Lagerstätte

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Fossil fish from the Green River Formation, an Eocene Lagerstätte

Stranded scyphozoans with Climactichnites trackways from Blackberry Hill, Wisconsin (Cambrian). Scyphozoan in foreground is 10 cm in diameter. Slab is in hyporelief.

A Lagerstätte (German: [ˈlaːɡɐˌʃtɛtə], from Lager 'storage' Stätte 'place'; plural Lagerstätten) is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying decomposition. Lagerstätten span geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Jurassic Solnhofen limestone, and the Cretaceous Santana and Yixian formations.

Types

Palaeontologists distinguish two kinds:^[1]

1. Konzentrat-Lagerstätten (concentration Lagerstätten) are deposits with a particular "concentration" of disarticulated organic hard parts, such as a bone bed. These Lagerstätten are less spectacular than the more famous Konservat-Lagerstätten. Their contents invariably display a large degree of time averaging, as the accumulation of bones in the absence of other sediment takes some time. Deposits with a high concentration of fossils that represent an in situ community, such as reefs or oyster beds, are not considered Lagerstätten.
2. Konservat-Lagerstätten (conservation Lagerstätten) are deposits known for the exceptional preservation of fossilized organisms or traces. The individual taphonomy of the fossils varies with the sites. Conservation Lagerstätten are crucial in providing answers to important moments in the history and evolution of life. For example, the Burgess Shale of British Columbia is associated with the Cambrian explosion, and the Solnhofen limestone with the earliest known bird, Archaeopteryx.

Preservation

Konservat-Lagerstätten preserve lightly sclerotized and soft-bodied organisms or traces of organisms that are not otherwise preserved in the usual shelly and bony fossil record; thus, they offer more complete records of ancient biodiversity and behavior and enable some reconstruction of the palaeoecology of ancient aquatic communities. In 1986, Simon Conway Morris calculated only about 14% of genera in the Burgess Shale had possessed biomineralized tissues in life. The affinities of the shelly elements of conodonts were mysterious until the associated soft tissues were discovered near Edinburgh, Scotland, in the Granton Lower Oil Shale of the Carboniferous.^[2] Information from the broader range of organisms found in Lagerstätten have contributed to recent phylogenetic reconstructions of some major metazoan groups. Lagerstatten seem to be temporally autocorrelated, perhaps because global environmental factors such as climate might affect their deposition.^[3]

A number of taphonomic pathways may produce Lagerstätten. The following is an incomplete list:

• Orsten-type and Doushantuo-type preservations preserve organisms in phosphate.
• Bitter Springs-type preservation preserves them in silica.
• Carbonaceous films are the result of Burgess Shale-type preservation
• Pyrite preserves exquisite detail in Beecher’s trilobite-type preservation.
• Ediacaran-type preservation preserves casts and moulds with the aid of microbial mats.

Important Konservat-Lagerstätten

The world's major Lagerstätten include:

See also

• List of fossil sites (with link directory)

References

Further reading

• Penney, D.(ed.) 2010. Biodiversity of Fossils in Amber from the Major World Deposits. Siri Scienfic Press, Manchester, 304 pp.
• "Fossil Lagerstätten". Department of Earth Sciences, University of Bristol. 2003. Retrieved 2005-11-21.  – A catalogue of sites of exceptional fossil preservation produced by MSc palaeobiology students at University of Bristol's Department of Earth Sciences.
• Orr, Patrick J.; David J. Siveter (1 January 2000). "Three-dimensional preservation of a non-biomineralized arthropod in concretions in Silurian volcaniclastic rocks from Herefordshire, England". Journal of the Geological Society 157 (1): 173–186. doi:10.1144/jgs.157.1.173. Retrieved 2006-10-26.