Entelodont

Entelodonts Temporal range: 37.2–16.3Ma PreꞒ Ꞓ O S D C P T J K Pg N Late Eocene - Early Miocene
Daeodon skeleton
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
(unranked):	Cetancodontamorpha
Family:	†Entelodontidae Lydekker, 1883
Genera
†Archaeotherium †Brachyhyops †Cypretherium †Daeodon †Entelodon †Eoentelodon †Paraentelodon †Proentelodon
Synonyms
Elotheridae

Entelodonts — sometimes facetiously termed hell pigs or terminator pigs^[1] — are an extinct family of pig-like omnivores endemic to forests and plains of North America, Europe, and Asia from the late Eocene to early Miocene epochs (37.2—16.3 million years ago), existing for about 21 million years.^[2]

Taxonomy

Archaeotherium mortoni skull

The Entelodontidae were named by Richard Lydekker and assigned to Nonruminantia by Gregory (1910).^[3] They were then assigned to Artiodactyla by Lucas et al.^[4] (1998) and to Entelodontoidea by Carroll (1988) and Boisserie et al. (2005). While entelodonts have long been classified as members of the Suina, Spaulding et al. have found them to be closer to whales and hippos than to pigs.^[5]

Description

Entelodonts are an extinct group of rather pig-like omnivorous mammals with bulky bodies, but short, slender legs, and long muzzles. The largest were the North American Daeodon shoshonensis, and the Eurasian Paraentelodon intermedium, standing up to 2.1 m (6.9 ft) tall at the shoulder, with brains the size of an orange.^[6]

A single specimen recorded by M. Mendoza, C. M. Janis, and P. Palmqvist for body mass was estimated to have a weight of 421 kg (930 lb).^[7]

Entelodonts had full sets of teeth, including large canines, heavy incisors, and relatively simple, yet powerful, molars. These features suggest an omnivorous diet, similar to that of modern pigs. Like many other artiodactyls, they had cloven hooves, with two toes touching the ground, and the remaining two being vestigial.^[8]

The most distinguishing characteristic of the animals, however, would have been the heavy, bony lumps on either side of their heads which are similar to a warthog's. Some of these may have been attachment points for powerful jaw muscles,^[8][9] but some were larger in males, suggesting they may have fought over mates.^[10]

Paleobiology

Entelodon by Charles R. Knight

Entelodonts lived in the forests and plains where they were the apex predators of North America's early Miocene and Oligocene, consuming carrion and live animals and rounding off their diets with plants and tubers. They would have hunted large animals, like the cow-sized artiodactyl Eporeodon major and the sheep-sized cameloid Poebrotherium wilsoni, dispatching them with a bite from their jaws. Some fossil remains of these other animals have been found with the bite marks of entelodonts on them. Like modern-day pigs, they were omnivores, eating both meat and plants, but their adaptations show a bias towards live prey and carrion. They were most likely opportunists, mainly eating live animals, but not rejecting carrion and roots and tubers in times of drought. Some entelodonts even exhibited caching behavior, as an Archaeotherium's cache has been discovered, made up of the remains of several early camels.

In popular culture

Entelodonts appear in the third episode of the popular BBC documentary Walking with Beasts, where in the program, the narrator always refers to the creatures as "entelodonts" rather than a more specific genus, such as Entelodon or Archaeotherium. The same creatures appear in another BBC production, the 2001 remake of The Lost World. Entelodonts were also the main focus of episode 4 of National Geographic Channel's show Prehistoric Predators in an episode titled "Killer Pig". The episode featured Archaeotherium (identified as "entelodont") as being the top predator of the American Badlands, and how it evolved into the even larger Daeodon ("Dinohyus").

References

External links

• http://www.abc.net.au/beasts/factfiles/factfiles/entelodont.htm Video, images & sound
• http://digital.lib.uiowa.edu/u?/geoscience,34 Entelodont Skeleton