsul" nyanzae Le Gros Clark & Leakey (1950); genus also contains "Proconsul" heseloni Walker et al. (1993).

Ekgmowechashala zancanellai[420]

Sp. nov

Valid

Samuels, Albright & Fremd

Arikareean

John Day Formation

 United States

A primate of uncertain phylogenetic placement, probably an adapiform; a species of Ekgmowechashala.

Homo naledi[421]

Sp. nov

Valid

Berger et al.

Uncertain

Rising Star Cave

 South Africa

A species of Homo.

Homo tsaichangensis

Sp. nov

Valid

McMenamin et al..

Late Pleistocene

Taiwan

A species of hominin recovered sometime before 2008 by fishermen working in the Penghu Channel (between the Penghu Islands and mainland Taiwan) and described in 2015 by an international team of Japanese, Taiwanese, and Australian scientists.

Myanmarcolobus[422]

Gen. et sp. nov

Valid

Takai et al.

Late Miocene to early Pliocene

 Myanmar

A member of Colobinae. The type species is Myanmarcolobus yawensis.

Necrolemur anadoni[423]

Sp. nov

Valid

Minwer-Barakat, Marigó & Moyà-Solà

Middle Eocene (Robiacian)

 Spain

A member of Omomyidae belonging to the subfamily Microchoerinae; a species of Necrolemur.

Perupithecus[424]

Gen. et sp. nov

Valid

Bond et al.

Probably late Eocene

Yahuarango Formation

 Peru

A member of Anthropoidea of uncertain phylogenetic placement, probably a member of Platyrrhini. The type species is Perupithecus ucayaliensis.

Pliobates[425]

Gen. et sp. nov

Valid

Alba et al.

Miocene

 Spain

An ape. The type species is Pliobates cataloniae.

Other eutherians
Name Novelty Status Authors Age Unit Location Notes Images

Altacreodus[426]

Gen. et comb. nov

Valid

Fox

Late Cretaceous

 Canada
 United States

A basal member of Eutheria; a new genus for "Cimolestes" magnus Clemens & Russell (1965).

Ambilestes[426]

Gen. et comb. nov

Valid

Fox

Late Cretaceous

 Canada

A basal member of Eutheria; a new genus for "Cimolestes" cerberoides Lillegraven (1969).

Carodnia inexpectans[427]

Sp. nov

Valid

Antoine et al.

Early Eocene

Mogollón Formation

 Peru

A member of Xenungulata, a species of Carodnia.

Chilestylops[428]

Gen. et sp. nov

Valid

Bradham et al.

Tinguirirican

Abanico Formation

 Chile

A notostylopid notoungulate. The type species is Chilestylops davidsoni.

Galerix wesselsae[429]

Sp. nov

Valid

Zijlstra & Flynn

Miocene

 Pakistan

A member of Erinaceidae (a relative of gymnures and hedgehogs); a species of Galerix.

Hilarcotherium[430]

Gen. et sp. nov

Valid

Vallejo-Pareja et al.

Miocene

La Victoria Formation

 Colombia

An astrapotheriid astrapothere. The type species is Hilarcotherium castanedaii.

Kerberos[431]

Gen. et sp. nov

Valid

Solé et al.

Eocene (Bartonian)

Formation des Molasses de Saix et de Lautrec

 France

A hyainailourine hyaenodont. The type species is Kerberos langebadreae.

Lacrimodon[432]

Gen. et sp. nov

Valid

Van den Hoek Ostende and Fejfar

Early Miocene

 Czech Republic

A member of Eulipotyphla related to Dimylus. The type species is Lacrimodon vandermeuleni.

Mimolagus aurorae[433]

Sp. nov

Valid

Fostowicz-Frelik et al.

Early Middle Eocene

Irdin Manha Formation

 China

A member of Glires belonging to the family Mimotonidae, a species of Mimolagus.

Nesodon taweretus[434]

Sp. nov

Valid

Forasiepi et al.

Early Miocene

Aisol Formation

 Argentina

A toxodontid, a species of Nesodon.

Paedotherium kakai[435]

Sp. nov

Valid

Reguero et al.

Miocene (Huayquerian)

Palo Pintado Formation

 Argentina

A hegetotheriid notoungulate, a species of Paedotherium.

Pampahippus secundus[436]

Sp. nov

Valid

Deraco & García-López

Eocene

Lumbrera Formation

 Argentina

A member of Toxodonta, a species of Pampahippus.

Periphragnis vicentei[428]

Sp. nov

Valid

Bradham et al.

Tinguirirican

Abanico Formation

 Chile

A member of Toxodonta belonging or related to the family Homalodotheriidae; a species of Periphragnis.

Plagioctenodon thewisseni[437]

Sp. nov

Valid

Manz & Bloch

Paleocene (Clarkforkian)

Willwood Formation

 United States

A member of Nyctitheriidae, a species of Plagioctenodon.

Preregidens[438]

Gen. et sp. nov

Valid[439]

Solé, Falconnet & Vidalenc

Eocene

 France

A member of Hyaenodonta. The type species is Preregidens langebadrae.

Prohegetotherium malalhuense[440]

Sp. nov

Valid

Cerdeño & Reguero

Oligocene (Deseadan)

 Argentina

A hegetotheriid notoungulate, a species of Prohegetotherium.

Prolagus latiuncinatus[441]

Sp. nov

Valid

Angelone & Čermák

Late Miocene

 Hungary

A member of Lagomorpha, a species of Prolagus.

Prolagus pannonicus[441]

Sp. nov

Valid

Angelone & Čermák

Late Miocene

 Hungary

A member of Lagomorpha, a species of Prolagus.

Protypotherium sinclairi[404]

Sp. nov

Valid

Kramarz, Bond & Arnal

Early Miocene

Cerro Bandera Formation

 Argentina

An interatheriid notoungulate, a species of Protypotherium.

Ricardocifellia[442]

Nom. nov

Valid

Mones

Paleogene (Itaboraian)

 Brazil

A didolodontid "condylarth"; a replacement name for Paulacoutoia Cifelli, 1983 and Depaulacoutoia Cifelli & Ortiz-Jaureguizar, 2014 (both preoccupied).

Scollardius[426]

Gen. et comb. nov

Valid

Fox

Late Cretaceous

 Canada
 United States

A basal member of Eutheria; a new genus for "Cimolestes" propalaeoryctes Lillegraven (1969).

Seuku[443]

Gen. et comb. nov

Valid

Beatty & Cockburn

Oligocene

 United States

A desmostylian; a new genus for "Behemotops" emlongi Domning, Ray & McKenna (1986).

Sigynorum[444]

Gen. et sp. nov

Valid[445]

McComas & Eberle

Paleocene (early Puercan)

Fort Union Formation

 United States

A member of Arctocyonidae. The type species is Sigynorum magnadivisus.

Sinclairella simplicidens[446]

Sp. nov

Valid

Czaplewski & Morgan

Late Oligocene (Arikareean)

Crystal River Formation

 United States

A member of Apatemyidae, a species of Sinclairella.

Other animals

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Aegirocassis[448][449]

Gen. et sp. nov

Valid

Van Roy, Daley & Briggs

Early Ordovician

 Morocco

A member of Anomalocarididae (a group of animals of uncertain phylogenetic placement, most likely stem-arthropods). The type species is Aegirocassis benmoulae.

Allonnia tintinopsis[450]

Sp. nov

Valid

Bengtson & Collins

Cambrian

Burgess Shale

 Canada

A chancelloriid, a species of Allonnia.

Archiasterella coriacea[450]

Sp. nov

Valid

Bengtson & Collins

Cambrian

Burgess Shale

 Canada

A chancelloriid, a species of Archiasterella.

Banffia episoma[451]

Sp. nov

In press

Conway Morris et al.

Cambrian

Spence Shale

 United States

An early deuterostome, a species of Banffia.

Collinsium[452]

Gen. et sp. nov

Valid

Yang et al.

Early Cambrian

Xiaoshiba Lagerstätte

 China

A lobopodian belonging to the lineage leading to the velvet worms. The type species is Collinsium ciliosum.

Colospongia cribrifera[453]

Sp. nov

Valid

Senowbari-Daryan, Link & Riedel

Late Triassic (Norian)

 Turkey

A sponge, a species of Colospongia.

Costatulites kimi[454]

Sp. nov

Valid

Ghavidel-Syooki et al.

Ordovician (Katian)

 Iran

A member of Tentaculita (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Costatulites.

Cyrtograptus fui[455]

Sp. nov

Valid

Wang et al.

Silurian (Llandovery)

 China

A graptolite, a species of Cyrtograptus.

Eocyathispongia[456]

Gen. et sp. nov

Valid

Yin et al.

Precambrian (Ediacaran)

Doushantuo Formation

 China

A sponge. The type species is Eocyathispongia qiania.

Eokinorhynchus[457]

Gen. et sp. nov

Valid

Zhang et al.

Cambrian (Fortunian)

Dengying Formation

 China

A kinorhynch-like scalidophoran. The type species is Eokinorhynchus rarus.

Eolarva[458]

Gen. et sp. nov

Valid

Zhang & Dong

Cambrian (Fortunian)

Kuanchuanpu Formation

 China

An animal of uncertain phylogenetic placement, possibly a cnidarian-grade animal. The type species is Eolarva kuanchuanpuensis.

Galeactena[459]

Gen. et sp. nov

Valid

Ou et al.

Early Cambrian

Heilinpu Formation

 China

A comb jelly. The type species is Galeactena hemispherica.

Gamascolex vanroyi[460]

Sp. nov

Valid

Gutiérrez-Marco & García-Bellido

Late Ordovician

 Morocco

A palaeoscolecid worm, a species of Gamascolex.

Gemmactena[459]

Gen. et sp. nov

Valid

Ou et al.

Early Cambrian

Heilinpu Formation

 China

A comb jelly. The type species is Gemmactena actinala.

Glyptograptus perneri[461]

Sp. nov

Valid

Štorch

Silurian

 Czech Republic

A graptolite belonging to the family Retiolitidae, a species of Glyptograptus.

Guanshanchaeta[462]

Gen. et sp. nov

Valid

Liu et al.

Early Cambrian

Wulongqing Formation

 China

A polychaete annelid. The type species is Guanshanchaeta felicia.

Hindenites parkheadensis[463]

Sp. nov

Valid

Eriksson & von Bitter

Silurian (Wenlock)

Eramosa Lagerstätte

 Canada

A polychaete, a species of Hindenites.

Neomonograptus aequabilis latus[464]

Subsp. nov

Valid

Chen et al.

Early Devonian

 China

A graptolite, a subspecies of Neomonograptus aequabilis.

Normalograptus frydai[461]

Sp. nov

Valid

Štorch

Silurian

 Czech Republic

A graptolite belonging to the family Normalograptidae, a species of Normalograptus.

Normalograptus parvus[461]

Sp. nov

Valid

Štorch

Silurian

 Czech Republic

A graptolite belonging to the family Normalograptidae, a species of Normalograptus.

Ottoia tricuspida[465]

Sp. nov

Valid

Smith, Harvey & Butterfield

Cambrian

Burgess Shale

 Canada

A stem-group priapulid, a species of Ottoia.

Oymurania[466]

Gen. et sp. nov

Valid

Ushatinskaya in Kouchinsky et al.

Cambrian

 Russia
 Morocco?

A stem-group brachiopod. The type species is Oymurania gravestocki from Siberian Platform; genus might also contain a second, yet-unnamed species from Morocco.

Parastylothalamia minima[453]

Sp. nov

Valid

Senowbari-Daryan, Link & Riedel

Late Triassic (Norian)

 Turkey

A sponge, a species of Parastylothalamia.

Pauxillites thaddei[467]

Sp. nov

Valid

Valent & Carbacho

Ordovician (Floian)

Upper Fezouata Formation

 Morocco

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Pauxillites.

Pernerograptus pribyli[461]

Sp. nov

Valid

Štorch

Silurian

 Czech Republic

A graptolite belonging to the family Monograptidae, a species of Pernerograptus.

Protopholoe colombiana[468]

Sp. nov

Valid

Luque, Hourdez & Vinn

Late Cretaceous (Coniacian)

 Colombia

An aphroditiform polychaete, a species of Protopholoe.

Pseudorthograptus mitchelli[461]

Sp. nov

Valid

Štorch

Silurian

 Czech Republic

A graptolite belonging to the family Retiolitidae, a species of Pseudorthograptus.

Pyrgopolon (Septenaria) nekvasilovae[469]

Sp. nov

Valid

Kočí & Jäger

Late Cretaceous (late Cenomanian)

Bohemian Cretaceous Basin

 Czech Republic

A polychaete belonging to the family Serpulidae, a species of Pyrgopolon.

Pyrgopolon (Septenaria) zitti[469]

Sp. nov

Valid

Kočí & Jäger

Late Cretaceous (late Cenomanian)

Bohemian Cretaceous Basin

 Czech Republic

A polychaete belonging to the family Serpulidae, a species of Pyrgopolon.

Quasimolites[470]

Gen. et sp. nov

Valid

Valent et al.

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is Quasimolites quasimodo.

Rollinschaeta[471]

Gen. et sp. nov

Valid

Parry et al.

Late Cretaceous (Cenomanian)

 Lebanon

A polychaete belonging to the family Amphinomidae. The type species is Rollinschaeta myoplena.

Sarmentofascis? digitatus[472]

Sp. nov

Valid

Schlagintweit & Krajewski

Late Jurassic-Early Cretaceous (Tithonian-early Berriasian)

Crimean Peninsula

A cladocoropsid stromatoporoid sponge, possibly a species of Sarmentofascis.

Scathascolex[473]

Gen. et sp. nov

Valid[474]

Smith

Cambrian

Burgess Shale

 Canada

A palaeoscolecid worm. The type species is Scathascolex minor.

Skryjelites[470]

Gen. et sp. nov

Valid

Valent et al.

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is Skryjelites auritus.

Sphenoecium wheelerensis[475]

Sp. nov

Valid[476]

Maletz & Steiner

Cambrian

Wheeler Shale

 United States

A graptolite, a species of Sphenoecium.

Stipesia[477]

Gen. et sp. nov

Valid[478]

Świerczewska-Gładysz

Late Cretaceous (early Campanian)

 Poland

A lithistid demosponge belonging to the family Pleromidae. The type species is Stipesia belchatowiensis.

Thaumactena[459]

Gen. et sp. nov

Valid

Ou et al.

Early Cambrian

Heilinpu Formation

 China

A comb jelly. The type species is Thaumactena ensis.

Thylacocercus[451]

Gen. et sp. nov

In press

Conway Morris et al.

Cambrian

Wheeler Formation

 United States

A vetulicystid deuterostome. The type species is Thylacocercus ignota.

Triplicatella papilio[479]

Sp. nov

Valid

Kouchinsky in Kouchinsky et al.

Early Cambrian

Emyaksin Formation

 Russia

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Triplicatella.

Uncinatograptus jadae[464]

Sp. nov

Valid

Chen et al.

Early Devonian

 China

A graptolite, a species of Uncinatograptus.

Wiwaxia herka[480]

Sp. nov

Valid

Conway Morris et al.

Cambrian

Spence Shale

 United States

A stem-lophotrochozoan, a species of Wiwaxia.

Yuknessia stephenensis[481]

Sp. nov

Valid

LoDuca et al.

Cambrian

Burgess Shale
Pierson Cove Formation
Spence Shale
Wheeler Shale

 Canada
 United States

A species of Yuknessia (originally thought to be an alga, reinterpreted as a member of Pterobranchia by LoDuca et al., 2015).[481]

Other organisms

Name Novelty Status Authors Age Unit Location Notes Images

Adeliesphaera[482]

Gen. et sp. nov

Valid

Bijl & Brinkhuis

Early Eocene

Antarctica

A dinoflagellate. The type species is Adeliesphaera ohanlonii.

Annulatubus[483]

Gen. et sp. nov

Valid

Carbone et al.

Ediacaran

Blueflower Formation

 Canada

A tubular organism of uncertain phylogenetic placement. The type species is Annulatubus flexuosus.

Caledonicratis[484]

Gen. et sp. nov

Valid

Zapalski & Clarkson

Carboniferous (Viséan)

Gullane Formation

 United Kingdom

An organism of uncertain phylogenetic placement; possibly an animal belonging to the cnidarian family Solanderiidae (the possibility considered to be more probable by the authors of its description) or an alga similar to Perissothallus. The type species is Caledonicratis caridum.

Casparyotorula[485]

Gen. et comb et sp. nov

Valid

Rikkinen, Schmidt & Kettunen in Kettunen et al.

Eocene

 Germany
 Russia (Kaliningrad Oblast)

A hyphomycete fungus known from Baltic amber and Bitterfeld amber. A new genus for "Torula" globulifera Caspary (1886); genus also contains "Torula" heteromorpha Caspary (1886), as well as the new species Casparyotorula arnoldii.

Circulodinium? laeve[486]

Sp. nov

Valid

Cheng et al.

Eocene

Anjihaihe Formation

 China

A dinoflagellate, possibly a species of Circulodinium.

Cyclostephanos ramosus[487]

Sp. nov

Valid

Hayashi & Tanimura

Pleistocene

   Nepal

A diatom, a species of Cyclostephanos.

Cyclotella paleo-ocellata[488]

Sp. nov

Valid

Vossel & Van de Vijver in Vossel et al.

Holocene

 Israel

A diatom. Originally described as a species of Cyclotella; transferred to the genus Lindavia by Nakov et al. (2015).[489]

Donskinica[490]

Gen. nov

Valid

Strelnikova & Kozyrenko

A diatom.

Haslea antiqua[491]

Sp. nov

Valid

Fenner

Paleocene

Southwestern Pacific Ocean (Campbell Plateau)

A diatom, a species of Haslea.

Kaiwaradinium abbreviatum[486]

Sp. nov

Valid

Cheng et al.

Eocene

Anjihaihe Formation

 China

A dinoflagellate, a species of Kaiwaradinium.

Kuckaraukia[492]

Gen. et sp. nov

Valid

Razumovskiy et al.

Late Vendian

Basa Formation

 Russia

An organism of uncertain phylogenetic placement, might be a unitary organism or as a colony of motionless benthic organisms. The type species is Kuckaraukia multituberculata.

Mallomonas schumachii[493]

Sp. nov

Valid

Siver

Eocene

 Canada

A synurophyte, a species of Mallomonas.

Meliolinites buxi[494]

Sp. nov

Valid

Ma & Sun in Ma et al.

Oligocene

Ningming Formation

 China

A fungus belonging to the family Meliolaceae; a species of Meliolinites.

Oligosphaeropsis[486]

Gen. et 3 sp. nov

Valid

Cheng et al.

Eocene

Anjihaihe Formation

 China

A dinoflagellate. The type species is Oligosphaeropsis accreta; genus also contains Oligosphaeropsis complex and Oligosphaeropsis megaprocessa.

Palaeoborrelia[495]

Gen. et sp. nov

Valid

Poinar

Miocene

 Dominican Republic

A spirochaete belonging to the family Spirochaetaceae known from Dominican amber. The type species is Palaeoborrelia dominicana.

Palaeoclaviceps[496]

Gen. et sp. nov

Valid

Poinar, Alderman & Wunderlich

Cretaceous (roughly 100 million years ago)

 Myanmar

A fungus belonging to the family Clavicipitaceae (a relative of ergots). The type species is Palaeoclaviceps parasiticus.

Palaeorickettsia[497]

Gen. et sp. nov

Valid

Poinar

Early Cretaceous (late Albian)

 Myanmar

Probably a member of Rickettsiales. The type species is Palaeorickettsia protera.

Palaeozoosporites[498]

Gen. et sp. nov

Valid

Strullu-Derrien in Strullu-Derrien et al.

Devonian (approximately 407 Mya)

 United Kingdom

A chytrid-like (i.e. similar to members of Blastocladiomycota and Chytridiomycota) fungus of uncertain phylogenetic placement. The type species is Palaeozoosporites renaultii.

Rhoicosphenia gandhii[499]

Sp. nov

Valid

Thomas, Karthick & Kociolek in Thomas, Kociolek & Karthick

Probably Pliocene or Pleistocene

 India

A diatom, a species of Rhoicosphenia.

Rhoicosphenia indica[499]

Sp. nov

Valid

Thomas, Karthick & Kociolek in Thomas, Kociolek & Karthick

Probably Pliocene or Pleistocene

 India

A diatom, a species of Rhoicosphenia.

Rhoicosphenia patrickae[499]

Sp. nov

Valid

Thomas & Kociolek in Thomas, Kociolek & Karthick

Probably Pliocene or Pleistocene

 United States

A diatom, a species of Rhoicosphenia.

Rhoicosphenia reimeri[499]

Sp. nov

Valid

Thomas & Kociolek in Thomas, Kociolek & Karthick

Probably Pliocene or Pleistocene

 United States

A diatom, a species of Rhoicosphenia.

Rhyniosarcina[500]

Gen. et sp. nov

Valid

Taylor & Krings

Early Devonian

 United Kingdom

A member of Cyanobacteria, probably a member or a relative of Chroococcales. The type species is Rhyniosarcina devonica.

Scepasmatocarpion[501]

Gen. et sp. nov

Valid

Krings & Taylor

Early Devonian

 United Kingdom

A fungus of uncertain phylogenetic placement. The type species is Scepasmatocarpion fenestrulatum.

Sekwitubulus[483]

Gen. et sp. nov

Valid

Carbone et al.

Ediacaran

Blueflower Formation

 Canada

A tubular organism of uncertain phylogenetic placement. The type species is Sekwitubulus annulatus.

Sinotubulites hexagonus[502]

Sp. nov

Valid

Cai et al.

Late Ediacaran

Dengying Formation

 China

A tubular organism of uncertain phylogenetic placement, of probable animal affinities; a species of Sinotubulites.

Sinotubulites pentacarinalis[502]

Sp. nov

Valid

Cai et al.

Late Ediacaran

Dengying Formation

 China

A tubular organism of uncertain phylogenetic placement, of probable animal affinities; a species of Sinotubulites.

Sinotubulites triangularis[502]

Sp. nov

Valid

Cai et al.

Late Ediacaran

Dengying Formation

 China

A tubular organism of uncertain phylogenetic placement, of probable animal affinities; a species of Sinotubulites.

Spiniferites adnatus latispinus[486]

Subsp. nov

Valid

Cheng et al.

Eocene

Anjihaihe Formation

 China

A dinoflagellate, a subspecies of Spiniferites adnatus.

Tertiarius juriljii[503]

Sp. nov

Valid

Ognjanova-Rumenova et al.

Pliocene

Vitacevo Formation

 Macedonia

A diatom; a species of Tertiarius.

Tertiarius mariovensis[503]

Sp. nov

Valid

Ognjanova-Rumenova et al.

Pliocene

Vitacevo Formation

 Macedonia

A diatom; a species of Tertiarius.

Tianshandinium[486]

Gen. et sp. nov

Valid

Cheng et al.

Eocene

Anjihaihe Formation

 China

A dinoflagellate. The type species is Tianshandinium biconicum.

Turbiosphaera guersteinae[482]

Sp. nov

Valid

Bijl & Brinkhuis

Early Eocene

Antarctica

A dinoflagellate, a species of Turbiosphaera.

See also

References

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