Hadrosauroidea

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Hadrosauroids
Temporal range: Early-Late Cretaceous, 130–65.5Ma
Holotype skeleton of Tethyshadros insularis
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Ornithischia
Clade: Ornithopoda
Clade: Hadrosauriformes
Superfamily: Hadrosauroidea
Cope, 1869
Type species
Hadrosaurus foulkii
Leidy, 1858
Families

Hadrosauridae
And see text.

Synonyms

Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to Iguanodon. Many primitive hadrosauroids, such as the sail-backed Ouranosaurus, have traditionally been included in a paraphyletic (unnatural grouping) "Iguanodontidae". With cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only Iguanodon and perhaps its closest relatives.

Classification

Taxonomy

The following taxonomy follows Ramírez-Velasco et al, 2012 unless otherwise noted.[1]

Phylogeny

The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012).[3]

Hadrosauriformes 
 Iguanodontidae 

Iguanodon




Ouranosaurus



Mantellisaurus




 Hadrosauroidea 

Bolong




Equijubus




Jinzhousaurus




Altirhinus




Batyrosaurus




Probactrosaurus




Eolambia




Protohadros




Bactrosaurus



Levnesovia




Shuangmiaosaurus




Tethyshadros




Telmatosaurus



Hadrosauridae















Cladogram after Prieto-Marquez and Norell (2010).[5]

Hadrosauroidea 


Jinzhousaurus



Penelopognathus





Equijubus




Probactrosaurus





Eolambia



Protohadros





Tanius




Bactrosaurus




Gilmoreosaurus




Telmatosaurus




Shuangmiaosaurus



Nanyangosaurus



Lophorhothon



Hadrosauridae











A phylogenetic analysis performed by Ramírez-Velasco et al. (2012) found a big polytomy of all hadrosauroids which are more derived than Probactrosaurus but less derived than Hadrosauridae. The exclusion of Claosaurus, Jeyawati, Levnesovia, Nanyangosaurus, Shuangmiaosaurus and Telmatosaurus from the polytomy resulted in a more resolved topology. The cladogram below shows Huehuecanauhtlus phylogenetic position among other hadrosauroids following this topology (the relationships within Hadrosauridae aren't shown).[1]

Hadrosauroidea 

Jinzhousaurus yangi




Fukuisaurus tetoriensis




Penelopognathus weishampeli




Equijubus normani




Probactrosaurus gobiensis





Eolambia caroljonesa



Protohadros byrdi





Tanius sinensis





Bactrosaurus johnsoni



Glishades ericksoni





Gilmoreosaurus mongoliensis




Huehuecanauhtlus tiquichensis





Jintasaurus meniscus



Tethyshadros insularis




Hadrosauridae













References

  1. 1.0 1.1 Angel Alejandro Ramírez-Velasco, Mouloud Benammi, Albert Prieto-Márquez, Jesús Alvarado Ortega and René Hernández-Rivera (2012). "Huehuecanauhtlus tiquichensis, a new hadrosauroid dinosaur (Ornithischia: Ornithopoda) from the Santonian (Late Cretaceous) of Michoacán, Mexico". Canadian Journal of Earth Sciences 49 (2): 379–395. doi:10.1139/e11-062. 
  2. Hai-Lu You; and Da-Qing Li (2009). "A new basal hadrosauriform dinosaur (Ornithischia: Iguanodontia) from the Early Cretaceous of northwestern China". Canadian Journal of Earth Sciences 46 (12): 949–957. doi:10.1139/E09-067. 
  3. 3.0 3.1 3.2 Wu Wenhao and Pascal Godefroit (2012). "Anatomy and Relationships of Bolong yixianensis, an Early Cretaceous Iguanodontoid Dinosaur from Western Liaoning, China". In Godefroit, P. (eds). Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Indiana University Press. pp. 293–333. 
  4. McDonald, A. T. (2012). "Phylogeny of Basal Iguanodonts (Dinosauria: Ornithischia): An Update". In Farke, Andrew A. PLoS ONE 7 (5): e36745. doi:10.1371/journal.pone.0036745. PMC 3358318. PMID 22629328. 
  5. Albert Prieto-Marquez and Mark A. Norell (2010). "Anatomy and Relationships of Gilmoreosaurus mongoliensis (Dinosauria: Hadrosauroidea) from the Late Cretaceous of Central Asia". American Museum Novitates 3694: 1–52. ISSN 0003-0082. 
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