Elachista

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For the genus of Chordariaceae brown algae, see Elachista (alga). As invalidly described by V. Kollar in 1832, Elachista refers to the Lyonetiidae moth genus Lyonetia.
Elachista
Adult Elachista argentella
at IJmuiden (The Netherlands)
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Division: Ditrysia
Family: Elachistidae
Genus: Elachista
Treitschke in Ochsenheimer, 1833
Type species
Elachista bifasciella
Treitschke in Ochsenheimer, 1833
Species

Numerous, see text

Synonyms

Numerous, see text

Elachista is a genus of gelechioid moths. It is the type genus of the grass-miner moth family (Elachistidae). This family is sometimes (in particular in older sources) circumscribed very loosely, including for example the Agonoxenidae and Ethmiidae which seem to be quite distinct among the Gelechioidea, as well as other lineages which are widely held to be closer to Oecophora than to Elachista and are thus placed in the concealer moth family Oecophoridae here.[1]

These grass-miners are very small moths with the "feathery" hindwings characteristic of their family. They are essentially found worldwide, except in very cold places and on some oceanic islands; as usual for Gelechioidea, they are most common in the Palearctic however. They usually have at least one, sometimes as many as three light bands running from leading to trailing edge of their forewing uppersides. Some species, however, have upper forewings that are mostly white.[2]

Selected species

Not all species of this large genus have been discovered yet, let alone validly described and named. Several small genera, e.g. Biselachista and Cosmiotes, are here included in Elachista as junior synonyms (see below). They seem to be at least as closely related to E. bifasciella (the type species of the present genus) than other species commonly placed here, if not actually closer. In addition, Elachista contains several cryptic species complexes such as the one around E. dispunctella and E. triseriatella whose systematics and taxonomy are still not fully resolved.[3]

A number of Elachista species have been assigned to one of several groups, which may or may not be monophyletic, and whose names in the literature do not consistently follow the usual taxonomic practice (i.e. using as namesake the group-member which was described first). Some of these groups are placed in either of the two large subgenera Elachista (Aphelosetia) and Elachista (Elachista). Other proposed subgenera the small Elachista (Dibrachia) and Elachista (Hemiprosopa) are here included in the species of unclear relationships; like some small species-groups, they stand a chance of rendering the two larger subgenera paraphyletic if accepted as distinct, but seem too unlike them to warrant placement in either.[4]

Subgenus Elachista (Aphelosetia)

argentella group (sometimes in bedellella group)

Adult male Elachista pullicomella specimen from Tvärminne (Hanko Peninsula, Finland)

bedellella/unifasciella group (including collitella group)

Elachista (Aphelosetia) incertae sedis

Subgenus Elachista (Atachia)

These are sorted in the presumed phylogenetic sequence.[6]

gerasmia group, gerasmia section

Incertae sedis

Elachista cynopa complex

Elachista melanthes complex

Elachista gerasmia complex

Elachista paragauda complex

gerasmia group, catarata section

erebophthalma group

Subgenus Elachista (Hemiprosopa)

Subgenus Elachista (Elachista)

Adult female Elachista apicipunctella specimen from Sonkajärvi (Finland)
Adult Elachista maculicerusella specimen from Hellerup (Denmark)
Adult male Elachista gleichenella specimen from Kangasniemi (Finland)
Adult male Elachista humilis specimen from Sonkajärvi (Finland)

albifrontella/bifasciella group

cerusella/monosemiella group

gleichenella group

orba group

praelineata group

pulchella group

solena group

Adult female Elachista exactella specimen from Sonkajärvi (Finland)
Adult male Elachista exactella specimen
Adult Elachista synethes specimen from ventral

"Cosmiotes"/freyerella group

Incertae sedis

  • Elachista albipalpella Chambers, 1880
  • Elachista albrechti Kaila, 1998
  • Elachista amseli (Parenti, 1981)
  • Elachista antipetra Meyrick, 1922
  • Elachista baltica E.Hering, 1891 (sometimes in E. freyerella; tentatively placed here)
  • Elachista consortella Stainton, 1851 (tentatively placed here)
  • Elachista cristatella Chambers, 1876
  • Elachista deficiens Meyrick, 1922
  • Elachista epicaeria (Diakonoff, 1955)
  • Elachista exactella (tentatively placed here)
  • Elachista herbigrada Braun, 1925
  • Elachista heteroplaca Meyrick, 1934
  • Elachista illectella (Clemens, 1860) (type of Cosmiotes)
  • Elachista illicitella (Clemens, 1860)
  • Elachista infuscata Frey, 1882 (sometimes in E. exactella; tentatively placed here)
  • Elachista inscia (Meyrick, 1913)
  • Elachista ksarella Chrétien, 1908
  • Elachista laquaeorum (Dugdale, 1971)
  • Elachista lorigera (Meyrick, 1921) (type of Ptilodoxa)
  • Elachista nipponicella Sugisima, 2006
  • Elachista nymphaea Meyrick, 1911
  • Elachista obtusella Sruoga, 2008
  • Elachista ochroleuca Meyrick 1923
  • Elachista pallens (Sruoga, 1990)
  • Elachista planicara Kaila, 1998
  • Elachista praematurella Clemens, 1860
  • Elachista quadrata Meyrick, 1932
  • Elachista scopulicola (Braun, 1948)
  • Elachista spiculifera Meyrick, 1922
  • Elachista stabilella Stainton, 1858 (tentatively placed here)
  • Elachista stichospora Meyrick, 1932
  • Elachista tuberella Sruoga, 2008

Elachista freyerella complex[7]

Elachista synethes complex[7]

Adult female Elachista eleochariella specimen from Sonkajärvi (Finland)

tetragonella group

Elachista (Elachista) incertae sedis

  • Elachista absaroka Kaila, 1996
  • Elachista adelpha Kaila & Jalava, 1994
  • Elachista albicapilla Höfner, 1918
  • Elachista albicapitella Engel, 1907
  • Elachista albisquamella Zeller, 1877
  • Elachista amideta Braun, 1948
  • Elachista amseli (Parenti, 1981)
  • Elachista anserinelloides Nel, 2003
  • Elachista aphanta Turner, 1923
  • Elachista archaeonoma Meyrick, 1889
  • Elachista arctodyta Meyrick, 1897
  • Elachista argentifasciella Höfner, 1898
  • Elachista argentosa Braun, 1920
  • Elachista argopis Meyrick, 1897
  • Elachista arnoldi (Koster, 1993)
  • Elachista beothucella Kaila, 1996
  • Elachista bicingulella Sruoga, 1992
  • Elachista bipunctella (Sinev & Sruoga, 1995)
  • Elachista bifurcatella (Sinev & Sruoga, 1995)
  • Elachista bisetella Sinev & Sruoga, 1995
  • Elachista brachyelythrifoliella Clemens, 1864
  • Elachista brachyplectra Meyrick, 1921
  • Elachista brachypterella (Klimesch, 1990)
  • Elachista bromella Chretien, 1915
  • Elachista caelebs Meyrick, 1933
  • Elachista calusella Kaila, 1996
  • Elachista cana Braun, 1920
  • Elachista canariella Nielsen & Traugott-Olsen, 1987
  • Elachista canis Parenti, 1983
  • Elachista cataptila Meyrick, 1897
  • Elachista cerasella Kaila, 1996
  • Elachista ciliigera Kaila, 1996
  • Elachista colouratella Sinev & Sruoga, 1995
  • Elachista confirmata Meyrick, 1931
  • Elachista contaminatella Zeller, 1847
  • Elachista cornutifera (Sruoga, 1995)
  • Elachista demogenes Meyrick, 1897
  • Elachista dubitella Sinev & Sruoga, 1995
  • Elachista devexella Kaila, 2003
  • Elachista differens Parenti, 1978
  • Elachista dimicatella Rebel, 1903
  • Elachista encumeadae Kaila & Karsholt, 2002

  • Elachista ermolenkoi Sinev & Sruoga, 1995
  • Elachista exaula Meyrick, 1889
  • Elachista falaxella Sinev & Sruoga, 1995
  • Elachista fuliginea Braun, 1948
  • Elachista fumosella Sinev & Sruoga, 1995
  • Elachista geminatella (Herrich-Schäffer, 1855)
  • Elachista glenni Kaila, 1996
  • Elachista grandiferella Sruoga, 1992
  • Elachista griseella (Duponchel, 1843)
  • Elachista griseola Diakonoff, 1955
  • Elachista gruenewaldi Parenti, 2002
  • Elachista helonoma Meyrick, 1889
  • Elachista huron Kaila, 1996
  • Elachista ibericella Traugott-Olsen, 1995
  • Elachista igaloensis (Amsel, 1951)
  • Elachista inaudita Braun, 1927
  • Elachista irenae Buszko, 1989
  • Elachista irrorata Braun, 1920
  • Elachista japonica (Parenti, 1983)
  • Elachista kaszabi Parenti, 1991
  • Elachista kebneella (Traugott-Olsen, 1977)
  • Elachista kopetdagica (Sruoga, 1990)
  • Elachista kosteri Traugott-Olsen, 1995
  • Elachista lambeseella Nielsen & Traugott-Olsen, 1987
  • Elachista latebrella Sinev & Sruoga, 1995
  • Elachista lenape Kaila, 1996
  • Elachista leucofrons Braun, 1920
  • Elachista maculosella Chrétien, 1896
  • Elachista maritimella McDunnough, 1942
  • Elachista megagnathos Sruoga, 1990
  • Elachista metallica Parenti, 1981
  • Elachista minuta (Parenti, 2003)
  • Elachista mongolica Parenti, 1991
  • Elachista morandinii Huemer & Kaila, 2003
  • Elachista multidentella Sinev & Sruoga, 1995
  • Elachista napaea Philpot, 1930
  • Elachista nearcha Meyrick, 1910
  • Elachista nigrothoracella Sinev & Sruoga, 1995
  • Elachista nitidiuscula Braun, 1948
  • Elachista olgae (Sinev, 1992)
  • Elachista ombrodoca Meyrick, 1889
  • Elachista opacella Sinev & Sruoga, 1995
  • Elachista optatella Sinev & Sruoga, 1995
  • Elachista orientella Sinev & Sruoga, 1995
  • Elachista oxytypa Braun, 1948
  • Elachista parasella Traugott-Olsen, 1974
  • Elachista pelaena Kaila, 1996
  • Elachista plagiaula (Meyrick, 1938)
  • Elachista pravella (Sinev & Sruoga, 1995)
  • Elachista pusillella (Sinev & Sruoga, 1995)
  • Elachista putris Meyrick, 1923
  • Elachista pyrrha Kaila, 1996
  • Elachista radiantella Braun, 1922
  • Elachista rufella Sinev & Sruoga, 1995
  • Elachista saccharella Busck, 1934
  • Elachista sagittifera Philpott, 1927
  • Elachista sagittiferella Sinev & Sruoga, 1995
  • Elachista sasae Sinev & Sruoga, 1995
  • Elachista scythrodes (Turner, 1947)
  • Elachista serra Kaila, 1996
  • Elachista sicula Parenti, 1978
  • Elachista simplimorphella Sinev & Sruoga, 1995
  • Elachista solitaria Braun, 1922
  • Elachista staintonella Chamber, 1878
  • Elachista stenopterella Rebel, 1932
  • Elachista stramineola Braun, 1921
  • Elachista strepens Meyrick, 1922
  • Elachista sylvestris Braun, 1920
  • Elachista tabghaella Amsel, 1935
  • Elachista talgarella Kaila, 1992
  • Elachista tersella (Sinev & Sruoga, 1995)
  • Elachista thallophora Meyrick, 1889
  • Elachista vakshi (Sruoga, 1992)
  • Elachista vinlandica Kaila, 1996
  • Elachista watti Philpott, 1924

Other groups

Some Elachista are divided into groups whose relationships to the two large subgenera requires further study, including:[8]

"Dibrachia" group

"Irenicodes" group

  • Elachista antipodensis (Dugdale, 1971)
  • Elachista eurychora (Meyrick, 1919) (type of Irenicodes)
  • Elachista galatheae (Viette, 1954) (type of Euproteodes)
  • Elachista holdgatei (Bradley, 1965)
  • Elachista hookeri (Dugdale, 1971)
  • Elachista pumila (Dugdale, 1971)

Incertae sedis

Additional Elachista species are of even more uncertain relationships than usual for this genus:[9]

Placement in Elachista uncertain

  • Elachista arundinella (Duponchel, 1843
  • Elachista cupreella Blanchard, 1852
  • Elachista endobela Meyrick, 1926
  • Elachista hilda (Meyrick, 1932)
  • Elachista infamiliaris Gozmany, 1957
  • Elachista ladiniella Hartig, 1938
  • Elachista leucosoma Meyrick, 1922
  • Elachista maculoscella Clemens, 1860
  • Elachista melanura Meyrick, 1889
  • Elachista metallifera Lower, 1908
  • Elachista notosema (Meyrick, 1922)
  • Elachista oxycrates Meyrick, 1932
  • Elachista petalistis Meyrick, 1932
  • Elachista picroleuca (Meyrick, 1921)
  • Elachista rubella Blanchard, 1852
  • Elachista tersectella Zeller, 1875
  • Elachista unifasciella Chambers, 1875
  • Elachista vestata Meyrick, 1932

Excluded species

The following species are excluded from the Elachista genus, but have not yet been placed in another genus:

Synonyms

Invalid scientific names (junior synonyms and others) of Elachista in the circumscription as presented here are:[10]

  • Aphelosetia Stephens, 1834
  • Aphigalia Dyar, [1903]
  • Atachia Wocke in Heinemann, [1876]
  • Atrinia Sinev, 1992
  • Biselachista Traugott-Olsen & Schmidt Nielsen, 1977
  • Cleroptila Meyrick, 1909
  • Cosmiotes Clemens, 1860
  • Cycnodia Herrich-Schäffer, 1853
  • Dibrachia Sinev & Sruoga, 1992
  • Dicasteris Meyrick, 1906
  • Dicranoctetes Braun, 1918
  • Donacivola Busck, 1934
  • Elachistoides Sruoga, 1992
  • Elaschista (lapsus)

  • Euproteodes Viette, 1954
  • Eupneusta Bradley, 1974
  • Hecista Wallengren, 1881
  • Hemiprosopa A.F.Braun, 1948
  • Irenicodes Meyrick, 1919
  • Neaera Chambers, 1880 (non Robineau-Desvoidy, 1830: preoccupied)
  • Paraperittia Rebel, 1916
  • Phigalia Chambers, 1875 (non Duponchel, 1829: preoccupied)
  • Platyphyllis Meyrick, 1932
  • Ptilodoxa Meyrick, 1921
  • Svenssonia Parenti, 2009

Footnotes

  1. AEBR (2008), FE (2009)
  2. AEBR (2008), FE (2009), and see references in Savela (2008)
  3. Kusdas & Reichl (1973): 194, Pitkin & Jenkins (2004), AEBR (2008), FE (2009), and see references in Savela (2008)
  4. FE (2009), Wikispecies (2009-SEP-27), and see references in Savela (2008)
  5. 5.0 5.1 5.2 5.3 See references in Savela (2008)
  6. 6.0 6.1 6.2 6.3 6.4 6.5 6.6 6.7 6.8 6.9 6.10 Kaila (2011)
  7. 7.0 7.1 Sorted in the presumed phylogenetic sequence (Kaila, 2011).
  8. ABRS (2008), FE (2009), Wikispecies (2009-SEP-27)
  9. Wikispecies (2009-SEP-27), and see references in Savela (2008)
  10. AEBR (2008), FE (2009), Wikispecies (2009-SEP-27), and see references in Savela (2008)

References

Data related to Elachista at Wikispecies

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