List of eurypterids

This list of eurypterids is an attempt to create a comprehensive listing of all genera that have ever been included in the subclass Eurypterida, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomina dubia), or were not formally published (nomina nuda), as well as junior synonyms of more established names, and genera that are no longer considered eurypterids.

The list includes 81 genera, all of them extinct.

Contents 1 Naming conventions and terminology 2 Genera list 3 Classification in detail 3.1 Dubious and invalid taxa 4 See also 5 References

Naming conventions and terminology

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

• Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
• Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
• Nomen oblitum (Latin for "forgotten name"): A name that has not been used in the scientific community for more than fifty years after its original proposal.
• Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
• Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

Genera list

Genus Authors Year Status Age Location Acutiramus[1] Ruedemann 1935 valid Prid.[1] lDev.[1] Adelophthalmus[1] Jordan & Meyer 1856 valid uDev.[1] lPerm.[1] Alkenopterus[1] Størmer 1974 valid lDev.[1] Angustidontus[2] Cooper 1936 valid Baltoeurypterus[1] Størmer 1973 valid Wen.[1] Prid.[1] Bassipterus[1] Kjellesvig-Waering Leutze 1966 valid Prid.[1] Belinuropsis[2] Matthew 1909 valid Brachopterella[1] valid Prid.[1] Brachyopterus[1] Størmer 1951 valid uOrd.[1] Buffalopterus[1] Kjellesvig-Waering Heubusch 1962 valid Prid.[1] Campulocephalus[2] valid typo? Campylocephalus[1] Eichwald 1860 valid Miss.[1] Carcinosoma[1] Claypole 1890 valid Lud.[1] lDev.[1] Clarkeipterus[1] valid Prid.[1] Ctenopterus[1] Clarke Ruedemann 1912 valid Wen.[1] Curviramus[2] Ruedemann 1935 valid Dolichocephala[3] Macquart 1823 valid Dolichopterus[1] Hall 1859 valid Wen.[1] lDev.[1] Dorfopterus[1] Erik N. Kjellesvig-Waering 1955 valid lDev.[1] Drepanopterus[1] Laurie 1892 valid Lland.[1] lDev.[1] Dunsopterus[1] Waterston 1968 valid Miss.[1] Echinognathus[1] Walcott 1882 valid uOrd.[1] Eidothea[2] Scouler 1831 valid Eocarcinosoma[1] valid uOrd.[1] Erettopterus[1] Salter 1859 valid Lland.[1] lDev.[1] Erieopterus[1] valid Wen.[1] lDev.[1] Eurypterus[1] De Kay 1825 valid Lland.[1] lDev.[1] Glaucodes[2] Pruvost 1923 valid Glyptoscorpius[2] Peach 1882 valid Grossopterus[1] Størmer 1934 valid lDev.[1] Hallipterus[1] Kjellesvig-Waering 1963 valid uDev.[1] Hardieopterus[1] Waterson 1979 valid Lland.[1] Prid.[1] Hastimima[1] White 1908 valid Cis.[1] Hibbertopterus[1] valid Miss.[1] Himantopterus[2] valid Holmipterus[1] valid Wen.[1] Hughmilleria[1] Sarle 1902 valid Lud.[1] Jaekelopterus[1] valid lDev.[1] Kiaeropterus[1] valid Lland.[1] Lud.[1] Kokomopterus[1] valid Prid.[1] Lamontopterus[1] valid Lland.[1] Lanarkopterus[1] valid Wen.[1] Lud.[1] Laurieopterus[1] valid Lland.[1] Laurieipterus[2] valid Lepidoderma[2] Reuss 1855 valid Mazonipterus[2] valid Megalograptus[1] valid uOrd.[1] Melbournopterus [2] valid Moselopterus [2] valid Mixopterus[1] valid Wen.[1] Prid.[1] Moselopterus[1] valid lDev.[1] mDev.[1] Mycterops[2] Cope 1886 valid Nanahughmilleria[1] valid Lud.[1] lDev.[1] Onychopterella[4] Stoermer 1951 valid Prid.[1] Onychopterus[4] preoccupied[4] N/A N/A Orcanopterus[5] valid Pagea[1] valid lDev.[1] Paracarcinosoma[1] valid Wen.[1] lDev.[1] Parahughmillera[1] valid Lland.[1] lDev.[1] Parastylonurus[1] valid Prid.[1] Pittsfordipterus[1] valid Prid.[1] Pterygotus[1] valid Lud.[1] uDev.[1] Rhenopterus[1] Størmer 1936 valid lDev.[1] uDev.[1] Rhinocarcinosoma[1][6] Novojilov 1962 valid Lud.[1] Ruedemannipterus[1] valid uOrd.[1] Wen.[1] Salteropterus[1] Kjellesvig-Waering 1951 valid Prid.[1] Slimonia[1] Page 1856 valid Lland.[1] Prid.[1] Strobilopterus[1] Ruedemann 1935 valid lDev.[1] Stylonurella[1] valid Lland.[1] Stylonuroides[1] valid Lud.[1] Stylonurus[1] Page 1856 valid lDev.[1] Syntomopterus[1] valid lDev.[1] Tarsopterella[4] Stoermer 1951 valid Prid.[1] lDev.[1] Tarsopterus [2] Clarke and Ruedemann 1912 preoccupied[4] N/A N/A Truncatiramus[1] valid Wen.[1] Prid.[1] Tylopterella[4] Stoermer 1951 valid Lud.[1] Tylopterus[2] Clarke and Ruedemann 1912 preoccupied[4] N/A N/A Unionopterus [2] Chernyshev 1948 valid Vernopterus[2] Waterson 1968 valid Waeringopterus[1] Leutze 1961 valid Prid.[1] lDev.[1] Willwerathia[1] Størmer 1969 valid lDev.[1] Woodwardopterus[2] Kjellesvig-Waering 1959 valid

Classification in detail

As an addition to the data above, a provisional list of all published eurypterid species (including synonyms and misidentified fossils) is presented below.

• EURYPTERIDA Burmeister, 1843
  • =GIGANTOSTRACA Haeckel, 1866
  • =CYRTOCTENIDA Størmer & Waterston, 1968

STYLONURINA Diener, 1924 HIBBERTOPTEROIDEA Kjellesvig-Waering, 1959 CYRTOCTENIDAE Waterston, Oelofsen & Oosthuizen, 1985 Cyrtoctenus Størmer & Waterston, 1968 Cyrtoctenus caledonicus (Salter, 1863) — Carboniferous, Scotland Cyrtoctenus dewalquei (Fraipont, 1889) — Devonian, Belgium = Eurypterus dewalquei var. longimanus Fraipont, 1889 Cyrtoctenus dicki (Peach, 1883) — Carboniferous Thurso, Scotland Cyrtoctenus ostraviensis (Augusta & Přibyl, 1951) — Carboniferous, Czech Republic Cyrtoctenus peachi Størmer & Waterston, 1968 — Carboniferous, Scotland Cyrtoctenus wittebergensis Waterston, Oelofsen & Oosthuizen, 1985 — Carboniferous, South Africa Dunsopterus Waterston, 1968 Dunsopterus stevensoni (Etheridge Jr, 1877) — Carboniferous, Scotland Hastimima White, 1908 Hastimima whitei White, 1908 — Permian, Brazil HIBBERTOPTERIDAE Kjellesvig-Waering, 1959 Campylocephalus Eichwald, 1860 Campylocephalus oculatus (Kutorga, 1838) — Permian, Russia ?Campylocephalus salmi Stur, 1877 — Carboniferous, Czech Republic Hibbertopterus Kjellesvig-Waering, 1959 ?Hibbertopterus hibernicus (Baily, 1872) — Carboniferous, Ireland Hibbertopterus permianus Ponomarenko, 1985 — Permian, Russia Hibbertopterus scouleri (Hibbert, 1836) — Carboniferous, Scotland Vernonopterus Waterston, 1957 Vernonopterus minutisculptus (Peach, 1907) — Carboniferous, Scotland WOODWARDOPTERIDAE Kjellesvig-Waering, 1959 Megarachne Hünicken, 1980 Megarachne servinei Hünicken, 1980 — Carboniferous – Permian, Argentina Mycterops Cope, 1886 Carboniferous ?Mycterops blairi Waterston, 1968 — Carboniferous, Scotland Mycterops matthieui Pruvost, 1924 — Carboniferous, Belgium Mycterops ordinatus Cope, 1886 — Carboniferous, USA ?Mycterops whitei Schram, 1984 — Carboniferous, USA Woodwardopterus Kjellesvig-Waering, 1959 Woodwardopterus scabrosus (Woodward, 1887) — Carboniferous, Scotland STYLONUROIDEA Kjellesvig-Waering, 1959 STYLONURIDAE Diener, 1924 Kiaeropterus Waterston, 1979 Silurian Kiaeropterus cyclophthalmus (Laurie, 1892) — Silurian, Scotland Kiaeropterus ruedemanni (Størmer, 1934) — Silurian, Norway Stylonurella Kjellesvig-Waering, 1966 Stylonurella ?arnoldi (Ehlers, 1935) — Devonian, USA Stylonurella ?beecheri (Hall, 1884) — Devonian, USA Stylonurella spinipes (Page, 1859) — Silurian, Scotland = Stylonurus logani Woodward, 1872 Silurian, Scotland Stylonurus Page, 1856 Devonian Stylonurus powriensis Page, 1856 — Devonian, Scotland = Stylonurus ensiformis Woodward, 1864 — Devonian Scotland ?Stylonurus shaffneri Willard, 1933 — Devonian, USA DREPANOPTERIDAE Kjellesvig-Waering, 1966 Drepanopterus Laurie, 1892 Drepanopterus abonensis Simpson, 1951 — Devonian, England Drepanopterus pentlandicus Laurie, 1892 — Silurian, Scotland PARASTYLONURIDAE Waterston, 1979 Brachyopterus Størmer, 1951 Brachyopterus stubblefieldi Størmer, 1951 — Ordovician, Montgomeryshire Parastylonurus Kjellesvig-Waering, 1966 Parastylonurus hendersoni Waterston, 1979 — Silurian, Scotland Parastylonurus ornatus (Laurie, 1892) — Silurian, Scotland ?Parastylonurus sigmoidalis Kjellesvig-Waering, 1971 — Silurian Shropshire Stylonuroides Kjellesvig-Waering, 1966 Stylonuroides dolichopteroides (Størmer, 1934) — Silurian, Norway LAURIEIPTERIDAE Kjellesvig-Waering, 1966 Ctenopterus Clarke & Ruedemann, 1912 Ctenopterus cestrotus (Clarke, 1907) — Silurian, USA Laurieipterus Kjellesvig-Waering, 1966 Laurieipterus elegans (Laurie, 1899) — Silurian, Scotland Pagea Waterston, 1962 Devonian Pagea sturrocki Waterston, 1962 — Devonian , Scotland Pagea symondsii (Salter, 1859) — Devonian, Scotland KOKOMOPTERIDAE Kjellesvig-Waering, 1966 Kokomopterus Kjellesvig-Waering, 1966 Kokomopterus longicaudatus (Clarke & Ruedemann, 1912) — Silurian, USA Lamontopterus Waterston, 1979 Lamontopterus knoxae (Lamont, 1955) — Silurian, Scotland HARDIEOPTERIDAE Tollerton, 1989 Hallipterus Kjellesvig-Waering, 1963 Hallipterus excelsior (Hall, 1884) — Devonian, New York = Dolichocephala lacoana Claypole, 1883 — Devonian, USA Hardieopterus Waterston, 1979 ?Hardieopterus lanarkensis Waterston, 1979 — Silurian, Scotland Hardieopterus macrophthalmus (Laurie, 1892) — Silurian, Scotland Hardieopterus megalops (Salter, 1859) — Silurian, England Hardieopterus myops (Clarke, 1907) — Silurian, USA Tarsopterella Størmer, 1951 Tarsopterella scotica (Woodward, 1872) — Devonian, Scotland = ?Erieopterus brewsteri Woodward, 1864 — Devonian, Scotland = Stylonurus armatus Page, 1867 — Devonian, Scotland BRACHYOPTERELLIDAE Tollerton, 1989 Brachyopterella Kjellesvig-Waering, 1966 Brachyopterella pentagonalis (Størmer, 1934) — Silurian, Norway Brachyopterella ritchiei Waterston, 1979 — Silurian, Scotland Clarkeipterus Kjellesvig-Waering, 1966 Clarkeipterus ?otisius (Clarke, 1907) — Silurian, USA Clarkeipterus testudineus (Clarke & Ruedeman, 1912) — Silurian, USA RHENOPTERIDAE Størmer, 1951 = ALKENOPTERIDAE Poschmann & Tetlie, 2004 Alkenopterus Størmer, 1974 Alkenopterus brevitelson Størmer, 1974— Devonian, Germany Alkenopterus burglahrensis Poschmann & Tetlie, 2004 — Devonian, Germany Rhenopterus Størmer, 1936 Rhenopterus diensti Størmer, 1936 — Devonian, Germany = Rhenopterus latus Størmer, 1936 — Devonian, Germany Rhenopterus macrotuberculatus Størmer, 1974 — Devonian, Germany Rhenopterus tuberculatus Størmer, 1936 — Devonian, Germany EURYPTERINA Burmeister, 1843 Plesion taxa Moselopterus Størmer, 1974 Moselopterus ancylotelson Størmer, 1974 — Devonian, Germany Moselopterus elongatus Størmer, 1974 — Devonian, Germany Moselopterus lancmani (Delle, 1937) — Devonian, Latvia Onychopterella Størmer, 1951 Onychopterella augusti Braddy, Aldridge & Theron, 1995 — Ordovician, South Africa Onychopterella kokomoensis (Miller & Gurley, 1896) — Silurian, USA = Eurypterus ranilarva Clarke & Ruedemann, 1912 — Silurian, USA ?Onychopterella pumilus (Savage, 1916) — Silurian, USA Vinetopterus Poschmann & Tetlie, 2004 Vinetopterus martini Poschmann & Tetlie, 2004 — Devonian, Germany Vinetopterus struvei (Størmer, 1974) — Devonian, Germany Plesion taxa currently assigned to Drepanopterus ?Drepanopterus bembycoides Laurie, 1899 — Silurian, Scotland ?Drepanopterus conicus Laurie, 1892 — Silurian, Scotland ?Drepanopterus latus (Størmer, 1934) — Silurian, Norway ?Drepanopterus lobatus Laurie, 1899 — Silurian, Scotland ?Drepanopterus nodosus Kjellesvig-Waering & Leutze, 1966 — Silurian, USA MEGALOGRAPTOIDEA Caster & Kjellesvig-Waering, 1955 MEGALOGRAPTIDAE Caster & Kjellesvig-Waering, 1955 Echinognathus Walcott, 1882 Echinognathus clevelandi (Walcott, 1882) — Ordovician, USA Megalograptus Miller, 1874 Megalograptus alveolatus (Shuler, 1915) — Ordovician, USA Megalograptus ohioensis Caster & Kjellesvig-Waering, 1955 — Ordovician, USA Megalograptus shideleri Caster & Kjellesvig-Waering, 1964 — Ordovician, USA Megalograptus welchi Miller, 1874 — Ordovician, USA Megalograptus williamsae Caster & Kjellesvig-Waering, 1964 — Ordovician, USA EURYPTEROIDEA Burmeister, 1843 DOLICHOPTERIDAE Kjellesvig-Waering & Størmer, 1952 Dolichopterus Hall, 1859 Dolichopterus gotlandicus Kjellesvig-Waering, 1979 — Silurian, Sweden Dolichopterus jewetti Caster & Kjellesvig-Waering, 1956 — Silurian, USA Dolichopterus macrocheirus Hall, 1859 — Silurian, USA, Canada Dolichopterus siluriceps Clarke & Ruedemann, 1912 — Silurian, USA, Canada ?Dolichopterus stoermeri Caster & Kjellesvig-Waering, 1956 — Silurian, Estonia Ruedemannipterus Kjellesvig-Waering, 1966 Ruedemannipterus stylonuroides (Clarke & Ruedemann, 1912) — Silurian, USA Buffalopterus Kjellesvig-Waering & Heubusch, 1962 Buffalopterus pustulosus (Hall, 1859) — Silurian, USA & Canada = Eurypterus giganteus Pohlman, 1882 — Silurian, USA & Canada = Pterygotus globicaudatus Pohlman, 1882 — Silurian, USA & Canada Strobilopterus Ruedemann, 1935 Strobilopterus princetonii (Ruedemann, 1934) — Devonian, USA Syntomopterus Kjellesvig-Waering, 1961 Syntomopterus richardsoni Kjellesvig-Waering, 1961 — Devonian, USA EURYPTERIDAE Burmeister, 1843 Eurypterus de Kay, 1825 = Baltoeurypterus Størmer, 1973 ?Eurypterus cephalaspis Salter, 1856 — Silurian, England Eurypterus dekayi Hall, 1859 — Silurian, USA & Canada Eurypterus flintstonensis Swartz, 1923 — Silurian, USA Eurypterus hankeni Tetlie, 2006 — Silurian, Norway Eurypterus henningsmoeni (Tetlie, 2002) — Silurian, Norway Eurypterus laculatus Kjellesvig-Waering, 1958 — Silurian, USA & Canada Eurypterus lacustris Harlan, 1834 — Silurian, USA & Canada = Eurypterus pachycheirus Hall, 1859 — Silurian, USA & Canada = Eurypterus robustus Hall, 1859 — Silurian, USA & Canada Eurypterus leopoldi Tetlie, 2006 — Silurian, Canada Eurypterus megalops Clarke & Ruedemann, 1912 — Silurian, USA ?Eurypterus minor Laurie, 1899 — Silurian, Scotland Eurypterus ornatus Leutze, 1958 — Silurian, USA Eurypterus pittsfordensis Sarle, 1903 — Silurian, USA Eurypterus quebecensis Kjellesvig-Waering, 1958 — Silurian, Canada Eurypterus remipes DeKay, 1825 — Silurian, USA, Canada = Carcinosoma trigona (Ruedemann, 1916) — Silurian, USA Eurypterus serratus (Jones & Woodward, 1888) — Silurian, Sweden Eurypterus tetragonophthalmus Fischer, 1839 — Silurian, Ukraine = Eurypterus fischeri Eichwald, 1854 — Silurian, Ukraine = Eurypterus fischeri var. rectangularis Schmidt, 1883 — Silurian, Estonia ERIEOPTERIDAE Tollerton, 1989 Erieopterus Kjellesvig-Waering, 1958 Erieopterus eriensis (Whitfield, 1882) — Silurian, USA Erieopterus hypsophthalmus Kjellesvig-Waering, 1958 — Silurian, USA ?Erieopterus laticeps (Schmidt, 1883) — Silurian, Norway Erieopterus latus Ruedemann, 1935 — Devonian, USA ?Erieopterus limuloides (Kjellesvig-Waering, 1948) — Silurian, USA Erieopterus microphthalmus (Hall, 1859) — Devonian, USA & Canada ?Erieopterus phillipsensis Copeland, 1971 — Silurian. Canada ?Erieopterus statzi Størmer, 1936 —Devonian, Germany ?Erieopterus turgidus Stumm & Kjellesvig-Waering, 1962 — Silurian, USA MIXOPTEROIDEA Caster & Kjellesvig-Waering, 1955 CARCINOSOMATIDAE Størmer, 1934 Carcinosoma Claypole, 1890 ?Carcinosoma harleyi Kjellesvig-Waering, 1961 — Silurian ,England Carcinosoma libertyi Copeland & Bolton, 1960 — Silurian, Canada Carcinosoma newlini Claypole, 1890 — Silurian, USA = Carcinosoma ingens Claypole, 1894 — Silurian, USA ?Carcinosoma punctatum (Salter in Huxley & Salter, 1859) — Silurian ,England Carcinosoma scorpioides (Woodward, 1868) — Silurian, Scotland = Pterygotus raniceps Woodward, 1868 — Silurian, Scotland Carcinosoma scoticus (Laurie, 1899) — Silurian, Scotland ?Carcinosoma spiniferumKjellesvig-Waering & Heubusch, 1962 — Silurian, USA Eocarcinosoma Caster & Kjellesvig-Waering, 1964 Eocarcinosoma batrachophthalmus Caster & Kjellesvig-Waering, 1964 — Ordovician, USA

MIXOPTEROIDEA Caster & Kjellesvig-Waering, 1955 (cont.) CARCINOSOMATIDAE Størmer, 1934 (cont.) Paracarcinosoma Caster & Kjellesvig-Waering, 1964 Paracarcinosoma acrocephalus (Semper, 1898) — Silurian – Devonian Barrandian area Paracarcinosoma obesus (Woodward, 1868) — Silurian, Scotland Paracarcinosoma scorpionis (Grote & Pitt, 1875) — Silurian, USA, Canada Rhinocarcinosoma Novojilov, 1962 Rhinocarcinosoma cicerops (Clarke, 1907) — Silurian, USA Rhinocarcinosoma dosonensis Braddy, Selden & Doan Nhat, 2002 — Silurian, Vietnam Rhinocarcinosoma vaningeni (Clarke & Ruedemann, 1912) — Silurian, USA MIXOPTERIDAE Caster & Kjellesvig-Waering, 1955 = LANARKOPTERIDAE Tollerton, 1989 Lanarkopterus Ritchie, 1968 Lanarkopterus dolichoschelus (Størmer, 1936) — Silurian, Scotland Mixopterus Ruedemann, 1921 Mixopterus kiaeri Størmer, 1934 — Silurian, Norway Mixopterus multispinosus (Clarke & Ruedemann, 1912) — Silurian, USA Mixopterus simonsoni Schmidt, 1883 — Silurian, Estonia WAERINGOPTEROIDEA author, date? families? NB: the superfamily name appears to be derived from a family group name proposed in a thesis Grossopterus Størmer, 1934 Grossopterus overathi (Gross, 1933) — Devonian, Germany Grossopterus inexpectans (Ruedemann, 1921) — Devonian, USA Orcanopterus Stott, Tetlie, Braddy, Nowlan, Glasser & Devereux, 2005 Orcanopterus manitoulinensis Stott, Tetlie, Braddy, Nowlan, Glasser & Devereux, 2005 — Ordovician, Canada Waeringopterus Leutze, 1961 Waeringopterus apfeli Leutze, 1961 — Silurian, Canada Waeringopterus cumberlandicus (Swartz, 1923) — Silurian, USA = Eurypterus swartzi Kjellesvig-Waering, 1958 — Silurian, USA ADELOPHTHALMOIDEA Tollerton, 1989 ADELOPHTHALMIDAE Tollerton, 1989 Adelopthalmus Jordan in Jordan & von Mayer, 1854 = Lepidoderma Reuss, 1855 = Anthraconectes Meek & Worthen, 1868 = Polyzosternites Goldenberg, 1873 = Glyptoscorpius Peach, 1882 Adelophthalmus approximatus (Hall & Clarke, 1888) — Carboniferous, USA Adelophthalmus asturica (Melendez, 1971) — Carboniferous, Spain Adelophthalmus bradorensis (Bell, 1922) — Carboniferous, New Zealand Adelophthalmus cambieri (Pruvost, 1930) — Carboniferous, Belgium ?Adelophthalmus carbonarius (Chernyshev, 1933) — Carboniferous, Ukraine Adelophthalmus chinensis (Grabau, 1920) — Carboniferous – Permian, China Adelophthalmus corneti (Pruvost, 1939) — Carboniferous, Belgium Adelophthalmus douvillei (de Lima, 1890) — Permian Bussaco, Portugal Adelophthalmus dumonti (Stainier, 1917) — Carboniferous, Belgium Adelophthalmus granosus Jordan in Jordan & von Meyer, 1854 — Carboniferous, Germany Adelophthalmus imhofi (Reuss, 1855) — Carboniferous, Czech Republic Adelophthalmus irinae Shpinev, 2006 — Carboniferous, Russia Adelophthalmus kidstoni (Peach, 1888) — Carboniferous, England ?Adelophthalmus lohesti (Dewalque in Fraipont 1889) — Devonian, Belgium Adelophthalmus luceroensis Kues & Kietzke, 1981 — Permian, USA Adelophthalmus mansfieldi (Hall, 1877) — Carboniferous, USA = Eurypterus stylus Hall, 1884 — Carboniferous Pennsylvania, USA Adelophthalmus mazonensis (Meek & Worthen, 1868) — Carboniferous, USA Adelophthalmus moyseyi (Woodward, 1907) — Carboniferous, Wales = Eurypterus derbiensis Woodward, 1907 — Carboniferous, England Adelophthalmus nebraskensis (Barbour, 1914) — Permian, USA Adelophthalmus pennsylvanicus (Hall, 1877) — Carboniferous, USA ?Adelophthalmus ?perornatus (Peach, 1882) — Carboniferous, Scotland Adelophthalmus pruvosti Kjellesvig-Waering, 1948 — Carboniferous, France ?Adelophthalmus raniceps Goldenberg, 1873 — Carboniferous, Germany Adelophthalmus sellardsi (Dunbar, 1924) — Permian, USA Adelophthalmus sievertsi (Størmer, 1969) — Devonian, Germany = ?Eurypterus trapezoides Størmer, 1974 — Devonian, Germany Adelophthalmus waterstoni (Tetlie et al., 2004) — Devonian, Australia Adelophthalmus wilsoni (Woodward, 1888) — Carboniferous, England Adelophthalmus zadrai Přibyl, 1952 — Carboniferous, Poland / Czech Republic Bassipterus Kjellesvig-Waering & Leutze, 1966 Bassipterus virginicus Kjellesvig-Waering & Leutze, 1966 — Silurian, USA Eysyslopterus Tetlie & Poschmann, 2008 Eysyslopterus patteni (Størmer, 1934) — Silurian, Estonia Nanahughmilleria Kjellesvig-Waering, 1961 Nanahughmilleria clarkei Kjellesvig-Waering, 1964b — Silurian, USA Nanahughmilleria norvegica (Kiær, 1911) — Silurian, Norway = Eurypterus minutus Kiær, 1911 — Silurian, Norway ?Nanahughmilleria prominens (Hall, 1884) — Silurian, USA Nanahughmilleria pygmaea (Salter, 1859) — Silurian, England ?Nanahughmilleria schiraensis (Pirozhnikov, 1957) — Devonian, Russia Parahughmilleria Kjellesvig-Waering, 1961 Parahughmilleria bellistriata (Kjellesvig-Waering, 1950) — Silurian, USA Parahughmilleria hefteri Størmer, 1973 — Devonian, Germany Parahughmilleria maria (Clarke, 1907) — Silurian, USA Parahughmilleria matarakensis (Pirozhnikov, 1957) — Devonian, Russia Parahughmilleria salteri Kjellesvig-Waering, 1961 — Silurian, England Pittsfordipterus Kjellesvig-Waering & Leutze, 1966 Pittsfordipterus phelpsae (Ruedemann, 1921) — Silurian, USA PTERYGOTIOIDEA Clarke & Ruedemann, 1912 HUGHMILLERIIDAE Kjellesvig-Waering, 1951 Herefordopterus Tetlie, 2006b Herefordopterus banksii (Salter, 1856) — Silurian, England = Eurypterus acuminatus Salter, 1859a — Silurian, England Hughmilleria Sarle, 1903 Hughmilleria shawangunk Clarke, 1907 — Silurian, USA Hughmilleria socialis Sarle, 1903 — Silurian, USA = Hughmilleria robusta Sarle, 1903 — Silurian, USA Hughmilleria wangi Tetlie, Selden & Ren, 2007 — Silurian, China SLIMONIDAE Novojilov, 1968 Salteropterus Kjellesvig-Waering, 1951 Salteropterus abbreviatus (Salter, 1859) — Silurian, England Slimonia Page, 1856 Slimonia acuminata Salter, 1856 — Silurian, Scotland = Himantopterus maximus Salter, 1856 — Silurian, Scotland Slimonia boliviana Kjellesvig-Waering, 1973 — Silurian, Bolivia Slimonia dubia Laurie, 1899 — Silurian, Scotland PTERYGOTIDAE Clarke & Ruedemann, 1912 = JAEKELOPTERIDAE Størmer, 1974 Acutiramus Ruedemann, 1935 Acutiramus bohemicus (Barrande, 1872) — Silurian, Czech Republic = Pterygotus comes Barrande, 1872 — Silurian, Czech Republic = Pterygotus mediocris Barrande, 1872 — Silurian, Czech Republic = Pterygotus blahai Semper, 1898 — Silurian, Czech Republic = Pterygotus fissus Seemann, 1906 — Silurian, Czech Republic Acutiramus cummingsi (Grote & Pitt, 1875) — Silurian USA & Canada = Pterygotus acuticaudatus Pohlman, 1882 — Silurian, USA = Pterygotus buffaloensis Pohlman, 1881 — Silurian, USA = Pterygotus quadraticaudatus Pohlman, 1882 — Silurian, USA Acutiramus floweri Kjellesvig-Waering & Caster, 1955 — Silurian, USA Acutiramus macrophthalmus (Hall, 1859) — Silurian USA & Canada = Pterygotus osborni Hall, 1859 — Silurian , USA = Pterygotus cobbi var. juvenis Clarke & Ruedemann, 1912 — Silurian, USA Acutiramus perneri Chlupáč, 1994 — Devonian, Czech Republic Acutiramus perryensis Leutze, 1958 — Silurian, USA Acutiramus suwanneensis Kjellesvig-Waering, 1955 — Silurian?, USA Erettopterus Salter in Huxley & Salter, 1859 Erettopterus bilobus (Salter, 1856) — Silurian, Scotland = Eurypterus perornatus Salter, 1856 — Silurian , Scotland = Pterygotus bilobus var. acidens Woodward, 1878 — Silurian , Scotland = Pterygotus bilobus var. crassus Woodward, 1878 — Silurian, Scotland = Pterygotus bilobus var. inornatus Woodward, 1878 — Silurian, Scotland = Pterygotus bilobus var. perornatus Woodward, 1878 — Silurian, Scotland = Pterygotus perornatus var. plicatissimus Salter in Huxley & Salter, 1859 — Silurian, Scotland Erettopterus brodiei Kjellesvig-Waering, 1961 — Silurian, England Erettopterus canadensis (Dawson, 1879) — Silurian, Canada Erettopterus exophthalmus Kjellesvig-Waering & Leutze, 1966 — Silurian, USA Erettopterus gigas Salter in Huxley & Salter, 1859 — Silurian, England Erettopterus globiceps Clarke & Ruedemann, 1912 — Silurian, USA Erettopterus grandis Pohlman, 1881 — Silurian, USA Erettopterus holmi (Størmer, 1934) — Silurian, Norway Erettopterus laticauda Schmidt, 1883 — Silurian, Estonia Erettopterus marstoni Kjellesvig-Waering, 1961 — Silurian, England Erettopterus megalodon Kjellesvig-Waering, 1961 — Silurian, England Erettopterus osiliensis Schmidt, 1883 — Siluria, Estonia, Erettopterus saetiger Kjellesvig-Waering, 1964 — Silurian, USA Erettopterus serratus Kjellesvig-Waering, 1961 — Devonian, USA Erettopterus spatulatus Kjellesvig-Waering, 1961 — Silurian, England ?Erettopterus vogti Størmer, 1934a — Devonian, Norway Erettopterus waylandsmithi Kjellesvig-Waering & Caster, 1955 — Silurian, USA Jaekelopterus Waterston, 1964 Jaekelopterus rhenaniae (Jaekel, 1914) — Devonian, Germany Pterygotus Agassiz, 1839 Pterygotus anglicus Agassiz, 1844 — Devonian, Scotland & Canada = Pterygotus atlanticus Clarke & Ruedemann, 1912 — Devonian, Canada = Pterygotus minor Woodward, 1864 — Devonian , Scotland Pterygotus arcuatus Salter in Huxley & Salter, 1859 — Devonian, England ?Pterygotus australis McCoy, 1899 — Silurian, Australia Pterygotus barrandei Semper, 1898 — Silurian, Czech Republic = Pterygotus beraunensis Semper, 1898 — Silurian, Czech Republic Pterygotus bolivianus Kjellesvig-Waering, 1964 — Devonian, Bolivia Pterygotus carmani Kjellesvig-Waering, 1961 — Devonian, USA Pterygotus cobbi Hall, 1859 — Silurian, USA, Canada Pterygotus denticulatus Kjellesvig-Waering, 1961 — Silurian, England Pterygotus floridanus Kjellesvig-Waering, 1950 — Devonian, USA Pterygotus gaspesiensis Russell, 1953 — Devonian, Canada ?Pterygotus grandidentatus Kjellesvig-Waering, 1961 — Silurian England ?Pterygotus howelli Kjellesvig-Waering & Størmer, 1952 — Devonian, USA ?Pterygotus impacatus Kjellesvig-Waering, 1964a — Silurian, Estonia Pterygotus kopaninensis Barrande, 1872 — Silurian, Czech Republic Pterygotus lanarkensis Kjellesvig-Waering, 1964 — Silurian, Scotland Pterygotus lightbodyi Kjellesvig-Waering, 1961 — Silurian England Pterygotus ludensis Salter in Huxley & Salter, 1859 — Silurian, England Pterygotus marylandicus Kjellesvig-Waering, 1964 — Silurian, USA Pterygotus monroensis Sarle 1902 — Silurian, USA ?Pterygotus sarlei Ciurca & Tetlie, 2007 — Silurian, USA ?Pterygotus ventricosus Kjellesvig-Waering, 1948 — Silurian, USA EURYPTERIDA incertae sedis Dorfopterus Kjellesvig-Waering, 1955 Dorfopterus angusticollis Kjellesvig-Waering, 1955 — Devonian, USA ?Dolichopterus ?Dolichopterus asperatus Kjellesvig-Waering, 1961 — Devonian, USA ?Dolichopterus bulbosus Kjellesvig-Waering, 1961 — Silurian, England ?Dolichopterus herkimerensis Caster & Kjellesvig-Waering, 1956 — Silurian ,USA & Canada ?Eurypterus ?Eurypterus loi Chang, 1957 [non eurypterid?] — Silurian, China ?Eurypterus podolicus Chernyshev, 1947 — Silurian Ukraine ?Eurypterus satpaevi Simorin, 1956 — Carboniferous, Kazakhstan ?Eurypterus styliformis Chang, 1957 [non eurypterid?] — Silurian, China ?Eurypterus tschernyschevi Simorin, 1956 — Carboniferous, Kazakhstan ?Eurypterus yangi Chang, 1957 [non eurypterid?] — Silurian, China Holmipterus Kjellesvig-Waering, 1979 Holmipterus suecicus Kjellesvig-Waering, 1979 — Silurian, Sweden Marsupipterus Caster & Kjellesvig-Waering, 1955 Marsupipterus sculpturatus Caster & Kjellesvig-Waering, 1955 — Silurian, England ?Nanahughmilleria ?Nanahughmilleria lanceolata Salter, 1856 — Silurian, Scotland = Eurypterus chartarius Salter, 1859 — Silurian, Scotland = Eurypterus linearis Salter, 1859 — Silurian, Scotland ?Salteropterus ?Salteropterus longilabium Kjellesvig-Waering, 1961 — Silurian, England ?Stylonurus ?Stylonurus perspicillum Størmer, 1969 — Devonian, Germany Tylopterella Størmer, 1951 Tylopterella boylei (Whiteaves, 1884) — Silurian, Canada ?Tylopterella menneri (Novojilov, 1959) — Devonian, Russia Unionopterus Chernyshev, 1948 Unionopterus anastasiae Chernyshev, 1948 — Carboniferous, Kazakhstan

Dubious and invalid taxa

Nomina dubia Bunodella horrida Matthew, 1888 [non Xiphosura] — Silurian, Canada ?Dunsopterus wrightianus Dawson 1881 — Devonian, USA Eurypterella ornata Matthew, 1888 — Carboniferous, Canada Eurypterus potens Hall, 1884 — Carboniferous, USA Eurypterus pulicaris Salter, 1863 — Devonian, Canada Hastimima sewardi Strand, 1926 — Devonian South Africa ?Pterygotus formosus Dawson, 1871 — Devonian, Canada Pterygotus nobilis Barrande, 1872 — S Barrandian area Pterygotus siemiradzkii Strand, 1926 — Devonian, Ukraine Pterygotus taurinus Salter, 1868 — Silurian, UK ?Slimonia stylops Salter in Huxley & Salter, 1859 — Silurian, England Nomina nuda Baltoeurypterus latus Hanken & Størmer, 1975 — Silurian, Norway Pterygotus mcgrewi Kjellesvig-Waering, 1986 — Devonian, USA Nomina vana Pterygotus problematicus Agassiz, 1844 — Silurian, UK Misidentifications Buffalopterus verrucosus Kjellesvig-Waering & Heubusch, 1962 [crustacean] — Ordovician, USA Carcinosoma ?logani(Williams, 1915) [crustacean] — Silurian, Canada Eurypterus (Stylonurus?) maccarthyi Kjellesvig-Waering, 1934 [cephalopod] — Devonian, USA Eurypterus pugioBarrande, 1872 [crustacean] — Silurian, Czech Republic Eurypterus thomasi Walter, 1924 [aglaspidid] — Cambrian, USA Kockurus grandis Chlupáč, 1995 [?aglaspidid] — Cambrian, Czech Republic Kodymirus vagans Chlupáč & Havlíček, 1965 [?aglaspidid] — Cambrian, Czech Republic Mazonipterus cyclophthalmusKjellesvig-Waering, 1963b [plant] — Carboniferous, USA Pterygotus expectatus Barrande, 1872 [crustacean] — Silurian, Czech Republic Pterygotus (Curviramus) elleri Ruedemann, 1935 [crustacean] — Devonian, USA Pterygotus (Curviramus) montanensis Ruedemann, 1935 [crustacean] — Devonian, USA Pterygotus (Leptocheles) leptodactylum M’Coy, 1849 [crustacean] — Silurian, England

Pseudofossils Brachyopterella magna (Clarke & Ruedemann, 1912) — Ordovician, USA ?Carcinosoma linguata (Clarke & Ruedemann, 1912) — Ordovician, USA ?Carcinosoma longiceps (Clarke & Ruedemann, 1912) — Ordovician, USA Dolichopterus antiquus Ruedemann, 1942 — Ordovician, USA Dolichopterus frankfortensis (Clarke & Ruedemann, 1912) — Ordovician, USA Dolichopterus insolitusRuedemann, 1926 — Ordovician, USA ?Dolichopterus stellatus (Clarke & Ruedemann, 1912) — Ordovician, USA ?Drepanopterus ruedemanni (O’Connell, 1916) — Ordovician, USA ?Eocarcinosoma breviceps (Ruedemann, 1926) — Ordovician, USA Eocarcinosoma ruedemanni (Flower, 1945) — Ordovician, USA Eocarcinosoma triangulatus (Clarke & Ruedemann, 1912) — Ordovician, USA Erettopterus walcotti (Ruedemann, 1926) — Ordovician, USA Erieopterus chadwicki (Clarke & Ruedemann, 1912) — Ordovician, USA Erieopterus hudsonicus (Ruedemann, 1934) — Ordovician, USA ?Eurypterus decepiens (Ruedemann, 1942) — Ordovician, USA Eurypterus indicus Dubey, 1985 — Precambrian, India ?Eurypterus pristinus (Clarke & Ruedemann, 1912) — Ordovician, USA Eurypterus vermai Dubey, 1985 — Precambrian, India Hughmilleria chiplonkariDubey, 1985 — Precambrian, India Hughmilleria kilfoylei Ruedemann, 1934 — Ordovician, USA Hughmilleria prisca Ruedemann, 1934 — Ordovician, USA Hughmilleria uticana Ruedemann, 1926 — Ordovician, USA Parastylonurus rusti (Ruedemann, 1926) — Ordovician, USA Pterygotus deepkillensisRuedemann, 1934 — Ordovician, USA Pterygotus nasutusClarke & Ruedemann, 1912 — Ordovician, USA ?Pterygotus normanskillensis Clarke & Ruedemann, 1912 — Ordovician, USA Ruedemannipterus breviceps (Clarke & Ruedemann, 1912) — Ordovician, USA Ruedemannipterus latifrons(Clarke & Ruedemann, 1912) — Ordovician, USA Stylonurella modestus (Clarke & Ruedemann, 1912) — Ordovician, USA Stylonuroides limbatus (Clarke & Rudemann, 1912) — Ordovician, USA ?Waeringopterus pristinus (Ruedemann, 1942) — Ordovician, USA Waeringopterus prolificus (Clarke & Ruedemann, 1912) — Ordovician, USA

See also

• Chelicerates
• Eurypterids

References