Taxonomy of the Euphorbiaceae

Here is a full taxonomy of the family Euphorbiaceae, according to the most recent molecular research . This complex family previously comprising 5 subfamilies: the Acalyphoideae, the Crotonoideae, the Euphorbioideae, the Phyllanthoideae and the Oldfieldioideae. The 3 first ones are uni-ovulate families while the 2 last one are bi-ovulate.

Now the Euphorbiaceae has been split into 5 families: The 3 uni-ovulate subfamilies have become the Euphorbiaceae in the strict sense, with the tribe Galearieae in the Acalyphoideae forming the most of the family Pandaceae.[1] Part of the bi-ovulate subfamily Phyllanthoideae has become the family Phyllanthaceae,[1] with the tribe Drypeteae as family Putranjivaceae[1] and the tribe Centroplaceae part of the Pandaceae. The other bi-ovulate subfamily Oldfieldioideae has become the Picrodendraceae.[1]

Subfamily Acalyphoideae

Tribe Acalypheae

There are 12 subtribes and 32 genera:

Subtribe Acalyphinae
Acalypha (also Acalyphes, Acalyphopsis, Calyptrospatha, Caturus, Corythea, Cupameni, Galurus, Gymnalypha, Linostachys, Mercuriastrum, Odonteilema, Paracelsea, Ricinocarpus, Schizogyne, Usteria)
Subtribe Adrianinae
Adriana (also Meialisa, Trachycaryon)
Subtribe Claoxylinae
Claoxylon (also Erythrochilus, Erythrochylus)
Claoxylopsis
Discoclaoxylon
Erythrococca (also Athroandra, Autrandra, Chloropatane, Deflersia, Poggeophyton, Rivinoides)
Micrococca
Subtribe Cleidiinae
Cleidion (also Lasiostyles, Psilostachys, Redia, Tetraglossa)
Sampantaea
Wetria (also Pseudotrewia)
Subtribe Dysopsidinae
Dysopsis (also Mirabellia, Molina)
Subtribe Lasiococcinae
Clonostylis
Homonoia (also called Haematospermum, Lumanaja])
Lasiococca
Spathiostemon (also Polydragma)
Subtribe Lobaniliinae
Lobanilia
Subtribe Macaranginae
Macaranga (also Adenoceras, Mappa, Mecostylis, Pachystemon, Panopia, Phocea, Tanarius)
Subtribe Mareyinae
Mareya
Mareyopsis
Subtribe Mercurialinae
Mercurialis (also Cynocrambe, Discoplis, Synema)
Leidesia
Seidelia
Subtribe Ricininae
Ricinus - castor bean
Subtribe Rottlerinae
Avellanita
Coccoceras
Cordemoya (also Boutonia)
Deuteromallotus
Mallotus (also Aconceveibum, Axenfeldia, Coelodiscus, Diplochlamys, Echinocroton, Echinus, Hancea, Lasipana, Plagianthera, Rottlera,Stylanthus)
Neotrewia
Octospermum
Rockinghamia
Trewia (also Canschi, Trevia)

Tribe Adelieae

There are 5 genera:

Adelia (also Ricinella)
Crotonogynopsis
Enriquebeltrania (also Beltrania)
Lasiocroton
Leucocroton

Tribe Agrostistachydeae

There are 4 genera:

Agrostistachys (also Heterocalyx, Sarcoclinium)
Chondrostylis (also Kunstlerodendron)
Cyttaranthus
Pseudagrostistachys

Tribe Alchorneae

There are 2 subtribes and 7 genera:

Subtribe Alchorneinae
Alchornea (also Bleekeria, Cladodes, Hermesia, Lepidoturus, Schousboea, Stipellaria)
Aparisthmium
Bocquillonia (also Ramelia)
Orfilea (also Diderotia, Laurembergia)
Subtribe Conceveibinae
Conceveiba (also Conceveibastrum, Conceveibum, Veconcibea)
Gavarretia
Polyandra

Tribe Ampereae

There are 2 genera:

Amperea (also Leptomeria)
Monotaxis (also Hippocrepandra, Reissipa)

Tribe Bernardieae

There are 6 genera:

Adenophaedra
Amyrea
Bernardia (also Alevia, Bernarda, Bernhardia, Bivonia, Passaea, Phaedra, Polyboea, Traganthus, Tyria)
Discocleidion
Necepsia (also Neopalissya, Palissya)
Paranecepsia

Tribe Caryodendreae

There are 3 genera:

Alchorneopsis
Caryodendron (also Centrodiscus)
Discoglypremna

Tribe Chaetocarpeae

This tribe has 2 genera:

Chaetocarpus (also Gaedawakka, Mettenia, Neochevaliera, Regnaldia)
Trigonopleura (also Peniculifera)

Tribe Cheiloseae

There are 2 genera:

Cheilosa
Neoscortechinia (also Alcineanthus, Scortechinia)

Tribe Chrozophoreae

This tribe has 4 subtribes and 12 genera:

Subtribe Chrozophorinae
Chrozophora (also Crossophora, Crozophora, Ricinoides, Tournesol, Tournesolia)
Subtribe Ditaxinae
Argythamnia (also Argithamnia, Argothamnia, Argyrothamnia, Argytamnia, Odotalon, Serophyton)
Caperonia (also Acanthopyxis, Androphoranthus, Cavanilla, Lepidococea, Meterana)
Chiropetalum (also Aonikena, Desfontaena, Desfontaina, Desfontainea)
Ditaxis (also Aphora, Paxiuscula, Stenonia)
Philyra (also Phyllera)
Subtribe Doryxylinae
Doryxylon (also Mercadoa, Sumbavia)
Melanolepis
Sumbaviopsis (also Adisa, Adisca)
Thyrsanthera
Subtribe Speranskiinae
Speranskia
Annual Mercury (Mercurialis annua)

Tribe Clutieae

Clutia (also Altora, Cluytia, Clytia)

Tribe Dicoelieae

This tribe has one genus:

Dicoelia

Tribe Epiprineae

There are 2 subtribes and 9 genera:

Subtribe Epiprininae
Adenochlaena (also Niedenzua, Centrostylis)
Cephalocroton
Cephalocrotonopsis
Cladogynos (also Adenogynum, Baprea, Chloradenia)
Cleidiocarpon (also Sinopimelodendron)
Epiprinus
Koilodepas (also Caelodepas, Calpigyne, Coelodepas, Nephrostylus)
Symphyllia
Subtribe Cephalomappinae
Cephalomappa
Muricococcum

Tribe Erismantheae

There are 3 genera:

Erismanthus
Moultonianthus
Syndyophyllum

Tribe Omphaleae

This tribe only has 1 genus:

Omphalea (also Duchola, Hebecocca, Hecatea, Neomphalea, Omphalandria, Ronnowia)

Tribe Pereae

There is only 1 genus:

Pera

Tribe Plukenetieae

There are 3 subtribes and 13 genera:

Subtribe Dalechampiinae
Dalechampia (also Cremophyllum, Dalechampsia, Megalostylis, Rhopalostylis)
Subtribe Plukenetiinae
Angostylis (also Angostyles)
Astrococcus
Plukenetia (also Accia, Angostylidium, Apopandra, Botryanthe, Ceratococcus, Elaeophora, Eleutherostigma, Fragariopsis, Hedraiostylus, Pseudotragia, Pterococcus, Sajorium, Tetracarpidium or Vigia)
Haematostemon
Romanoa (also Anabaena, Anabaenella)
Subtribe Tragiinae
Acidoton (also Durandeeldia, Gitara)
Cnesmone (also Cenesmon, Cnesmosa)
Megistostigma (also Clavistylus)
Pachystylidium
Platygyna (also Acanthocaulon)
Sphaerostylis
Tragia (also Agirta, Allosandra, Bia, Ctenomeria, Lassia, Leptobotrys, Leptorhachis, Leucandra, Schorigeram, Zuckertia)

Tribe Pogonophoreae

This tribe only has 1 genus:

Pogonophora (also Poraresia)

Tribe Pycnocomeae

There are 2 subtribes and 7 genera:

Subtribe Blumeodendrinae
Blumeodendron
Botryophora (also Botryospora, Botryphora)
Podadenia
Ptychopyxis (also Clarorivinia)
Subtribe Pycnocominae
Argomuellera (also Neopycnocoma, Wetriaria)
Droceloncia
Pycnocoma (also Comopyena)

Tribe Sphyranthereae

There is only 1 genus:

Sphyranthera

Subfamily Crotonoideae

Tribe Adenoclineae

There are 2 subtribes and 6 genera:

Subtribe Adenoclininae
Adenocline (also Adenoclina, Diplostylis, Paradenocline)
Ditta
Glycydendron
Klaineanthus

Tetrorchidium (also Hasskarlia, Tetrorchidiopsis)

Subtribe Endosperminae
Endospermum (also Capellenia)

Tribe Aleuritideae

This tribe has 6 subtribes and 16 genera:

Subtribe Aleuritinae
Aleurites (also Camirium)
Reutealis
Vernicia (also Ambinux, Dryandra, Elaeococca)
Subtribe Benoistiinae
Benoistia
Subtribe Crotonogyninae
Crotonogyne (also Neomanniophyton)
Cyrtogonone
Manniophyton
Subtribe Garciinae
Garcia (also called Carcia)
Subtribe Grosserinae
Anomalocalyx
Cavacoa
Grossera
Neoholstia (also Holstia)
Sandwithia
Tannodia (also Domohinea)
Tapoides
Subtribe Neoboutoniinae
Neoboutonia

Tribe Codiaeae

There are 15 genera:

Acidocroton
Baliospermum
Baloghia (also Steigeria)
Blachia (also Deonia)
Codiaeum (also Crozophyla, Junghuhnia, Phyllaurea, Synaspisma)
Colobocarpos
Dimorphocalyx
Dodecastigma
Fontainea
Hylandia
Ophellantha
Ostodes
Pantadenia
Sagotia
Strophioblachia

Tribe Crotoneae

There are 5 genera:

Brasiliocroton
Croton (also Agelandra, Aldinia, Angelandra, Anisepta, Anisophyllum, Argyra, Argyrodendron, Astraea, Astrogyne, Aubertia, Banalia, Barhamia, Brachystachys, Calypteriopetalon, Cascarilla, Centrandra, Cieca, Cleodora, Codonocalyx, Comatocroton, Crotonanthus, Crotonopsis, Cyclostigma, Decarinium, Drepadenium, Eluteria, Engelmannia, Eremocarpus, Eutrophia, Friesia, Furcaria, Geiseleria, Gynamblosis, Halecus, Hendecandas, Heptallon, Heterochlamys, Heterocroton, Julocroton, Klotzschiphytum, Kurkas, Lasiogyne, Leptemon, Leucadenia, Luntia, Macrocroton, Medea, Merleta, Monguia, Myriogomphus, Ocalia, Oxydectes, Palanostigma, Penteca, Pilinophyton, Piscaria, Pleopadium, Podostachys, Saipania, Schradera, Semilta, Tiglium, Timandra, Tridesmis, Triplandra, Vandera)
Mildbraedia (also Neojatropha, Plesiatropha)
Moacroton (also Cubacroton)
Paracroton (also Desmostemon, Fahrenheitia)

Tribe Elateriospermeae

This tribe only has 1 genus:

Elateriospermum (also Elaterioides, Elaterispermum)

Tribe Gelonieae

This tribe has 2 genera:

Cladogelonium
Suregada (also Ceratophorus, Erythrocarpus, Gelonium, Owataria)

Tribe Jatropheae

There are 8 genera:

Annesijoa
Deutzianthus
Jatropha (also Adenorhopium, Adenoropium, Castiglionia, Collenucia, Curcas, Jatropa, Loureira, Mesandrinia, Mozinna, Zimapania)
Joannesia (also Anda, Andicus)
Leeuwenbergia
Loerzingia
Oligoceras
Vaupesia

Tribe Manihoteae

This tribe has 2 genera:

Cnidoscolus (also Bivonea, Jussieuia, Mandioca, Victorinia)
Manihot (also Hotnima, Janipha, Manihotoides)

Tribe Micrandreae

There are 2 subtribes and 4 genera:

Subtribe Heveinae
Hevea (also Caoutchoua, Micrandra, Siphonanthus, Siphonia)
Subtribe Micrandrinae
Cunuria
Micrandra (also Clusiophyllum, Pogonophyllum)
Micrandropsis

Tribe Ricinocarpeae

There are 2 subtribes and 7 genera:

Subtribe Bertyinae
Bertya (also Lambertya)
Borneodendron
Cocconerion
Myricanthe
Subtribe Ricinocarpinae
Alphandia
Beyeria (also Beyeriopsis, Calyptrostigma, Clavipodium)
Ricinocarpus (also Echinosphaera, Ricinocarpus, Roeperia)

Tribe Ricinodendreae

There are 2 genera:

Ricinodendron
Schinziophyton

Tribe Trigonostemoneae

There is only 1 genus:

Trigonostemon (also Actephilopsis, Athroisma, Enchidium, Kurziodendron, Neotrigonostemon, Nepenthandra, Poilaniella, Prosartema, Silvaea, Telogyne, Tritaxis, Tylosepalum)

Subfamily Euphorbioideae

Tribe Euphorbieae

This tribe has 3 subtribes and 6 genera:

Subtribe Anthosteminae
Anthostema
Dichostemma
Subtribe Euphorbiinae
Cubanthus (to be transferred to Euphorbia)
Euphorbia (also Ademo, Adenopetalum, Adenorima, Agaloma, Aklema, Alectoroctonum, Allobia, Anisophyllum, Anthacantha, Aplarina, Arthrothamnus, Bojeria, Ceraselma, Chamaesyce, Characias, Chylogala, Crepidaria, Ctenadena, Cyathophora, Cystidospermum, Dactylanthes, Dematra, Desmonema, Diadenaria, Dichrophyllum, Dichylium, Diplocyathium, Ditritra, Elaeophorbia, Endadenium, Endoisila, Epurga, Esula, Euforbia, Eumecanthus, Euphorbiastrum, Euphorbiodendron, Euphorbiopsis, Euphorbium, Galarhoeus, Hexadenia, Kanopikon, Kobiosis, Lacanthis, Lathyris, Lepadena, Leptopus, Lophobios, Lortia, Lyciopsis, Medusea, Monadenium, Nisomenes, Ossifraga, Peccana, Pedilanthus, Petalandra, Pleuradena, Poinsettia, Pythius, Sclerocyathium, Stenadenium, Sterigmanthe, Synadenium, Tithymaloides, Tithymalopsis, Tithymalus, Torfasadis, Treisia, Tricherostigma, Trichosterigma, Tumalis, Vallaris, Ventenatia, Xamesike, Zalitea, Zygophyllidium)
Subtribe Neoguillauminiinae
Calycopeplus
Neoguillauminia

Tribe Hippomaneae

This tribe has 2 subtribes and 33 genera:

Subtribe Carumbiinae
Omalanthus (also Carumbium, Dibrachion, Dibrachium, Duania, Wartmannia)
Subtribe Hippomaninae
Actinostemon (also Dactylostemon)
Adenopeltis
Anomostachys
Balakata
Bonania (also Hypocoton)
Colliguaja
Conosapium
Dalembertia (also Alcoceria)
Dendrocousinsia
Dendrothrix
Ditrysinia
Duvigneaudia
Excoecaria (also Commia, Glyphostylus)
Falconeria
Grimmeodendron
Gymnanthes (also Adenogyne, Ateramnus)
Hippomane (also Mancanilla, Mancinella)
Mabea
Maprounea (also Aegopicron, Aegopricon, Aegopricum)
Neoshirakia (also Shirakia)
Pleradenophora
Pseudosenefeldera
Rhodothyrsus
Sapium (also Carumbium, Gymnobothrys, Sapiopsis, Seborium, Shirakiopsis, Stillingfleetia, Taeniosapium) - Chinese tallow
Sclerocroton
Sebastiania (also Clonostachys, Cnemidostachys, Elachocroton, Gussonia, Microstachys, Sarothrostachys, Tragiopsis)
Senefeldera
Senefelderopsis
Spegazziniophytum
Spirostachys
Stillingia (also Gymnostillingia)
Triadica

Tribe Hureae

There are 4 genera:

Algernonia
Hura
Ophthalmoblapton
Tetraplandra (also Dendrobryon)

Tribe Pachystromateae

There is one genus:

Pachystroma (also Acantholoma)

Tribe Stomatocalyceae

There are 2 subtribes and 4 genera:

Subtribe Hamilcoinae
Hamilcoa
Nealchornea
Subtribe Stomatocalycinae
Pimelodendron (also Stomatocalyx)
Plagiostyles

See also

References

  1. 1.0 1.1 1.2 1.3 Wurdack, K. J.; Hoffmann, P.; Chase, M. W. (2005), "Molecular phylogenetic analysis of uniovulate Euphorbiaceae (Euphorbiaceae sensu stricto) using plastid RBCL and TRNL-F DNA sequences", American Journal of Botany 92: 1397, doi:10.3732/ajb.92.8.1397, PMID 21646159

External links