2012 in archosaur paleontology

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In science: 2009; 2010; 2011; 2012; 2013; 2014; 2015

The year 2012 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur palaeontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2012 in paleontology included various significant developments regarding archosaurs.

This article records new taxa of fossil archosaurs of every kind that have been described during the year 2012, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2012.

Pseudosuchians

Research

The postcranial skeleton of Sebecus icaeorhinus is described by Diego Pol et al. (2012).^[1]
Skull anatomy of Dakosaurus and Plesiosuchus is described by Mark T. Young et al. (2012).^[2]
Kemkemia auditorei, previously thought to be a theropod dinosaur, is reinterpreted as a member of Crocodyliformes by Gabriel Lio et al. (2012).^[3]
A study including a supertree of Crocodyliformes is published by Mario Bronzati, Felipe Chinaglia Montefeltro and Max C. Langer (2012).^[4]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aegisuchus^[5]	Gen. et sp. nov	Valid	Holliday & Gardner	Cenomanian	Kem Kem Formation	Morocco	An aegyptosuchid eusuchian.	Aegisuchus witmeri.
Aetobarbakinoides^[6]	Gen. et sp. nov	Valid	Desojo, Ezcurra & Kischlat	Late Carnian or early Norian	Santa Maria Formation	Brazil	An aetosaur.
Apachesuchus^[7]	Gen. et sp. nov	Valid	Spielmann & Lucas	Late Triassic		United States	An aetosaur. The type species is Apachesuchus heckerti.
Barreirosuchus^[8]	Gen. et sp. nov	Valid	Iori & Garcia	Turonian to Santonian	Adamantina Formation	Brazil	A trematochampsid. The type species is Barreirosuchus franciscoi.
Borealosuchus threeensis^[9]	Sp. nov	Valid	Brochu, Parris, Grandstaff, Denton & Gallagher	Maastrichtian or Danian		USA	A crocodilian from New Jersey.
Bystrowisuchus^[10]	Gen. et sp. nov	Valid	Sennikov	Early Triassic		Russia	A ctenosauriscid. The type species is Bystrowisuchus flerovi.
Crocodylus thorbjarnarsoni^[11]	sp nov	Valid	Brochu & Storrs	Pliocene to Pleistocene		Kenya	A species of Crocodylus.	Crocodylus thorbjarnarsoni.
Diandongosuchus^[12]	Gen. et sp. nov.	Valid	Li et al.	Ladinian	Falang Formation	China	A poposauroid. The type species is Diandongosuchus fuyuanensis.	Diandongosuchus fuyuanensis.
Gasparinisuchus^[13]	Gen. et sp. nov	Valid	Martinelli et al.	Late Cretaceous		Argentina	A peirosaurid. The type species is Gasparinisuchus peirosauroides.

Newly named basal dinosauriforms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Diodorus^[14]	Gen. et sp. nov	Valid	Kammerer, Nesbitt, & Shubin	?Carnian-Norian	Timezgadiouine Formation	Morocco	A silesaurid.

Non-avian dinosaurs

Research

A study including a large phylogenetic analysis of non-coelurosaurian tetanuran theropod dinosaurs is published by Matthew T. Carrano, Roger B. J. Benson and Scott D. Sampson (2012).^[15]
A study including a systematic revision of the family Dromaeosauridae and a large phylogenetic analysis of paravian theropod dinosaurs (both avian and non-avian) is published by Alan Turner, Peter Makovicky and Mark Norell (2012).^[16]
A specimen of Microraptor is described by Quanguo Li et al. (2012), with melanosome imprints making it possible to determine the plumage coloration and iridescence in the plumage.^[17]
A new specimen of Austroraptor cabazai is described by Philip J. Currie and Ariana Paulina Carabajal (2012).^[18]
A specimen of Mei long is described by Chunling Gao et al. (2012).^[19]
A study of Khaan mckennai was published by Amy M. Balanoff and Mark Norell (2012).^[20]
A new information on Yixianosaurus longimanus is published by T. Alexander Dececchi, Hans C. E. Larsson and David W. E. Hone (2012).^[21]
Three specimens of Ornithomimus with evidence of feathers are described by Darla K. Zelenitsky et al. (2012).^[22]
A new information on the anatomy of the holotype specimen of Nqwebasaurus thwazi is published by Jonah N. Choiniere, Catherine A. Forster and William J. de Klerk (2012).^[23]
A study of Alioramus was published by Stephen Brusatte, Thomas D. Carr and Mark Norell (2012).^[24]
Abdominal contents of two specimens of Sinocalliopteryx gigas are described by Lida Xing et al. (2012).^[25]
A study of musculoskeletal anatomy, three-dimensional body proportions and body mass evolution in allosauroid theropod dinosaurs is published by Karl T. Bates, Roger B. J. Benson, and Peter L. Falkingham (2012).^[26]
A study of the braincase of Sinraptor dongi is published by Ariana Paulina Carabajal and Philip J. Currie (2012).^[27]
A study of Kelmayisaurus petrolicus, interpreting it as likely to be a member of Carcharodontosauridae, is published by Stephen L. Brusatte, Roger B.J. Benson and Xing Xu (2012).^[28]
Description of pectoral girdle and forelimb of Majungasaurus crenatissimus is published by Sara H. Burch and Matthew T. Carrano (2012).^[29]
A study of Early Cretaceous Australian theropod dinosaurs was published by Roger B. J. Benson et al. (2012).^[30]
A study including a phylogenetic analysis of titanosauriform sauropod dinosaurs is published by Michael D'Emic (2012).^[31]
The postcranial skeletal pneumaticity in the skeletons of five taxa of early sauropodomorph dinosaurs is described by Adam M. Yates, Mathew J. Wedel and Matthew F. Bonnan (2012).^[32]
The study on the presumed course of the recurrent laryngeal nerve in sauropod dinosaurs is published by Mathew J. Wedel (2012).^[33]
Pachysuchus, previously thought to be an Early Jurassic phytosaur, is reinterpreted as a sauropodomorph dinosaur by Paul M. Barrett and Xu Xing (2012).^[34]
A study of vertebral laminae of sauropod dinosaurs is published by Jeffrey A. Wilson (2012).^[35]
A study on the neural spine bifurcation in diplodocoid sauropod dinosaurs is published by D. Cary Woodruff and Denver W. Fowler (2012).^[36]
A study of the postcranial skeletal pneumaticity in the skeletons of Saltasaurus, Neuquensaurus and Rocasaurus is published by Ignacio A. Cerda, Leonardo Salgado and Jaime E. Powell (2012).^[37]
A study of Early Cretaceous sauropod dinosaurs from North America is published by Michael D. D’Emic and Brady Z. Foreman (2012). Among other findings, additional sauropod material from the Cloverly Formation of Wyoming was referred to Sauroposeidon, Paluxysaurus was synonymized with Sauroposeidon, Rugocaudia was considered a nomen dubium and the cause of the North American sauropod extinction in the middle of the Cretaceous period was discussed.^[38]
The first sauropod dinosaur (a member of Titanosauria) from Antarctica described by Ignacio A. Cerda et al. (2012).^[39]
A study of biomechanics, pectoral girdle articulation and body mass of the Triassic dinosaurs from Brazil (Staurikosaurus, Saturnalia, Pampadromaeus, Guaibasaurus and Unaysaurus) is published by Rafael Delcourt, Sergio A. K. de Azevedo, Orlando N. Grillo and Fernanda O. Deantoni (2012).^[40]
Anatomy of Fruitadens haagarorum is described by Richard J. Butler et al. (2012).^[41]
A study including a phylogenetic analysis of ankylosaurian dinosaurs is published by Richard S. Thompson et al. (2012).^[42]
A study including a phylogenetic analysis of iguanodontian ornithopod dinosaurs is published by Andrew T. McDonald (2012).^[43]
A study of the forearm orientation in hadrosaurids is published by Phil Senter (2012).^[44]
Skin impressions of two different species of Saurolophus are described by Phil Bell (2012).^[45]
A new description of Eolambia caroljonesa is published by Andrew T. McDonald et al. (2012).^[46]
A study of anatomy and relationships of Bolong yixianensis is published by Wu Wenhao and Pascal Godefroit (2012).^[47]
An overview of Early Cretaceous iguanodontian ornithopod dinosaurs of England and Belgium is published by David B. Norman (2012).^[48]
A study of the bone histology of Tenontosaurus tilletti is published by Sarah Werning (2012).^[49]
A study of the bone histology of Dysalotosaurus lettowvorbecki is published by Tom R. Hübner (2012).^[50]
A study questioning the interpretation of Torosaurus as a junior synonym and a growth stage of Triceratops was published by Nicholas R. Longrich and Daniel J. Field (2012).^[51]
A study of some of the earliest known dinosaur assemblages is published by Martín D. Ezcurra (2012). Among other findings, Teyuwasu is interpreted as a member of the clade Dinosauriformes of uncertain phylogenetic placement and a nomen dubium.^[52]
A study of anatomical variability exhibited by major dinosaur groups living during the latest Cretaceous is published by Stephen L. Brusatte et al. (2012).^[53]
A study of biodiversity of late Maastrichtian non-avian dinosaurs is published by Jean Le Loeuff (2012).^[54]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Alnashetri^[55]	Gen. et sp. nov	Valid	Makovicky, Apesteguía & Gianechini	Late Cretaceous (Cenomanian or Turonian)	Candeleros Formation	Argentina	A coelurosaurian, a probable relative of alvarezsaurids. The type species is Alnashetri cerropoliciensis.
Astrophocaudia^[56]	Gen. et sp. nov	Valid^[57]	D’Emic	Early Cretaceous	Trinity Group	United States	A titanosauriform sauropod, a member of the clade Somphospondyli. The type species is Astrophocaudia slaughteri.
Batyrosaurus^[58]	Gen. et sp. nov	Valid	Godefroit et al.	Late Cretaceous		Kazakhstan	A basal hadrosauroid. The type species is Batyrosaurus rozhdestvenskyi.
Bicentenaria^[59]	Gen. et sp. nov	Valid	Novas et al.	Cenomanian	Candeleros Formation	Argentina	A coelurosaur. The type species is Bicentenaria argentina.
Bonapartenykus^[60]	Gen. et sp. nov	Valid	Agnolin, Powell, Novas, & Kundrát	Campanian/Maastrichtian	Allen Formation	Argentina	An alvarezsaurid.
Comahuesaurus^[61]	Gen. et sp. nov	Valid	Carballido et al.	Early Cretaceous (Aptian or Albian)	Lohan Cura Formation	Argentina	A rebbachisaurid sauropod. The type species is Comahuesaurus windhauseni.
Coronosaurus^[62]	Gen. et comb. nov	Valid	Ryan, Evans & Shepherd	Late Cretaceous	Oldman Formation	Canada	A centrosaurine ceratopsian, a new genus for "Centrosaurus" brinkmani Ryan & Russell (2005).
Darwinsaurus^[63]	Gen. et sp. nov	Valid	Paul	Early Cretaceous (early Valanginian)	Wadhurst Clay Formation	United Kingdom	A styracosternan ornithopod. The type species is Darwinsaurus evolutionis, and the holotype is an associated skeleton that includes material catalogued under numbers NHMUK R1831 (a dentary), R1833, R1835 and R1836;^[63] this skeleton is assigned by Norman (2010) to Hypselospinus fittoni.^[64]
Elaltitan^[65]	Gen. et sp. nov	Valid	Mannion & Otero	Middle Cenomanian to Turonian	Bajo Barreal Formation	Argentina	A lithostrotian titanosaur. The type species is Elaltitan lilloi.
Eoabelisaurus^[66]	Gen. et sp. nov	Valid	Pol & Rauhut	Aalenian or Bajocian	Cañadon Asfalto Formation	Argentina	An abelisaurid. The type species is Eoabelisaurus mefi.
Gideonmantellia^[67]	Gen. et sp. nov	Valid	Ruiz-Omeñaca et al.	Barremian	Camarillas Formation	Spain	A basal ornithopod. The type species is Gideonmantellia amosanjuanae.
Gryphoceratops^[68]	Gen. et sp. nov	Valid	Ryan, Evans, Currie, Brown, & Brinkman	Late Santonian	Milk River Formation	Canada	A leptoceratopsid ceratopsian.
Hexing^[69]	Gen. et sp. nov	Valid	Jin, Chen & Godefroit	Early Valanginian to early Barremian	Yixian Formation	China	A basal ornithomimosaur. The type species is Hexing qingyi.
Huehuecanauhtlus^[70]	Gen. et sp.	Valid	Ramírez-Velasco et al..	Santonian		Mexico	A basal hadrosauroid. The type species is Huehuecanauhtlus tiquichensis.	Huehuecanauhtlus.
Huxleysaurus^[63]	Gen. et com. nov	Valid	Paul	Early Cretaceous (early Valanginian)	Wadhurst Clay Formation	United Kingdom	A styracosternan ornithopod, a new genus for "Iguanodon" hollingtoniensis Lydekker (1889);^[63] this species is considered by Norman (2010) to be a junior synonym of Hypselospinus fittoni.^[64]
Ichthyovenator^[71]	Gen. et sp. nov	Valid	Allain et al.	Late Early Cretaceous	Savannakhet Basin	Laos	A spinosaurid. The type species is Ichthyovenator laosensis.
Kaatedocus^[72]	Gen. et sp. nov	Valid^[73]	Tschopp & Mateus	Late Jurassic (Kimmeridgian)	Morrison Formation	United States	A diplodocine diplodocid sauropod. The type species is Kaatedocus siberi.
Kundurosaurus^[74]	Gen. et sp. nov	Valid	Godefroit, Bolotsky & Lauters	Late Cretaceous, possibly late Maastrichtian	Udurchukan Formation	Russia	A saurolophine hadrosaurid. The type species is Kundurosaurus nagornyi.	Kundurosaurus.
Lapampasaurus^[75]	Gen. et sp. nov	Valid	Coria, Riga & Casadío	Late Cretaceous (late Campanian or early Maastrichthian)	Allen Formation	Argentina	A hadrosaurid. The type species is Lapampasaurus cholinoi.
Latirhinus^[76]	Gen. et sp. nov	Valid	Prieto-Márquez & Brañas	Late Cretaceous (late Campanian)	Cerro del Pueblo Formation	Mexico	A saurolophine hadrosaurid. The type species is Latirhinus uitstlani.
Magnapaulia^[77]	Gen. et comb. nov	Valid	Prieto-Márquez, Chiappe & Joshi	Late Campanian	El Gallo Formation	Mexico	A lambeosaurine hadrosaurid, a new genus for "Lambeosaurus" laticaudus (Morris, 1981).	Magnapaulia.
Mantellodon^[63]	Gen. et sp. nov	Valid	Paul	Early Cretaceous (early Aptian)	Lower Greensand Formation	United Kingdom	A styracosternan ornithopod. The type species is Mantellodon carpenteri and the holotype is the specimen NHMUK R3741 consisting of partial associated skeleton (also known as Gideon Mantell's "Mantel-piece")^[63]
Martharaptor^[78]	Gen. et sp. nov	Valid	Senter, Kirkland & DeBlieux	Early Aptian	Cedar Mountain Formation	United States	A possible therizinosauroid. The type species is Martharaptor greenriverensis.	Martharaptor.
Microraptor hanqingi^[79]	sp nov	Valid	Gong et al.	Early Cretaceous		China	A species of Microraptor. Its status as a distinct species is disputed, with Pei et al. (2014) considering the holotype specimen's morphology not to be distinguishable from other Microraptor specimens.^[80]
Mochlodon vorosi^[81]	Sp. nov	Valid	Ősi et al.	Santonian	Csehbánya Formation	Hungary	A rhabdodontid ornithopod, a species of Mochlodon.
Ningyuansaurus^[82]	Gen. et sp. nov	Valid	Ji et al.	Early Cretaceous	Yixian Formation	China	An oviraptorosaur. The type species is Ningyuansaurus wangi.
Ostafrikasaurus^[83]	Gen. et sp. nov	Valid	Buffetaut	Late Jurassic	Tendaguru Formation	Tanzania	A spinosaurid. The type species is Ostafrikasaurus crassiserratus.
Pachyrhinosaurus perotorum^[84]	sp nov	Valid	Fiorillo & Tykoski	Maastrichtian	Prince Creek Formation	USA	Third known species of Pachyrhinosaurus.
Pegomastax^[85]	Gen. et sp. nov	Valid	Sereno	Hettangian to Sinemurian	Elliot Formation	South Africa	A heterodontosaurid. The type species is Pegomastax africana.
Philovenator^[86]	Gen. et sp. nov	Valid	Xu et al.	Campanian	Wulansuhai Formation	China	A troodontid. Its type species is Philovenator curriei.
Proa^[87]	Gen. et sp. nov	Valid	McDonald et al.	Early Cretaceous (early Albian)	Escucha Formation	Spain	An iguanodont ornithopod closely related to Iguanodon and the clade Hadrosauroidea. The type species is Proa valdearinnoensis.
Rugocaudia^[88]	Gen. et sp. nov	Valid	Woodruff	Aptian or Albian	Cloverly Formation	USA	A titanosauriform sauropod. The type species is Rugocaudia cooneyi.
Sciurumimus^[89]	Gen. et sp. nov	Valid	Rauhut et al.	Upper Kimmeridgian		Germany	A megalosauroid. The type species is Sciurumimus albersdoerferi.	Sciurumimus.
Unescoceratops^[68]	Gen. et sp. nov	Valid	Ryan, Evans, Currie, Brown, & Brinkman	Late Campanian	Dinosaur Park Formation	Canada	A leptoceratopsid ceratopsian.
Xenoceratops^[62]	Gen. et sp. nov	Valid	Ryan, Evans & Shepherd	Late Cretaceous (middle Campanian)	Foremost Formation	Canada	A centrosaurine ceratopsian. The type species is Xenoceratops foremostensis.
Yueosaurus^[90]	Gen. et sp. nov	Valid	Zheng, Jin, Shibata, Azuma, & Yu	Aptian-Cenomanian	Liangtoutang Formation	China	A basal ornithopod.
Yurgovuchia^[91]	Gen. et sp. nov	Valid	Senter et al..	Early Cretaceous, possibly Barremian	Cedar Mountain Formation	USA	A dromaeosaurid. The type species is Yurgovuchia doellingi.	Yurgovuchia.
Yutyrannus^[92]	Gen. et sp. nov	Valid	Xu et al..	Early Cretaceous	Yixian Formation	China	A basal tyrannosauroid. The type species is Yutyrannus huali.	Yutyrannus.

Newly named birds

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aegolius gradyi ^[93]	Sp. nov.	Valid	Storrs L. Olson	Pleistocene to Holocene		Bermuda	A Strigidae.
Aegypius jinniushanensis ^[94]	Sp. nov.	Valid	Zhang Zihui Huang Yunping Helen F. James Hou Lianhai	Middle Pleistocene		China	An Accipitridae.
Aix praeclara ^[95]	Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Middle Miocene	Oshin Formation	Mongolia	An Anatidae.
Alauda xerarvensis ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian	Bulgaria	An Alaudidae.
Anchigyps voorhiesi ^[97]	Gen. nov. et Sp. nov.	Valid	Zhang Zihui Alan Feduccia Helen F. James	Late Miocene. late Clarendonian	Ash Hollow Formation	USA: Nebraska	An Accipitridae close in morphology to the Old World Vultures. The type species of the new genus.
Anhinga walterbolesi ^[98]	Sp. nov.	Valid	Trevor H. Worthy	Late Oligocene or Early Miocene	Etadunna Formation, 24-26 Ma	Australia: South Australia	An Anhingidae.
Anthus antecedens ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7-8	Hungary	A Motacillidae.
Athene vallgornerensis ^[101]	Sp. nov.	Valid	Carmen Guerra Pere Bover Josep A. Alcover	Early Pleistocene	Cave deposits	Spain: Majorca	A Strigidae.
Australopicus nelsonmandelai ^[102]	Gen. nov. et Sp. nov.	Valid	Albrecht Manegold Antoine Louchart	Early Pliocene	Varswater Formation	South Africa	A Picidae, this is the type species of the new genus.
Avolatavis tenens ^[103]	Gen. nov. et Sp. nov.	Valid	Daniel T. Ksepka Julia A. Clarke	Eocene	Green River Formation	USA: Wyoming	A stem parrot, a member of Pan-Psittaciformes. The type species of the new genus.
Becassius charadriioides ^[104]	Gen. nov. et Sp. nov.	Valid	Vanesa L. De Pietri Gerald Mayr	Early Miocene	Saint-Gérard-le-Puy, MN 2a	France	A Scolopacidae or a relative of the Scolopacidae. The type species of the new genus.
Bombycilla hamori ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5	Hungary	A Bombycillidae.
Brodavis americanus ^[105]	Gen. nov. et Sp. nov.	Valid	Larry D. Martin Evgeny N. Kurochkin Tim T. Tokaryk	Late Cretaceous,	Maastrichtian Frenchman Formation	Canada: Saskatchewan	A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012, this is the type species of the new genus.
Brodavis baileyi ^[105]	Sp. nov.	Valid	Larry D. Martin Evgeny N. Kurochkin Tim T. Tokaryk	Late Cretaceous,	Maastrichtian Hell Creek Formation	USA: South Dakota	A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012.
Brodavis mongoliensis ^[105]	Sp. nov.	Valid	Larry D. Martin Evgeny N. Kurochkin Tim T. Tokaryk	Late Cretaceous,	Maastrichtian Nemegt Formation	Mongolia	A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012.
Certhia janossyi ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Late Miocene	MN 9	Hungary	A Certhiidae.
Chenoanas deserta ^[106]	Gen. nov. et Sp. nov.	Valid	Nikita V. Zelenkov	Middle Miocene	Oshin Formation	Mongolia	An Anatidae, this is the type species of the new genus.
Cinclus major ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5	Hungary	A Cinclidae.
Circaetus rhodopensis ^[107]	Sp. nov.	Valid	Zlatozar Boev	Late Miocene	Tulorian = Late Meotian, Late MN 11, ca. 7,5 Mya	Bulgaria	An Accipitridae.
Crypturellus reai ^[108]	Sp. nov.	Valid	Robert M. Chandler	Miocene (Santacrucian)	Santa Cruz Formation	Argentina	A Tinamidae.
?Elorius limosoides ^[104]	Sp. nov.	Valid	Vanessa L. De Pietri Gerald Mayr	Early Miocene	Saint-Gérard-le-Puy, MN 2a	France	A Scolopacidae, possibly a species of Elorius.
Emberiza bartkoi ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5	Hungary	An Emberizidae.
Eremarida xerophila ^[96]	Gen. nov. et Sp. nov.	Valid	Zlatozar Boev	Late Miocene	Gnilyane Formation	Bulgaria	An Alaudidae, this is the type species of the new genus.
Eremophila prealpestris ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian, MP 17, 2,23-2,40 Mya	Bulgaria	An Alaudidae.
Erithacus horusitzkyi ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8	Hungary	A Muscicapidae, an Erithacus robin.
Galerida bulgarica ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian	Bulgaria	An Alaudidae.
Galerida cserhatensis ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5	Hungary	An Alaudidae.
Heteroanser vicinus ^[109]	Gen. nov. et Comb. nov.	Valid	Nikita V. Zelenkov	Late Miocene	Hyargas Nuur Formation	Mongolia	An Anatidae a new genus for "Heterochen" vicinus Kurochkin, 1976.
Hirundo selengenica ^[110]	Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Late Pliocene	Chikoi Formation	Mongolia	A Hirundinidae.
Jeholornis palmapenis ^[111]	Sp. nov.	Valid	Jingmai K. O'Connor Sun Chengkai Xu Xing Wang Xiaolin Zhou Zhonghe	Early Aptian	Jiufotang Formation	China	A Jeholornithidae Zhou et Zhang, 2006.
Kairuku grebneffi ^[112]	Sp. nov.	Valid	Daniel T. Ksepka R. Ewan Fordyce Tatsuro Ando Craig M. Jones	Oligocene	Dutroonian	New Zealand	A Spheniscidae.
Kairuku waitaki ^[112]	Gen. nov. et Sp. nov	Valid	Daniel T. Ksepka R. Ewan Fordyce Tatsuro Ando Craig M. Jones	Oligocene	Late Whaingaroan or Early Duntroonian	New Zealand	A Spheniscidae, the type species of the genus.
Lanius schreteri ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7-8	Hungary	A Laniidae.
Leptoptilos lüi ^[113]	Sp. nov.	Not valid	Zhang Zihui Huang Yunping Helen F. James Hou Lianhai	Middle Pleistocene	Cave deposits	China	A Ciconiidae, the name is not valid because of the "ü" in it, that must, of course be a plain "u".
Lullula balcanica ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian	Bulgaria	An Alaudidae.
Lullula neogradensis ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8	Hungary	An Alaudidae.
Lullula slivnicensis ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Early Late Villafranchian, MN 18	Bulgaria	An Alaudidae, a relative of the woodlark.
Luscinia praeluscinia ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5	Hungary	A Muscicapidae.
Lutetodontopteryx tethyensis ^[114]	Gen. nov. et Sp. nov.	Valid	Gerald Mayr Evgenij Zvonok	Middle Eocene	Early Lutetian	Ukraine	A Pelagornithidae, this is the type species of the new genus.
Melanocorypha donchevi ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian	Bulgaria	An Alaudidae.
Melanocorypha serdicensis ^[96]	Sp. nov.	Valid	Zlatozar Boev	Late Miocene	Gnilyane Formation, Middle Late Pontian	Bulgaria	An Alaudidae.
Mioquerquedula minutissima ^[95]	Gen. nov. et Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Middle Miocene, possibly also Late Miocene	Gobi-Altai Aimag, Shargyn Govi, Sharga location	Mongolia	An Anatidae, this is the type species of the new genus, the new genus also contains "Anas" velox Milne-Edwards, 1868 as a Comb. nov. from France, Germany, Romania and ?Hungary.
Muscicapa leganyii ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8	Hungary	A Muscicapidae.
Otus mauli ^[115]	Sp. nov.	Valid	Carlos Rando Harald Pieper Josep A. Alcover Storrs L. Olson	Quaternary		Portugal: Madeira	A Strigidae.
?Parvelorius calidris ^[104]	Gen. nov. et Sp. nov.	Valid	Vanessa L. De Pietri Gerald Mayr	Early Miocene	MN 2a	France	A Scolopacidae, the new genus is erected for the species "Tringa" gracilis Milne-Edwards, 1868, the type species of the new genus and makes a Com. nov. with it; genus may also contain this new species.
Phylloscopus miocaenicus ^[99]	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 6-8	Hungary	A Phylloscopidae.
Picavus litencicensis ^[116]	Gen. nov. et Sp. nov.	Valid	Gerald Mayr Růžena Gregorová	Early Oligocene	Menilite Formation	Czech Republic	A stem group representative of the Piciformes, placed in a new family Picavidae Mayr et Gregorová, 2012. it is the type species of the new genus.
Pipilo naufragus ^[117]	Sp. nov.	Valid	Storrs L. Olson David B. Wingate	Middle Pleistocene to Holocene	Cave deposits	Bermuda	A Passerellidae, towhee.
Pliocalcarius orkhonensis ^[110]	Gen. nov. et Sp. nov	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Late Pliocene	Villafranchian, Chikoi Formation, MN 16a	Mongolia	A probable close relative of longspurs, Calcariidae. The type species of the new genus.
Praealauda hevesensis ^[96]^[99]^[100]	Gen. nov. et Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8	Hungary	An Alaudidae. The type species of the new genus.
Primophaps schoddei ^[118]	Gen. nov. et Sp. nov.	Valid	Trevor H. Worthy	Late Oligocene to early Miocene	Riversleigh	Australia Queensland	A Columbidae related to the bronzewing pigeons. The type species of the new genus.
Pterocnemia mesopotamica ^[119]	Sp. nov.	Valid	Federico L. Agnolín Jorge I. Noriega	Late Miocene	Ituzaingó Formation, Huayquerian	Argentina	A Rheidae.
Rallicrex litkensis ^[120]	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5	Hungary	A Rallidae, a new species of Rallicrex Lambrecht, 1933, the type species is Rallicrex kolozsvarensis Lambrecht, 1933.
Rhodospiza shaamarica ^[110]	Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Late Pliocene	Villafranchian, Chikoi Formation, MN 16a	Mongolia	A Fringillidae, Carduelinae.
Sanshuiornis zhangi ^[121]	Gen. nov. et Sp. nov.	Valid	Wang Min Gerald Mayr Zhang Jiangyong Zhou Zhonghe	Middle Eocene	Huayong Formation	China	A Ciconiiformes Incertae Sedis.
Schizooura lii ^[122]	Gen. nov. et Sp. nov	Valid	Zhou Shuang Zhou Zhonghe Jingmai K. O’Connor	Aptian	Jiufotang Formation	China	A basal Ornithuromorpha. The type species of the new genus.
Scolopacimilis lartetianus ^[104]	Gen. nov. et Comb. nov.	Valid	Vanessa L. De Pietri Mayr	Early Miocene	MN 2a	France	A Scolopacidae, a new genus for the species "Totanus" lartetianus, the type species of the genus, creating a Comb. nov.
Shengjingornis yangi ^[123]	Gen. nov et Sp. nov.	Valid	Li Li Wang Jinqi Zhang Xi Hou Shilin	Early Cretaceous	Jiufotang Formation	China	A Longipterygidae Zhang, Zhou, Hou L. H. et Gu, 2000, Enantiornithes Walker, 1981. The type species of the new genus.
Songzia acutunguis ^[124]	Sp. nov.	Valid	Wang Min Gerald Mayr Zhang Jiangyong Zhou Zhonghe	Early Eocene	Yangxi Formation	China	A Songziidae Hou, 1990, Gruiformes. The type species is Songzia heidangkouensis Hou, 1990.
Sturnus kretzoii ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Late Miocene	MN 9	Hungary	A Sturnidae.
Tadorna minor ^[120]	Sp. nov.	Valid	Jenö Kessler János Hír	Miocene	MN 7/8	Hungary	An Anatidae.
Turdicus minor ^[99]^[100]	Sp. nov.	Valid	Jenö Kessler János Hír	Early to Middle Miocene		Hungary	A Turdidae.
Tydea septentrionalis ^[125]	Gen. nov. et Sp. nov.	Valid	Gerald Mayr Thierry Smith	Early Oligocene	Rupelian	Belgium: Antwerp	An Diomedeidae. The type species of the new genus.
Xiangornis shenmi ^[126]	Gen. nov. et Sp. nov.	Valid	Hu Dongyu Xu Xing Hou Lianhai Corwin Sullivan	Early Cretaceous	Jiufotang Formation	China	An Enantiornithes Walker, 1981. The type species of the new genus.

Newly named pterosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aerotitan^[127]	Gen. et sp. nov	Valid	Novas et al.	Late Cretaceous		Argentina	An azhdarchid. The type species is Aerotitan sudamericanus.
Bellubrunnus^[128]	Gen. et sp. nov	Valid	Hone et al.	Late Kimmeridgian	Pfraundorf-Heitzenhofen Basin	Germany	A rhamphorhynchid. The type species is Bellubrunnus rothgaengeri.	Bellubrunnus.
Dendrorhynchoides mutoudengensis^[129]	Sp. nov	Valid	Lü & Hone	Middle Jurassic	Tiaojishan Formation	China	An anurognathid, a species of Dendrorhynchoides.
Europejara^[130]	Gen. et sp. nov	Valid	Vullo et al.	Upper Barremian	La Huérguina Formation	Spain	A tapejarid. The type species is Europejara olcadesorum.	Europejara.
Gladocephaloideus^[131]	Gen. et sp. nov	Valid	Lü, Ji, Wei, & Liu	Barremian	Yixian Formation	China	A gallodactylid ctenochasmatoid.
Guidraco^[132]	Gen. et sp. nov	Valid	Wang, Kellner, Jiang, & Cheng	Early Cretaceous	Jiufotang Formation	China	A close relative of Ludodactylus. The type species is Guidraco venator.	Guidraco.
Jianchangnathus^[133]	Gen. et sp. nov	Valid	Cheng, Wang, Jiang, & Kellner	Middle Jurassic	Tiaojishan Formation	China	A scaphognathid.
Moganopterus^[134]	Gen. et sp. nov	Valid	Lü et al.	Early Cretaceous	Yixian Formation	China	A boreopterid. The type species is Moganopterus zhuiana.
Qinglongopterus^[135]	Gen. et sp.	Valid	Lü, Unwin, Zhao, Gao & Shen.	Middle Jurassic	Tiaojishan Formation	China	A rhamphorhynchid.

Archosauria incertae sedis

Newly named archosaurs with uncertain affinities

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Samrukia^[136]	Gen. et sp. nov	Valid	Naish, Dyke, Cau, Escuillié, Godefroit	Late Cretaceous	Kyzylorda District	Kazakhstan	A possible basal ornithuromorph;^[136] alternatively, it could be a large pterosaur.^[137]^[138]
Smok^[139]	Gen. et sp. nov	Valid	Niedźwiedzki, Sulej & Dzik	Latest Norian/Early Rhaetian	Lipie Śląskie clay-pit	Poland	A large predatory archosaur of uncertain phylogenetic placement, the largest known predatory archosaur from the Late Triassic and Early Jurassic of central Europe.

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↑ José Ignacio Ruiz-Omeñaca, José Ignacio Canudo, Gloria Cuenca-Bescós, Penélope Cruzado-Caballero, José Manuel Gasca and Miguel Moreno-Azanza (2012). "A new basal ornithopod dinosaur from the Barremian of Galve, Spain". Comptes Rendus Palevol 11 (6): 435–444. doi:10.1016/j.crpv.2012.06.001.
1 2 Michael J. Ryan, David C. Evans, Philip J. Currie, Caleb M. Brown and Don Brinkman (2012). "New leptoceratopsids from the Upper Cretaceous of Alberta, Canada". Cretaceous Research 35: 69–80. doi:10.1016/j.cretres.2011.11.018.
↑ Jin Liyong, Chen Jun and Pascal Godefroit (2012). "A New Basal Ornithomimosaur (Dinosauria: Theropoda) from the Early Cretaceous Yixian Formation, Northeast China". In Godefroit, P. (eds). Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Indiana University Press. pp. 467–487. ISBN 978-0-253-35721-2.
↑ Angel Alejandro Ramírez-Velasco, Mouloud Benammi, Albert Prieto-Márquez, Jesús Alvarado Ortega and René Hernández-Rivera (2012). "Huehuecanauhtlus tiquichensis, a new hadrosauroid dinosaur (Ornithischia: Ornithopoda) from the Santonian (Late Cretaceous) of Michoacán, Mexico". Canadian Journal of Earth Sciences 49 (2): 379–395. doi:10.1139/e11-062.
↑ Ronan Allain, Tiengkham Xaisanavong, Philippe Richir and Bounsou Khentavong (2012). "The first definitive Asian spinosaurid (Dinosauria: Theropoda) from the early cretaceous of Laos". Naturwissenschaften 99 (5): 369–377. doi:10.1007/s00114-012-0911-7. PMID 22528021.
↑ Emanuel Tschopp and Octávio Mateus (2013). "The skull and neck of a new flagellicaudatan sauropod from the Morrison Formation and its implication for the evolution and ontogeny of diplodocid dinosaurs". Journal of Systematic Palaeontology 11 (7): 853–888. doi:10.1080/14772019.2012.746589.
↑ http://zoobank.org/NomenclaturalActs/E559FA2A-67AF-4A8E-818F-A3E6999CC3FC
↑ Pascal Godefroit, Yuri L. Bolotsky and Pascaline Lauters (2012). "A New Saurolophine Dinosaur from the Latest Cretaceous of Far Eastern Russia". PLoS ONE 7 (5): e36849. doi:10.1371/journal.pone.0036849. PMC 3364265. PMID 22666331.
↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. doi:10.5710/amgh.9.4.2012.487.
↑ Albert Prieto-Márquez and Claudia Inés Serrano Brañas (2012). "Latirhinus uitstlani, a 'broad-nosed' saurolophine hadrosaurid (Dinosauria, Ornithopoda) from the late Campanian (Cretaceous) of northern Mexico". Historical Biology: An International Journal of Paleobiology 24 (6): 607–619. doi:10.1080/08912963.2012.671311.
↑ Albert Prieto-Márquez, Luis M. Chiappe and Shantanu H. Joshi (2012). "The Lambeosaurine Dinosaur Magnapaulia laticaudus from the Late Cretaceous of Baja California, Northwestern Mexico". PLoS ONE 7 (6): e38207. doi:10.1371/journal.pone.0038207. PMC 3373519. PMID 22719869.
↑ Phil Senter, James I. Kirkland and Donald D. DeBlieux (2012). "Martharaptor greenriverensis, a New Theropod Dinosaur from the Lower Cretaceous of Utah". PLoS ONE 7 (8): e43911. doi:10.1371/journal.pone.0043911. PMC 3430620. PMID 22952806.
↑ En-Pu Gong, Larry D. Martin, David A. Burnham, Amanda R. Falk and Lian-Hai Hou (2012). "A new species of Microraptor from the Jehol Biota of northeastern China". Palaeoworld 21 (2): 81–91. doi:10.1016/j.palwor.2012.05.003.
↑ Rui Pei, Quanguo Li, Qingjin Meng, Ke-Qin Gao and Mark A. Norell (2014). "A new specimen of Microraptor (Theropoda: Dromaeosauridae) from the Lower Cretaceous of western Liaoning, China". American Museum Novitates 3821: 1–28. doi:10.1206/3821.1.
↑ Attila Ősi, Edina Prondvai, Richard Butler and David B. Weishampel (2012). "Phylogeny, Histology and Inferred Body Size Evolution in a New Rhabdodontid Dinosaur from the Late Cretaceous of Hungary". PLoS ONE 7 (9): e44318. doi:10.1371/journal.pone.0044318. PMC 3448614. PMID 23028518.
↑ Ji Qiang, Lü Jun-Chang, Wei Xue-Fang, Wang Xu-Ri (2012). "A new oviraptorosaur from the Yixian Formation of Jianchang, Western Liaoning Province, China". Geological Bulletin of China 31 (12): 2102–2107.
↑ Buffetaut, Eric (2012). "An early spinosaurid dinosaur from the Late Jurassic of Tendaguru (Tanzania) and the evolution of the spinosaurid dentition". Oryctos 10: 1–8.
↑ Anthony R. Fiorillo and Ronald S. Tykoski (2012). "A new species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope (Prince Creek Formation: Maastrichtian) of Alaska". Acta Palaeontologica Polonica 57 (3): 561–573. doi:10.4202/app.2011.0033.
↑ Paul C. Sereno (2012). "Taxonomy, morphology, masticatory function and phylogeny of heterodontosaurid dinosaurs". ZooKeys 226: 1–225. doi:10.3897/zookeys.226.2840. PMC 3491919. PMID 23166462.
↑ Xu Xing, Zhao Ji, Corwin Sullivan, Tan Qing-Wei, Martin Sander and Ma Qing-Yu (2012). "The taxonomy of the troodontid IVPP V 10597 reconsidered" (PDF). Vertebrata PalAsiatica 50 (2): 140–150.
↑ Andrew T. McDonald, Eduardo Espílez, Luis Mampel, James I. Kirkland and Luis Alcalá (2012). "An unusual new basal iguanodont (Dinosauria: Ornithopoda) from the Lower Cretaceous of Teruel, Spain" (PDF). Zootaxa 3595: 61–76.
↑ D. Cary Woodruff (2012). "A new titanosauriform from the Early Cretaceous Cloverly Formation of Montana". Cretaceous Research 36: 58–66. doi:10.1016/j.cretres.2012.02.003.
↑ Oliver W. M. Rauhut, Christian Foth, Helmut Tischlinger and Mark A. Norell (2012). "Exceptionally preserved juvenile megalosauroid theropod dinosaur with filamentous integument from the Late Jurassic of Germany". Proceedings of the National Academy of Sciences of the United States of America 109 (29): 11746–11751. doi:10.1073/pnas.1203238109. PMC 3406838. PMID 22753486.
↑ Zheng, Wenjie; Xingsheng Jin, Masateru Shibata, Yoichi Azuma and Fangming Yu (2012). "A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China". Cretaceous Research 34: 208–219. doi:10.1016/j.cretres.2011.11.001.
↑ Senter, Phil; James I. Kirkland, Donald D. DeBlieux, Scott Madsen and Natalie Toth (2012). "New Dromaeosaurids (Dinosauria: Theropoda) from the Lower Cretaceous of Utah, and the Evolution of the Dromaeosaurid Tail". PLoS ONE 7 (5): e36790. doi:10.1371/journal.pone.0036790. PMC 3352940. PMID 22615813.
↑ Xu, Xing; Kebai Wang, Ke Zhang, Qingyu Ma, Lida Xing, Corwin Sullivan, Dongyu Hu, Shuqing Cheng and Shuo Wang (2012). "A gigantic feathered dinosaur from the Lower Cretaceous of China". Nature 484 (7392): 92–95. doi:10.1038/nature10906. PMID 22481363.
↑ Storrs L. Olson (2012). "A new species of small owl of the genus Aegolius (Aves: Strigidae) from Quaternary deposits on Bermuda". Proceedings of the Biological Society of Washington 125 (2): 97–105. doi:10.2988/11-36.1.
↑ Zihui Zhang, Yunping Huang, Helen F. James and Lianhai Hou (2012). "Two Old World vultures from the middle Pleistocene of northeastern China and their implications for interspecific competition and biogeography of Aegypiinae" (PDF). Journal of Vertebrate Paleontology 32 (1): 117–124. doi:10.1080/02724634.2012.624146.
1 2 N. V. Zelenkov and E. N. Kurochkin (2012). "Dabbling ducks (Aves: Anatidae) from the Middle Miocene of Mongolia". Paleontological Journal 46 (4): 421–429. doi:10.1134/s0031030112040132.
1 2 3 4 5 6 7 8 9 Zlatozar Boev (2012). "Neogene Larks (Aves: Alaudidae (Vigors, 1825)) from Bulgaria" (PDF). Acta Zoologica Bulgarica 64 (3): 295–318.
↑ Zihui Zhang, Alan Feduccia and Helen F. James (2012). "A Late Miocene Accipitrid (Aves: Accipitriformes) from Nebraska and Its Implications for the Divergence of Old World Vultures". PLoS ONE 7 (11): e48842. doi:10.1371/journal.pone.0048842.
↑ Trevor H. Worthy (2012). "A New Species of Oligo-Miocene Darter (Aves: Anhingidae) from Australia". The Auk 129 (1): 96–104. doi:10.1525/auk.2012.11204.
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Jeno Kessler and János Hír (2012). "Észak-Magyarország madárvilága a miocénben. II. rész." (PDF). Földtani közlöny 142 (2): 149–168.
1 2 3 4 5 6 7 8 9 10 11 12 13 14 Eugen Kessler (2013). "Neogene songbirds (Aves, Passeriformes) from Hungary" (PDF). Hantkeniana. Contributions of the Department of Palaeontology. Eötvös University 8: 37–149.
↑ Carmen Guerra, Pere Bover and Josep A. Alcover (2012). "A new species of extinct little owl from the Pleistocene of Mallorca (Balearic Islands)" (PDF). Journal of Ornithology 153 (2): 347–354. doi:10.1007/s10336-011-0749-3.
↑ Albrecht Manegold and Antoine Louchart (2012). "Biogeographic and paleoenvironmental implications of a new woodpecker species (Aves, Picidae) from the early Pliocene of South Africa". Journal of Vertebrate Paleontology 32 (4): 926–938. doi:10.1080/02724634.2012.664597.
↑ Daniel T. Ksepka and Julia A. Clarke (2012). "A new stem parrot from the Green River Formation and the complex evolution of the grasping foot in Pan-Psittaciformes". Journal of Vertebrate Paleontology 32 (2): 395–406. doi:10.1080/02724634.2012.641704.
1 2 3 4 Vanesa L. De Pietri and Gerald Mayr (2012). "An assessment of the diversity of early Miocene Scolopaci (Aves, Charadriiformes) from Saint-Gérand-le-Puy (Allier, France)". Palaeontology 55 (6): 1177–1197. doi:10.1111/j.1475-4983.2012.01182.x.
1 2 3 Larry D. Martin, Evgeny N. Kurochkin and Tim T. Tokaryk (2012). "A new evolutionary lineage of diving birds from the Late Cretaceous of North America and Asia". Palaeoworld 21 (1): 59–63. doi:10.1016/j.palwor.2012.02.005.
↑ Nikita V. Zelenkov (2012). "A new duck from the Middle Miocene of Mongolia, with comments on Miocene evolution of ducks". Paleontological Journal 46 (5): 520–530. doi:10.1134/s0031030112050139.
↑ Zlatozar Boev (2012). "Circaetus rhodopensis sp. n. (Aves: Accipitriformes) from the Late Miocene of Hadzhidimovo (SW Bulgaria)" (PDF). Acta Zoologica Bulgarica 64 (1): 5–12.
↑ Robert M. Chandler (2012). "A new species of tinamou (Aves: Tinamiformes, Tinamidae) from the Early-Middle Miocene of Argentina" (PDF). PalArch’s Journal of Vertebrate Palaeontology 9 (2): 1–8.
↑ Nikita V. Zelenkov (2012). "Neogene geese and ducks (Aves: Anatidae) from localities of the Great Lakes Depression, western Mongolia". Paleontological Journal 46 (6): 607–619. doi:10.1134/s0031030112060123.
1 2 3 Nikita V. Zelenkov and Evgeny N. Kurochkin (2012). "The first representative Pliocene assemblages of passerine birds in Asia (Northern Mongolia and Russian Transbaikalia)". Geobios 45 (3): 323–334. doi:10.1016/j.geobios.2011.10.001.
↑ Jingmai K. O'Connor, Chengkai Sun, Xing Xu, Xiaolin Wang and Zhonghe Zhou (2012). "A new species of Jeholornis with complete caudal integument" (PDF). Historical Biology 24 (1): 29–41. doi:10.1080/08912963.2011.552720.
1 2 Daniel T. Ksepka, R. Ewan Fordyce, Tatsuro Ando and Craig M. Jones (2012). "New fossil penguins (Aves, Sphenisciformes) from the Oligocene of New Zealand reveal the skeletal plan of stem penguins" (PDF). Journal of Vertebrate Paleontology 32 (2): 235–254. doi:10.1080/02724634.2012.652051.
↑ Zihui Zhang, Yunping Huang, Helen F. James and Lianhai Hou (2012). "A marabou (Ciconiidae: Leptoptilos) from the Middle Pleistocene of northeastern China" (PDF). The Auk 129 (4): 699–706. doi:10.1525/auk.2012.11227.
↑ Gerald Mayr and Evgenij Zvonok (2012). "A new genus and species of Pelagornithidae with well-preserved pseudodentition and further avian remains from the middle Eocene of the Ukraine". Journal of Vertebrate Paleontology 32 (4): 914–925. doi:10.1080/02724634.2012.676114.
↑ Juan Carlos Rando, Harald Pieper, Josep A. Alcover and Storrs L. Olson (2012). "A new species of extinct fossil scops owl (Aves: Strigiformes: Strigidae: Otus) from the Archipelago of Madeira (North Atlantic Ocean)" (PDF). Zootaxa 3182: 29–42.
↑ Gerald Mayr and Ružena Gregorová (2012). "A tiny stem group representative of Pici (Aves, Piciformes) from the early Oligocene of the Czech Republic". Paläontologische Zeitschrift 86 (3): 333–343. doi:10.1007/s12542-012-0133-5.
↑ Storrs L. Olson and David B. Wingate (2012). "A new species of towhee (Aves: Emberizidae: Pipilo) from Quaternary deposits on Bermuda". Proceedings of the Biological Society of Washington 125 (1): 85–96. doi:10.2988/11-21.1.
↑ Trevor H. Worthy (2012). "A phabine pigeon (Aves : Columbidae) from Oligo-Miocene Australia". Emu 112 (1): 23–31. doi:10.1071/mu11061.
↑ Federico L. Agnolín and Jorge I. Noriega (2012). "Una nueva especie de ñandú (Aves: Rheidae) del Mioceno tardío de la Mesopotamia Argentina". Ameghiniana 49 (2): 236–246. doi:10.5710/amgh.v49i2(492).
1 2 Jeno Kessler and János Hír (2012). "Észak-Magyarország madárvilága a miocénben. I. rész." (PDF). Földtani közlöny 142 (1): 67–78.
↑ Min Wang, Gerald Mayr, Jiangyong Zhang and Zhonghe Zhou (2012). "New bird remains from the Middle Eocene of Guangdong, China" (PDF). Acta Palaeontologica Polonica 57 (3): 519–526. doi:10.4202/app.2011.0061.
↑ Zhou Shuang, Zhou Zhong-He and Jingmai K. O’Connor (2012). "A new basal beaked ornithurine bird from the Lower Cretaceous of Western Liaoning, China" (PDF). Vertebrata PalAsiatica 50 (1): 9–24.
↑ Li Li, Wang Jinqi, Zhang Xi and Hou Shilin (2012). "A New Enantiornithine Bird from the Lower Cretaceous Jiufotang Formation in Jinzhou Area, Western Liaoning Province, China". Acta Geologica Sinica 86 (5): 1039–1044. doi:10.1111/j.1755-6724.2012.00729.x.
↑ Min Wang, Gerald Mayr, Jiangyong Zhang and Zhonghe Zhou (2012). "Two New Skeletons of the Enigmatic, Rail-like Avian Taxon Songzia Hou, 1990 (Songziidae) from the Early Eocene of China". Alcheringa 36 (4): 487–499. doi:10.1080/03115518.2012.673302.
↑ Gerald Mayr and Thierry Smith (2012). "A Fossil Albatross from the Early Oligocene of the North Sea Basin". The Auk 129 (1): 87–95. doi:10.1525/auk.2011.11192.
↑ Dongyu Hu, Xing Xu, Lianhai Hou and Corwin Sullivan (2012). "A new enantiornithine bird from the Lower Cretaceous of Western Liaoning, China, and its implications for early avian evolution". Journal of Vertebrate Paleontology 32 (3): 639–645. doi:10.1080/02724634.2012.652321.
↑ Fernando E. Novas, Martin Kundrat, Federico L. Agnolín, Martín D. Ezcurra, Per Erik Ahlberg, Marcelo P. Isasi, Alberto Arriagada and Pablo Chafrat (2012). "A new large pterosaur from the Late Cretaceous of Patagonia". Journal of Vertebrate Paleontology 32 (6): 1447–1452. doi:10.1080/02724634.2012.703979.
↑ David W. E. Hone, Helmut Tischlinger, Eberhard Frey and Martin Röper (2012). "A New Non-Pterodactyloid Pterosaur from the Late Jurassic of Southern Germany". PLoS ONE 7 (7): e39312. doi:10.1371/journal.pone.0039312. PMC 3390345. PMID 22792168.
↑ Lü Junchang and David W.E. Hone (2012). "A New Chinese Anurognathid Pterosaur and the Evolution of Pterosaurian Tail Lengths". Acta Geologica Sinica 86 (6): 1317–1325. doi:10.1111/1755-6724.12002.
↑ Romain Vullo, Jesús Marugán-Lobón, Alexander W. A. Kellner, Angela D. Buscalioni, Bernard Gomez, Montserrat de la Fuente and José J. Moratalla (2012). "A New Crested Pterosaur from the Early Cretaceous of Spain: The First European Tapejarid (Pterodactyloidea: Azhdarchoidea)". PLoS ONE 7 (7): e38900. doi:10.1371/journal.pone.0038900. PMC 3389002. PMID 22802931.
↑ Lü Junchang, Ji Qiang, Wei Xuefang and Liu Yongqing (2012). "A new ctenochasmatoid pterosaur from the Early Cretaceous Yixian Formation of western Liaoning, China". Cretaceous Research 34: 26–30. doi:10.1016/j.cretres.2011.09.010.
↑ Xiaolin Wang, Alexander W. A. Kellner, Shunxing Jiang and Xin Cheng (2012). "New toothed flying reptile from Asia: close similarities between early Cretaceous pterosaur faunas from China and Brazil". Naturwissenschaften 99 (4): 249–257. doi:10.1007/s00114-012-0889-1. PMID 22354475.
↑ Xin Cheng, Xiaolin Wang, Shunxing Jiang and Alexander W.A. Kellner (2012). "A new scaphognathid pterosaur from western Liaoning, China". Historical Biology 24 (1): 101–111. doi:10.1080/08912963.2011.635423.
↑ Lü Junchang, Pu Hanyong, Xu Li, Wu Yanhua and Wei Xuefang (2012). "Largest toothed pterosaur skull from the Early Cretaceous Yixian Formation of Western Liaoning, China, with comments on the family Boreopteridae". Acta Geologica Sinica 86 (2): 287–293. doi:10.1111/j.1755-6724.2012.00658.x.
↑ Lü, J., Unwin, D.M., Zhao, B., Gao, C. and Shen, C. (2012). "A new rhamphorhynchid (Pterosauria: Rhamphorhynchidae) from the Middle/Upper Jurassic of Qinglong, Hebei Province, China" (PDF). Zootaxa 3158: 1–19.
1 2 Darren Naish, Gareth Dyke, Andrea Cau, François Escuillié and Pascal Godefroit (2012). "A gigantic bird from the Upper Cretaceous of Central Asia". Biology Letters 8 (1): 97–100. doi:10.1098/rsbl.2011.0683. PMC 3259976. PMID 21835881.
↑ Eric Buffetaut (2011). "Samrukia nessovi, from the Late Cretaceous of Kazakhstan: A large pterosaur, not a giant bird". Annales de Paléontologie 97 (3–4): 133–138. doi:10.1016/j.annpal.2011.10.001.
↑ Naish, Darren (January 25, 2012). "Happy 6th Birthday, Tetrapod Zoology (part II)". Tetrapod Zoology. Retrieved March 15, 2012.
↑ Grzegorz Niedźwiedzki, Tomasz Sulej and Jerzy Dzik (2012). "A large predatory archosaur from the Late Triassic of Poland". Acta Palaeontologica Polonica 57 (2): 267–276. doi:10.4202/app.2010.0045.

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