Schizothorax
Schizothorax is a genus of cyprinid fish from Central and East Asia. Their scientific name means "cleft-breast", from Ancient Greek schízeïn (σχίζειν) "to cleave" + thórax (θώραξ) "breast-plate" (see also thorax). The western species are typically referred to as marinkas from their Russian name marinka (маринка; plural: маринки marinki), while the eastern species are usually called "snowtrouts". But though they do resemble trouts in habitus this is merely due to convergent evolution and they are by no means closely related apart from both being Teleostei: Cyprinids are in the teleost superorder Ostariophysi, while trouts are in the superorder Protacanthopterygii. Their ancestors must thus have diverged as early as the Triassic, more than 200 million years ago. The type species of Schizothorax is the Asaila (S. plagiostomus).[1]
Other "snowtrouts" – most if not all presumably closely related to Schizothorax – are found in the genera Diptychus, Ptychobarbus, Schizopyge and Schizopygopsis. The latter two were formerly often included in the present genus, and their delimitation is still not entirely clear. In any case, the genus Schizothorax (and other "snowtrouts") are in turn related to such cyprinids as the typical barbels (Barbus sensu stricto and the doubtfully distinct Luciobarbus and Messinobarbus), Carasobarbus, Labeobarbus (yellowfish), Aulopyge (Dalmatian Barbelgudgeon), and Cyprinion. They were thus placed in the subfamily Barbinae, which is quite paraphyletic however and thus here merged with the Cyprininae at least for the largest part (including the marinkas), becoming its junior synonym in the process.[2]
Species
Some important subspecies are also listed. New species of Schizothorax are still being discovered occasionally:
- Schizothorax argentatus Kessler, 1874 (Balkhash Marinka)
- Schizothorax beipanensis J. Yang, X. Y. Chen & J. X. Yang, 2009
- Schizothorax biddulphi Günther, 1876
- Schizothorax chongi (P. W. Fang, 1936)
- Schizothorax cryptolepis T. Y. Fu & M. R. Ye, 1984
- Schizothorax curvilabiatus (Wu & Tsao, 1992)
- Schizothorax davidi (Sauvage, 1880)
- Schizothorax dolichonema Herzenstein, 1889
- Schizothorax dulongensis S. Y. Huang, 1985
- Schizothorax edeniana McClelland, 1842
- Schizothorax elongatus S. Y. Huang, 1985
- Schizothorax esocinus Heckel, 1838 (Chirruh snowtrout)
- Schizothorax eurystomus Kessler, 1872
- Schizothorax gongshanensis W. H. Tsao, 1964
- Schizothorax grahami (Regan, 1904)
- Schizothorax griseus Pellegrin, 1931
- Schizothorax heterochilus M. R. Ye & T. Y. Fu, 1986
- Schizothorax heterophysallidos J. Yang, X. Y. Chen & J. X. Yang, 2009
- Schizothorax huegelii Heckel, 1838
- Schizothorax integrilabiatus (Wu et al., 1992)
- Schizothorax kozlovi Nikolskii, 1903
- Schizothorax kumaonensis Menon, 1971 (Kumaon Snowtrout)
- Schizothorax labiatus (McClelland, 1842) (Kunar Snowtrout)
- Schizothorax labrosus Y. H. Wang, D. D. Zhuang & L. C. Gao, 1981
- Schizothorax lantsangensis W. H. Tsao, 1964
- Schizothorax lepidothorax J. X. Yang, 1991
- Schizothorax lissolabiatus W. H. Tsao, 1964
- Schizothorax longibarbus (P. W. Fang, 1936)
- Schizothorax macrophthalmus Terashima, 1984 (Nepalese Snowtrout)
- Schizothorax macropogon Regan, 1905
- Schizothorax malacanthus S. Y. Huang, 1985
- Schizothorax meridionalis W. H. Tsao, 1964
- Schizothorax microcephalus F. Day, 1877
- Schizothorax microstomus S. Y. Hwang, 1982
- Schizothorax molesworthi (Chaudhuri, 1913)
- Schizothorax myzostomus W. H. Tsao, 1964
- Schizothorax nasus Heckel, 1838 (Dongu Snowtrout)
- Schizothorax nepalensis Terashima, 1984
- Schizothorax ninglangensis Y. H. Wang, K. X. Zhang & D. D. Zhuang, 1981
- Schizothorax nudiventris J. Yang, X. Y. Chen & J. X. Yang, 2009
- Schizothorax nukiangensis W. H. Tsao, 1964
- Schizothorax oconnori Lloyd, 1908
- Schizothorax oligolepis S. Y. Huang, 1985
- Schizothorax parvus W. H. Tsao, 1964
- Schizothorax pelzami Kessler, 1870 (Transcaspian Marinka)
- Schizothorax plagiostomus Heckel, 1838 (Asaila/Asela)
- Schizothorax prenanti (T. L. Tchang, 1930)
- Schizothorax progastus (McClelland, 1839) (Dinnawah Snowtrout)
- Schizothorax prophylax Pietschmann, 1933
- Schizothorax pseudoaksaiensis Herzenstein, 1889
- Schizothorax raraensis Terashima, 1984
- Schizothorax richardsonii (J. E. Gray, 1832) (Common Snowtrout, "Asaila"/"Asela")
- Schizothorax rotundimaxillaris Y. F. Wu & C. Z. Wu, 1992
- Schizothorax sinensis Herzenstein, 1889
- Schizothorax skarduensis Mirza & Awan, 1978
- Schizothorax waltoni Regan, 1905
- Schizothorax wangchiachii (P. W. Fang, 1936)
- Schizothorax yunnanensis Norman, 1923
- Schizothorax zarudnyi (Nikolskii, 1897)
- ^ Woodhouse (1910), FishBase [2009]
- ^ de Graaf et al. (2007)
References