Elachista
- For the genus of Chordariaceae brown algae, see Elachista (alga). As invalidly described by V. Kollar in 1832, Elachista refers to the Lyonetiidae moth genus Lyonetia.
Elachista is a genus of gelechioid moths. It is the type genus of the grass-miner moth family (Elachistidae). This family is sometimes (in particular in older sources) circumscribed very loosely, including for example the Agonoxenidae and Ethmiidae which seem to be quite distinct among the Gelechioidea, as well as other lineages which are widely held to be closer to Oecophora than to Elachista and are thus placed in the concealer moth family Oecophoridae here.[1]
These grass-miners are very small moths with the "feathery" hindwings characteristic of their family. They are essentially found worldwide, except in very cold places and on some oceanic islands; as usual for Gelechioidea, they are most common in the Palearctic however. They usually have at least one, sometimes as many as three light bands running from leading to trailing edge of their forewing uppersides. Some species, however, have upper forewings that are mostly white.[2]
Selected species
Not all species of this large genus have been discovered yet, let alone validly described and named. Several small genera, e.g. Biselachista and Cosmiotes, are here included in Elachista as junior synonyms (see below). They seem to be at least as closely related to E. bifasciella (the type species of the present genus) than other species commonly placed here, if not actually closer. In addition, Elachista contains several cryptic species complexes – such as the one around E. dispunctella and E. triseriatella – whose systematics and taxonomy are still not fully resolved.[3]
A number of Elachista species have been assigned to one of several groups, which may or may not be monophyletic, and whose names in the literature do not consistently follow the usual taxonomic practice (i.e. using as namesake the group-member which was described first). Some of these groups are placed in either of the two large subgenera Elachista (Aphelosetia) and Elachista (Elachista). Other proposed subgenera – the small Elachista (Dibrachia) and Elachista (Hemiprosopa) – are here included in the species of unclear relationships; like some small species-groups, they stand a chance of rendering the two larger subgenera paraphyletic if accepted as distinct, but seem too unlike them to warrant placement in either.[4]
Subgenus Elachista (Aphelosetia)
argentella group (sometimes in bedellella group)
bedellella/unifasciella group (including collitella group)
- Elachista adscitella Stainton, 1851
- Elachista antonia Kaila, 2007
- Elachista bedellella (Sircom, 1848)
- Elachista bisulcella (Duponchel, [1843])
- Elachista bruuni Traugott-Olsen, 1990
- Elachista camilla Kaila, 2007
- Elachista cahorsensis Traugott-Olsen, 1992
- Elachista chrysodesmella Zeller, 1850
- Elachista cingillella (Herrich-Schäffer, 1855)
- Elachista collitella (Duponchel, [1843])
- Elachista crocogastra Meyrick, 1908
- Elachista dispilella Zeller, 1839
- Elachista dispunctella (Duponchel, [1843])
- Elachista distigmatella Frey, 1859
- Elachista dorinda Kaila, 2007
- Elachista fasciola Parenti, 1983
- Elachista festucicolella Zeller, 1853
- Elachista gangabella Zeller, 1850
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- Elachista gregori Traugott-Olsen, 1988
- Elachista heinemanni Frey, 1866 (sometimes in E. subalbidella; tentatively placed here)
- Elachista istanella Nielsen & Traugott-Olsen, 1987
- Elachista laetella Rebel, 1930 (sometimes in E. subalbidella; tentatively placed here)
- Elachista levasi Sruoga, 1998
- Elachista littoricola Le Marchand, 1938
- Elachista lugdunensis Frey, 1859
- Elachista metella Kaila, 2002
- Elachista nedaella Traugott-Olsen, 1985
- Elachista nitidulella (Herrich-Schäffer, 1855)
- Elachista nolckeni Sulcs, 1992 (tentatively placed here)
- Elachista obliquella Stainton, 1854
- Elachista pollinariella Zeller, 1839
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- Elachista pollutissima Staudinger, 1880
- Elachista pullicomella
- Elachista purella Sruoga, 2000
- Elachista revinctella Zeller, 1850
- Elachista rudectella Stainton, 1851
- Elachista slivenica Kaila, 2007
- Elachista squamosella (Duponchel, [1843])
- Elachista subalbidella
- Elachista subocellea
- Elachista sutteri Kaila, 2002
- Elachista tinctella Sinev & Sruoga, 1995
- Elachista titanella Kaila & Jalava, 1994
- Elachista triseriatella Stainton, 1854
- Elachista unifasciella (Haworth, 1828)
- Elachista versicolora Kaila, 2007
- Elachista cf. dispunctella 'Volgograd Oblast'[5]
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Elachista (Aphelosetia) incertae sedis
- Elachista acutella Kaila, 2003
- Elachista agelensis Traugott-Olsen, 1996
- Elachista amparoae Traugott-Olsen, 1992
- Elachista amseli Rebel, 1833
- Elachista andorraensis Traugott-Olsen, 1988
- Elachista anitella Traugott-Olsen, 1985
- Elachista arduella Kaila, 2003
- Elachista atrisquamosa Staudinger, 1880
- Elachista baldizzonei Traugott-Olsen, 1996
- Elachista baldizzonella Traugott-Olsen, 1985
- Elachista bazaella Traugott-Olsen, 1992
- Elachista bazaensis Traugott-Olsen, 1990
- Elachista bengtssoni Traugott-Olsen, 1992
- Elachista berndtiella Traugott-Olsen, 1985
- Elachista bigorrensis Traugott-Olsen, 1990
- Elachista casascoensis Traugott-Olsen, 1992
- Elachista catalana Parenti, 1978
- Elachista catalunella Traugott-Olsen, 1992
- Elachista chamaea Kaila, 2003
- Elachista chionella Mann, 1861
- Elachista clintoni Traugott-Olsen, 1992
- Elachista constitella Frey, 1859
- Elachista contisella Chrétien, 1922
- Elachista cuencaensis Traugott-Olsen, 1992
- Elachista curonensis Traugott-Olsen, 1990
- Elachista dalmatiensis Traugott-Olsen, 1992
- Elachista deceptricula Staudinger, 1880
- Elachista disemiella Zeller, 1847
- Elachista drenovoi Parenti, 1981
- Elachista dumosa Parenti, 1981
- Elachista elsaella Traugott-Olsen, 1988
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- Elachista exigua Parenti, 1978
- Elachista flavescens Parenti, 1981
- Elachista fuscibasella Chrétien, 1915
- Elachista galacticella Eversmann, 1844
- Elachista gerdmaritella Traugott-Olsen, 1992
- Elachista gibbera Kaila, 2003
- Elachista gielisi Traugott-Olsen, 1992
- Elachista glaseri Traugott-Olsen, 1992
- Elachista gormella Nielsen & Traugott-Olsen, 1987
- Elachista graeca Parenti, 2002
- Elachista grandella Traugott-Olsen, 1992
- Elachista hallini Traugott-Olsen, 1992
- Elachista hedemanni Rebel, 1899
- Elachista heringi Rebel, 1899
- Elachista hispanica Traugott-Olsen, 1992
- Elachista imbi Traugott-Olsen, 1992
- Elachista intrigella Traugott-Olsen, 1992
- Elachista jaeckhi Traugott-Olsen, 1990
- Elachista karsholti Traugott-Olsen, 1992
- Elachista klimeschiella Parenti, 2002
- Elachista latipenella Sinev & Budashkin, 1991
- Elachista lerauti Traugott-Olsen, 1992
- Elachista liskai Kaila, 2011
- Elachista louisae Traugott-Olsen, 1992
- Elachista luqueti Traugott-Olsen, 1992
- Elachista maculata Parenti, 1978
- Elachista madridensis Traugott-Olsen, 1992
- Elachista mannella Traugott-Olsen, 1992
- Elachista manni Traugott-Olsen, 1990
- Elachista minusculella Traugott-Olsen, 1992
- Elachista modesta Parenti, 1978
- Elachista multipunctella Traugott-Olsen, 1992
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- Elachista neapolisella Traugott-Olsen, 1985
- Elachista nevadensis Parenti, 1978
- Elachista nielspederi Traugott-Olsen, 1992
- Elachista nuraghella Amsel, 1951
- Elachista occidentella Traugott-Olsen, 1992
- Elachista occulta Parenti, 1978
- Elachista ohridella Parenti, 2001
- Elachista olschwangi Kaila, 2003
- Elachista ozeini Parenti, 2004
- Elachista parvula Parenti, 1978
- Elachista passerini Traugott-Olsen, 1996
- Elachista pocopunctella Traugott-Olsen, 1992
- Elachista pollutella Duponchel, 1843
- Elachista povolnyi Traugott-Olsen, 1992
- Elachista punctella Traugott-Olsen, 1992
- Elachista rikkeae Traugott-Olsen, 1992
- Elachista skulei Traugott-Olsen, 1992
- Elachista spumella Caradja, 1920
- Elachista steueri Traugott-Olsen, 1990
- Elachista svenssoni Traugott-Olsen, 1988
- Elachista szocsi Parenti, 1978
- Elachista teruelensis Traugott-Olsen, 1990
- Elachista totanaensis Traugott-Olsen, 1992
- Elachista toveella Traugott-Olsen, 1985
- Elachista tribertiella Traugott-Olsen, 1985
- Elachista vanderwolfi Traugott-Olsen, 1992
- Elachista varensis Traugott-Olsen, 1992
- Elachista vegliae Parenti, 1978
- Elachista veletaella Traugott-Olsen, 1992
- Elachista vivesi Traugott-Olsen, 1992
- Elachista vulcana Kaila, 2011
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Subgenus Elachista (Atachia)
These are sorted in the presumed phylogenetic sequence.[6]
gerasmia group, gerasmia section
Incertae sedis
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Elachista cynopa complex
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Elachista melanthes complex
Elachista gerasmia complex
Elachista paragauda complex
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gerasmia group, catarata section
erebophthalma group
Subgenus Elachista (Elachista)
albifrontella/bifasciella group
- Elachista albifrontella (Hübner, [1817])
- Elachista alpinella
- Elachista apicipunctella
- Elachista atricomella
- Elachista baikalica Kaila, 1992
- Elachista bifasciella Treitschke, 1833
- Elachista compsa Traugott-Olsen, 1974
- Elachista deriventa Kaila & Mutanen, 2008
- Elachista diederichsiella E.Hering, 1889
- Elachista elegans Frey, 1859
- Elachista excelsicola
- Elachista fuscofrontella Sruoga, 1990
- Elachista habrella
- Elachista hiranoi Sugisima, 2005
- Elachista jaskai Kaila, 1998
- Elachista jupiter Sugisima, 2005
- Elachista kilmunella Stainton, 1849
- Elachista leifi Kaila & Kerppola, 1992
- Elachista luticomella
- Elachista micalis
- Elachista nobilella
- Elachista phalaridis Parenti, 1983
- Elachista platina
- Elachista poae Stainton, 1855
- Elachista scitula
- Elachista tanaella Aarvik & Berggren, 2004
- Elachista cf. bifasciella[6]
- Elachista cf. kilmunella 'Chelyabinsk Oblast'[5]
- Elachista cf. platina[6]
- Elachista sp. 'Chelyabinsk Oblast'[5]
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cerusella/monosemiella group
gleichenella group
orba group
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praelineata group
pulchella group
solena group
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"Cosmiotes"/freyerella group
Incertae sedis
- Elachista albipalpella Chambers, 1880
- Elachista albrechti Kaila, 1998
- Elachista amseli (Parenti, 1981)
- Elachista antipetra Meyrick, 1922
- Elachista baltica E.Hering, 1891 (sometimes in E. freyerella; tentatively placed here)
- Elachista consortella Stainton, 1851 (tentatively placed here)
- Elachista cristatella Chambers, 1876
- Elachista deficiens Meyrick, 1922
- Elachista epicaeria (Diakonoff, 1955)
- Elachista exactella (tentatively placed here)
- Elachista herbigrada Braun, 1925
- Elachista heteroplaca Meyrick, 1934
- Elachista illectella (Clemens, 1860) (type of Cosmiotes)
- Elachista illicitella (Clemens, 1860)
- Elachista infuscata Frey, 1882 (sometimes in E. exactella; tentatively placed here)
- Elachista inscia (Meyrick, 1913)
- Elachista ksarella Chrétien, 1908
- Elachista laquaeorum (Dugdale, 1971)
- Elachista lorigera (Meyrick, 1921) (type of Ptilodoxa)
- Elachista nipponicella Sugisima, 2006
- Elachista nymphaea Meyrick, 1911
- Elachista obtusella Sruoga, 2008
- Elachista ochroleuca Meyrick 1923
- Elachista pallens (Sruoga, 1990)
- Elachista planicara Kaila, 1998
- Elachista praematurella Clemens, 1860
- Elachista quadrata Meyrick, 1932
- Elachista scopulicola (Braun, 1948)
- Elachista spiculifera Meyrick, 1922
- Elachista stabilella Stainton, 1858 (tentatively placed here)
- Elachista stichospora Meyrick, 1932
- Elachista tuberella Sruoga, 2008
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Elachista freyerella complex[7]
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Elachista synethes complex[7]
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tetragonella group
- Elachista abiskoella Bengtsson, 1977
- Elachista agilis Braun, 1921
- Elachista albidella
- Elachista biatomella (Stainton, 1848)
- Elachista bilbaensis (Rebel, 1893)
- Elachista boursini Amsel, 1951
- Elachista christenseni Traugott-Olsen, 2000
- Elachista cinereopunctella (Haworth, 1828)
- Elachista cucullata Braun, 1921
- Elachista eleochariella
- Elachista ensifera
- Elachista falirakiensis Traugott-Olsen, 2000
- Elachista fulgens Parenti, 1983 (tentatively placed here)
- Elachista glaserella Traugott-Olsen, 2000
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- Elachista imatrella von Schantz, 1971
- Elachista jordanella Amsel, 1935
- Elachista juliensis Frey, 1870 = E. freyi Staudinger, 1871 (type of Biselachista)
- Elachista kebneella (Traugott-Olsen & Nielsen, 1977)
- Elachista kleini Amsel, 1935
- Elachista leucosticta Braun, 1948
- Elachista leucosyrma (Meyrick, 1932) (type of Platyphyllis)
- Elachista martinii Hofmann, [1898]
- Elachista nevadella Traugott-Olsen, 2000
- Elachista occidentalis Frey, 1882
- Elachista ornithopodella Frey, 1859
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- Elachista pigerella (Herrich-Schäffer, 1854) (type of Atachia)
- Elachista quadripunctella (Hübner, [1825])
- Elachista ribentella Kaila & Varalda, 2004
- Elachista saarelai
- Elachista salinaris Braun, 1925
- Elachista scirpi Stainton, 1887
- Elachista serricornis
- Elachista tetragonella (Herrich-Schäffer, 1855)
- Elachista trapeziella Stainton, 1849
- Elachista utonella Frey, 1856
- Elachista zabella Chrétien, 1908
- Elachista zonulae (Sruoga, 1992)
- Elachista cf. biatomella 'Ajat River'[5]
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Elachista (Elachista) incertae sedis
- Elachista adelpha Kaila & Jalava, 1994
- Elachista albicapilla Höfner, 1918
- Elachista anserinelloides Nel, 2003
- Elachista argentifasciella Höfner, 1898
- Elachista arnoldi (Koster, 1993)
- Elachista brachypterella (Klimesch, 1990)
- Elachista canariella Nielsen & Traugott-Olsen, 1987
- Elachista contaminatella Zeller, 1847
- Elachista devexella Kaila, 2003
- Elachista differens Parenti, 1978
- Elachista dimicatella Rebel, 1903
- Elachista encumeadae Kaila & Karsholt, 2002
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- Elachista geminatella (Herrich-Schäffer, 1855)
- Elachista griseella (Duponchel, 1843)
- Elachista gruenewaldi Parenti, 2002
- Elachista ibericella Traugott-Olsen, 1995
- Elachista igaloensis (Amsel, 1951)
- Elachista irenae Buszko, 1989
- Elachista kosteri Traugott-Olsen, 1995
- Elachista maculosella Chrétien, 1896
- Elachista megagnathos Sruoga, 1990
- Elachista minuta (Parenti, 2003)
- Elachista morandinii Huemer & Kaila, 2003
- Elachista sicula Parenti, 1978
- Elachista stenopterella Rebel, 1932
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Other groups
Some Elachista are divided into groups whose relationships to the two large subgenera requires further study, including:[8]
"Dibrachia" group
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"Irenicodes" group
- Elachista eurychora (Meyrick, 1919) (type of Irenicodes)
- Elachista galatheae (Viette, 1954) (type of Euproteodes)
- Elachista holdgatei (Bradley, 1965)
- Elachista hookeri (Dugdale, 1971)
- Elachista pumila (Dugdale, 1971)
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Incertae sedis
Additional Elachista species are of even more uncertain relationships than usual for this genus:[9]
- Elachista albella (Chambers, 1877) (type of Hemiprosopa)
- Elachista altaica Sinev, 1998
- Elachista baikalica Kaila, 1992
- Elachista bassii Parenti, 2006
- Elachista beltira Kaila, 2000
- Elachista beriga Kaila, 2000
- Elachista bimaculata Parenti, 1981
- Elachista brevis Sruoga & J. de Prins, 2009
- Elachista chelonites Meyrick, 1909 (type of Cleroptila)
- Elachista deresyensis Traugott-Olsen,1988
- Elachista donia Kaila, 2000
- Elachista gibbera Kaila, 2003
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- Elachista kakamegensis Sruoga & J. de Prins, 2009
- Elachista kherana Kaila, 2000
- Elachista longispina Sruoga & J. de Prins, 2009
- Elachista maculosa Parenti, 1981
- Elachista margareteae (Traugott-Olsen, 1994)
- Elachista mus Parenti, 1981
- Elachista planca Sruoga & J. de Prins, 2009
- Elachista puplesisi Sruoga, 2000
- Elachista semnani Parenti, 1981
- Elachista stelviella Amsel, 1932
- Elachista vasrana Kaila, 2000
- Elachista veruta Kaila, 2008
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Synonyms
Invalid scientific names (junior synonyms and others) of Elachista in the circumscription as presented here are:[10]
- Aphelosetia Stephens, 1834
- Aphigalia Dyar, [1903]
- Atachia Wocke in Heinemann, [1876]
- Biselachista Traugott-Olsen & Schmidt Nielsen, 1977
- Cleroptila Meyrick, 1909
- Cosmiotes Clemens, 1860
- Cycnodia Herrich-Schäffer, 1853
- Dibrachia Sinev & Sruoga, 1992
- Dicasteris Meyrick, 1906
- Elaschista (lapsus)
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- Euproteodes Viette, 1954
- Hecista Wallengren, 1881
- Hemiprosopa A.F.Braun, 1948
- Irenicodes Meyrick, 1919
- Neaera Chambers, 1880 (non Robineau-Desvoidy, 1830: preoccupied)
- Paraperittia Rebel, 1916
- Phigalia Chambers, 1875 (non Duponchel, 1829: preoccupied)
- Platyphyllis Meyrick, 1932
- Ptilodoxa Meyrick, 1921
- Svenssonia Parenti, 2009
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- ^ AEBR (2008), FE (2009)
- ^ AEBR (2008), FE (2009), and see references in Savela (2008)
- ^ Kusdas & Reichl (1973): 194, Pitkin & Jenkins (2004), AEBR (2008), FE (2009), and see references in Savela (2008)
- ^ FE (2009), Wikispecies (2009-SEP-27), and see references in Savela (2008)
- ^ a b c d See references in Savela (2008)
- ^ a b c d e f g h i j k Kaila (2011)
- ^ a b Sorted in the presumed phylogenetic sequence (Kaila, 2011).
- ^ ABRS (2008), FE (2009), Wikispecies (2009-SEP-27)
- ^ Wikispecies (2009-SEP-27), and see references in Savela (2008)
- ^ AEBR (2008), FE (2009), Wikispecies (2009-SEP-27), and see references in Savela (2008)
References
Data related to Elachista at Wikispecies
- Australian Biological Resources Study (ABRS) (2008): Australian Faunal Directory – Elachista. Version of 2008-OCT-09. Retrieved 2010-MAY-01.
- Fauna Europaea (FE) (2009): Elachista. Version 2.1, 2009-DEC-22. Retrieved 2010-MAY-01.
- Kaila, Lauri (2011): Elachistine Moths of Australia (Lepidoptera: Gelechioidea, Elachistidae) (Monographs on Australian Lepidoptera Series 11). CSIRO Publishing. ISBN 978-0643103054. Preview at Google Books
- Kusdas, Karl & Reichl, Ernst Rudolf (eds.) (1973): Schmetterlinge Oberösterreichs (Vol. 6: Microlepidoptera) [in German]. Landwirtschaftskammer für Oberösterreich.
- Pitkin, Brian & Jenkins, Paul (2004): Butterflies and Moths of the World, Generic Names and their Type-species – Elachista. Version of 2004-NOV-05. Retrieved 2010-MAY-01.
- Savela, Markku (2008): Markku Savela's Lepidoptera and some other life forms – Elachista. Version of 2008-JUL-19. Retrieved 2010-MAY-01.