Struthiosaurus

Struthiosaurus Temporal range: Late Cretaceous, 85–66Ma PreЄ Є O S D C P T J K Pg N
Struthiosaurus occitanicus humerus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Order:	†Ornithischia
Family:	†Nodosauridae
Subfamily:	†Struthiosaurinae
Genus:	†Struthiosaurus Bunzel, 1871
Species
†S. austriacus Bunzel, 1871 (type) †S. transylvanicus Nopcsa, 1915 †S. languedocensis Garcia and Pereda-Suberbiola, 2003
Synonyms
Crataeomus Seeley, 1881 Danubiosaurus Bunzel, 1871 Leipsanosaurus Nopcsa, 1918 Pleuropeltis Seeley, 1881 Rhodanosaurus Nopsca, 1929 Hoplosaurus Seeley, 1881

Struthiosaurus (Latin struthio = ostrich + Greek sauros = lizard) is one of the smallest known and most basal genera of nodosaurid dinosaurs, from the Late Cretaceous period (Campanian-Maastrichtian) of Austria, Romania and France in Europe.^[1] It was protected by body armour. Although estimates of its length vary, it may have been as small as 2.2 metres (7.2 ft) long.^[2]

Taxonomy and Valid Species

Struthiosaurus occitanicus osteoderms

Originally described in 1871 by Emanuel Bunzel,^[3] after remains uncovered by Eduard Suess and Ferdinand Stoliczka in 1855 from the Gute Hoffnung coal mine at Muthmannsdorf near Wiener Neustadt in Austria. Many species have been referred to Struthiosaurus, most based on very fragmentary and nondiagnostic material. Three valid species are currently recognized by paleontologists: S. austriacus Bunzel, 1871, based on holotype PIWU 2349/6; S. transylvanicus Nopcsa, 1915, based on BMNH R4966, a skull and partial skeleton from Romania;^[4] and S. languedocensis Garcia and Pereda-Suberbiola, 2003, based on UM2 OLV-D50 A–G CV, a partial skeleton found in 1998 in France.^[5] Along with Hungarosaurus Ösi, 2005, Struthiosaurus is currently one of only two valid members of the Ankylosauria known from Europe.^[6]

Outdated illustration by Nopsca drawn in 1915

A number of invalid taxa have been shown to be junior synonyms of Struthiosaurus austriacus, most of them created when Harry Govier Seeley in 1881 revised the Austrian material.^[7] They include: Danubiosaurus anceps Bunzel, 1871; Crataeomus pawlowitschii Seeley, 1881; Crataeomus lepidophorus Seeley 1881; Pleuropeltis suessii Seeley, 1881; Rhadinosaurus alcimus Seeley 1881, Hoplosaurus ischyrus Seeley 1881 and Leipsanosaurus noricus Nopcsa, 1918.^[8] Another European ankylosaurid, Rhodanosaurus ludguensis Nopsca, 1929, from Campanian-Maastrichtian-age rocks of southern France, is now regarded as a nomen dubium and referred to Nodosauridae incertae sedis.

The three valid species of Struthiosaurus differ from one another in that S. austriacus is smaller than S. transylvanicus and possesses less elongate cervical vertebrae. Also, though the quadrate-paroccipital process contact is fused in S. transylvanicus, it is unfused in S. austriacus. The skull of S. languedocensis is presently unknown, but the taxon differs from S. transylvanicus in the flatter shape of the dorsal vertebrae. It differs from S. austriacus in the shape of the ischium. (Vickaryous, Maryanska, and Weishampel 2004)

Phylogeny

Struthiosaurus transsylvanicus fossil

Struthiosaurus was originally by Bunzel considered a predator that he assigned to the new taxon Ornithocephala ("Bird Heads"). The generic name is a reference to the affinities with birds shown by traits in the skull. The first to understand it represented an armoured dinosaur was Nopcsa who in 1902 placed it in the Acanthopholididae.^[9] He later corrected its name to Acanthopholidae.^[10] Walter Coombs in 1978 stated it was a nodosaurid.^[11]

Cladistic analysis of Struthiosaurus (Ösi, 2004) indicates that the taxon is a basal member of the Nodosauridae and suggests it may be one of the most basal ankylosaurs in the clade Ankylosauria.

Some workers (Vickaryous, Maryanska, and Weishampel 2004) consider the assignment of Struthiosaurus to the Nodosauridae to be provisional, pending further study and future discoveries.^[12]

References