Mangeliidae
| Mangeliidae | |
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| Apertural view of the mangeliid Bela zonata | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| (unranked): | clade Caenogastropoda clade Hypsogastropoda clade Neogastropoda |
| Superfamily: | Conoidea |
| Family: | Mangeliidae [1] P. Fischer, 1883 |
| Genera | |
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See text | |
| Synonyms | |
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Mangeliidae is a monophyletic family of small to medium-sized sea snails, marine gastropod mollusks in the superfamily Conoidea.[1][2]
in 2011, Bouchet, Kantor et al elevated the subfamily Mangeliinae combined with the subfamily Oenopotinae (at that point both belonging to the family Conidae) to the rank of family. This was based on anatomical characters and a dataset of molecular sequences of three gene fragments.[1]
Description
The Mangeliidae are characterized in general by the lack of an operculum and by a deep sinus with a heavy callus on the shoulder slope of the outer lip. This outer lip (labrum) is reinforced. The aperture is oval-elongated, usually narrow, terminating in a rather short, truncated siphon. The aperture is only rarely denticulate. The axial ribs are dominant in the sculpture of the shell. The toxoglossate radula has a weak basal ribbon and relatively short marginal teeth with very variable morphology. The tooth cavity opens laterally.[3]
It should be noted however, that the genera Neoguraleus and Liracraea are operculate; the operculum in these genera has a terminal nucleus. These exceptions mean that the reinforced outer lip is the most characteristic feature of this subfamily.
Genera
This is a list of the accepted names of species in the family Mangeliidae : (the main reference for recent species is the World Register of Marine Species)[4]
- Acmaturris Woodring, 1928
- Agathotoma Cossman, 1899
- Anticlinura Thiele, 1934
- Antiguraleus Powell, 1942
- Apispiralia Laseron, 1954
- Apitua Laseron, 1954
- Bactrocythara Woodring, 1928
- Bela Gray, 1847
- Bellacythara McLean, 1971
- Benthomangelia Thiele, 1925
- Brachycythara Woodring, 1928
- Cacodaphnella Pilsbry & Lowe, 1933
- Citharomangelia Kilbu, 1992
- Cryoturris Woodring, 1928
- Curtitoma Bartsch, 1941
- Cythara Fischer, 1883 (nomen dubium)
- Cytharopsis A. Adams, 1865
- Eucithara Fischer, 1883
- Euryentmema Woodring, 1928
- Genotina Vera-Peláez, 2004
- Gingicithara Kilbu, 1992
- Glyphoturris Woodring, 1928
- Granotoma Bartsch, 1941
- Granoturris Fargo, 1953
- Guraleus Hedley, 1918
- Hemicythara Kuroda & Oyama, 1971
- Heterocithara Hedley, 1922
- Ithycythara Woodring, 1928
- Kurtzia Bartsch 1944
- Kurtziella Dall, 1918
- Leiocithara Hedley, 1922
- Liracraea Odhner, 1924
- Lorabela Powell, 1951
- Macteola Hedley, 1918
- Mangelia Risso, 1826 (78 species)
- Marita Hedley, 1922
- Mitraguraleus Laseron, 1954
- Neoguraleus Powell, 1939
- Notocytharella Hertlein & Strong, 1955
- Obesotoma Bartsch, 1941
- Oenopota Mörch, 1852
- Oenopotella A. Sysoev, 1988
- Papillocithara Kilbu, 1992
- Paraguraleus Powell, 1944
- Paraspirotropis Sysoev & Kantor, 1984
- Perimangelia McLean, 2000
- Platycythara Woodring, 1928
- Pleurotomoides Bronn, 1831
- Propebela Iredale, 1918
- Pseudorhaphitoma Boettger, 1895
- Pyrgocythara Woodring, 1928
- Rubellatoma Bartsch & Rehder, 1939
- Saccharoturris Woodring, 1928
- Stellatoma Bartsch & Rehder, 1939
- Suturocythara Garcia, 2008
- Tenaturris Woodring, 1928
- Toxicochlespira Sysoev & Kantor, 1990
- Venustoma Bartsch, 1941
- Vexiguraleus Powell, 1942
- Vitjazinella Sysoev 1988
- Vitricythara Fargo, 1953
- Genera moved to other families
- Anacithara Hedley, 1922 : brought into the new family Horaiclavidae
- Austropusilla Laseron, 1954 : brought into the new family Raphitomidae
- Belaturricula Powell, 1951 : brought into the new family Borsoniidae
- Clathromangelia Monterosato, 1884 : brought into the new family Clathurellidae
- Conopleura Hinds, 1844 : belongs to Drilliidae
- Euclathurella Woodring, 1928 : belongs to the new family Clathurellidae
- Glyptaesopus Pilsbry & Olsson, 1941 : brought into the new family Borsoniidae
- Lienardia Jousseaume, 1884 : brought into the new family Clathurellidae
- Lioglyphostomella Shuto, 1970 : brought into the new family Pseudomelatomidae
- Otitoma Jousseaume, 1898 : belongs to the new family Pseudomelatomidae
- Paraclathurella Boettger, 1895 : belongs to the new family Clathurellidae
- Paramontana Laseron, 1954 : belongs to the new family Raphitomidae
- Pseudoetrema Oyama, 1953 : belongs to the new family Clathurellidae
- Thelecythara Woodring, 1928 : belongs to the new family Horaiclavidae
- Turrella Laseron, 1954 : belongs to the new family Clathurellidae
- Genera brought into synonymy
- Canetoma Bartsch, 1941: synonym of Propebela Iredale, 1918
- Cestoma Bartsch, 1941: synonym of Propebela Iredale, 1918
- Cithara: synonym of Cythara Schumacher, 1817
- Clathromangilia: synonym of Clathromangelia Monterosato, 1884 : belongs now to the family Clathurellidae
- Clinuromella Beets, 1943: synonym of Anticlinura Thiele, 1934
- Clinuropsis Thiele, 1929: synonym of Anticlinura Thiele, 1934
- Cyrtocythara F. Nordsieck, 1977: synonym of Mangelia Risso, 1826
- Cytharella Monterosato, 1875: synonym of Mangelia (Cytharella) Monterosato, 1875
- Ditoma Bellardi, 1875: synonym of Agathotoma Cossman, 1899
- Euguraleus Cotton, 1947: synonym of Guraleus Hedley, 1918
- Fehria van Aartsen, 1988 : synonym of Bela Gray, 1847
- Funitoma Bartsch, 1941: synonym of Propebela Iredale, 1918
- Ginnania Monterosato, 1884: synonym of Bela Gray, 1847
- Lora Auctores non Gray, 1847: synonym of Oenopota Mörch, 1852
- Mangilia Lovén, 1846: synonym of Mangelia Risso, 1826
- Mangiliella Bucquoy, Dautzenberg & Dollfus, 1883 : synonym of Mangelia Risso, 1826
- Nematoma Bartsch, 1941: synonym of Curtitoma Bartsch, 1941
- Nodotoma Bartsch, 1941: synonym of Oenopota Mörch, 1852
- Pseudoraphitoma: synonym of Pseudorhaphitoma Boettger, 1895
- Rissomangelia Monterosato, 1917 : synonym of Mangelia Risso, 1826
- Rugocythara Nordsieck, 1977 : synonym of Mangelia Risso, 1826
- Scabrella Hedley, 1918: synonym of Asperdaphne Hedley, 1922
- Smithia Monterosato, 1884: synonym of Smithiella Monterosato, 1890
- Smithiella Monterosato, 1890 : synonym of Mangelia Risso, 1826
- Thelecytharella Shuto, 1969 : synonym of Otitoma Jousseaume, 1898 : belongs to the family Pseudomelatomidae
- Thetidos Hedley, 1899: synonym of Lienardia Jousseaume, 1884 : belongs to the family Clathurellidae
- Turritomella Bartsch, 1941: synonym of Propebela Iredale, 1918
- Villiersiella Monterosato, 1890: synonym of Mangelia Risso, 1826
- Widalli Bogdanov, 1986: synonym of Curtitoma Bartsch, 1941
- Cythara Schumacher, 1817 (nomen dubium)
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References
- ↑ 1.0 1.1 1.2 Bouchet P., Kantor Yu.I., Sysoev A. & Puillandre N. (2011) A new operational classification of the Conoidea. Journal of Molluscan Studies 77: 273-308.
- ↑ Bouchet P. & Rocroi J.-P. (Ed.) (2005). "Classification and nomenclator of gastropod families". Malacologia 47(1-2). ISBN 3-925919-72-4. 397 pp.
- ↑ Chen-Kwoh Chang, Small Turrids of Taiwan, chapter 12, p. 116; June, 1 2001
- ↑ WoRMS : Mangeliidae
- Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). ISBN 0-915826-22-4. XII, 195 pp