Staffordiidae

Staffordiidae
drawing of apertural view of the shell of Staffordia daflaensis
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
(unranked): clade Heterobranchia
clade Euthyneura
clade Panpulmonata
clade Eupulmonata
clade Stylommatophora
informal group Sigmurethra
clade limacoid clade
Superfamily: Staffordioidea
Thiele, 1931
Family: Staffordiidae
Thiele, 1931[1]
Genera

See text

Staffordiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Staffordioidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Staffordiidae is the only family in the superamily Staffordioidea. This family has no subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Staffordiidae is poorly known[2] family, because it occur only in Dafla Hills area, India, that is very insufficiently known.[2]

Various sources consider Staffordiidae as a part of Dyakiidae[3] or Ariophantidae/Dyakiinae.[4]

Contents

Distribution

Distribution of Staffordiidae include only India in Dafla Hills.[2]

This area is close to northern margin of Indian plate.[2] The historical area of origin of Staffordiidae is not surely known, because the possible coastal area in southern Asia is considered to became uninhabitable[2] after Indian Plate and Eurasian Plate collision 50 and 55 million years ago. The original ancestral area of limacoid families is thought to be Palaearctic region and south-eastern Asia.[2] So it has been hypothetized that Staffordiidae have colonized its actual area from southern margin of Asian part of Eurasian Plate in Oligocene.[2]

Genera

Genera within the family Staffordiidae include:

The generic name Staffordia is in honor of Brigadier-General Stafford, who was in command of the punitive force which entered the Dafla Hills for the first time in the winter of 1874-1875.[5]

The foot of Staffordia is pointed.[5] The peripodial margin is simple with a narrow pale margin.[5] There are present small right and left shell-lobes.[5]

Reproductive system of Staffordia: the dart-sac is small, globose, with a long cord-like attachment to a coronal gland.[5] The penis is simple.[5] The spermatheca is long.[5]

Radula of Staffordia is with aculeate laterals.[5]

Comparison of shells of Staffordia species:

Cladogram

Staffordiidae is considered as sister group of all other families in the limacoid clade.[2]

The following cladogram shows the phylogenic relationships of this family and superfamily to the other families within the limacoid clade:[2]

 limacoid clade 
 Staffordioidea 

Staffordiidae




 Dyakioidea 

Dyakiidae


 Gastrodontoidea 

Pristilomatidae




Chronidae




Euconulidae



Trochomorphidae





Gastrodontidae



Oxychilidae







 Parmacelloidea 

Trigonochlamydidae



Parmacellidae



Milacidae





 Zonitoidea 

Zonitidae


 Helicarionoidea 

Helicarionidae




Ariophantidae



Urocyclidae





 Limacoidea 

Vitrinidae




Boettgerillidae




Limacidae



Agriolimacidae









References

This article incorporates public domain text from the reference.[5]

  1. ^ Thiele J. (1931). Handbuch der systematischen Weichtierkunde Fischer, Jena, 1(2): 632.
  2. ^ a b c d e f g h i Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
  3. ^ Barker G. M. (2001) Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. 1-146. In: Barker G. M. (ed.) (2001) The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, cited pages: 139-144. ISBN 0-85199-318-4.
  4. ^ a b c d Ramakrishna, Dey A. & Mitra S. C. (PDF created 6 April 2010). "Checklist of Indian Land Mollusca". Zoological Survey of India. accessed 30 June 2010. 65 pp.
  5. ^ a b c d e f g h i j k Godwin-Austen H. H. (1907). Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean; Supplementary to Masers Theobald and Hanley's Conchologica Indica. Taylor and Francis, London. 2: page 184, plate CXIII.

Further reading