Hesperocyoninae

Hesperocyoninae Temporal range: Late Eocene–Middle Miocene PreꞒ Ꞓ O S D C P T J K Pg N
Skull of Mesocyon coryphaeus
Conservation status
Fossil
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Subfamily:	†Hesperocyoninae
Genera
†Cynodesmus †Caedocyon †Ectopocynus †Enhydrocyon †Hesperocyon †Mesocyon †Osbornodon †Paraenhydrocyon †Philotrox †Sunkahetanka

Phylogeny of Hesperocyoninae related to Borophaginae and Canines by R.L. Tedford, 1977

Fluctuation of species within Canidae over 40 million years

The Hesperocyoninae are a subfamily of extinct canids.

Taxonomic history

The Hesperocyoninae subfamily was named by Martin (1989). The members were reassigned to the family Canidae (with no subfamily) by Xiaoming Wang in 1999.^[1]

Hesperocyoninae are basal canids that, according to Wang and Tedford, gave rise to the other canid groups, such as the Borophaginae and Caninae.

This disused subfamily was endemic to North America, living from the Duchesnean stage of the Late Eocene through to the early Barstovian stage of the Miocene, lasting around 20 million years. It comprises 10 recognized genera and 26 recognized species; among these, four genera and species are new. Four major lineages can be defined based on shared characteristics:

• Mesocyon-Enhydrocyon clade
• Osbornodon clade
• Paraenhydrocyon
• Ectopocynus clade

The genus Caedocyon probably forms a distinct clade of its own.

Hesperocyon, which lacks the shared derived characters that would include it within any of the aforementioned clades, is possibly ancestral to many of the lineages. Some evidence indicates the Paraenhydrocyon clade may be directly descended from Hesperocyon gregarius. According to Xiaoming Wang,^[2] Hesperocyon coloradensis provides an important link between H. gregarius and the Mesocyon-Enhydrocyon clade.

Hesperocyoninae species per unit of time

References

Additional Reading

• Xiaoming Wang, Richard H. Tedford, Mauricio Antón, Dogs: Their Fossil Relatives and Evolutionary History, New York : Columbia University Press, 2008; ISBN 978-0-231-13528-3