Arthrodira

Arthrodira
Temporal range: Late Silurian? to Late Devonian
Dunkleosteus terrelli
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Class: Placodermi
Order: Arthrodira
Woodward, 1898
Suborders and infraorders

basal

  • Actinolepina

Phlyctaenioidei

  • Phlyctaeniina

Brachythoraci

and see text

Arthrodira is an order of extinct armoured jawed fishes of the Placodermi class that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches.

Greek for "jointed neck", the arthrodires had movable joint between armor surrounding the head and body. The mouth is interesting because as the lower jaw moved down the head shield moved allowing for a large opening. Lacking distinct teeth, like all placoderms, they used the sharpened edges of a bony plate as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus Arctolepis, were well-armoured fishes with flattened bodies. The largest member of this group, Dunkleosteus, was a true superpredator of the latest Devonian period, reaching 3 to as much as 9 metres in length. In contrast, the long-nosed Rolfosteus measured just 15 cm.

There is a common misconception that the arthrodires (along with all other placoderms) were sluggish bottom-dwellers that were out-competed by more advanced fish. Leading to this misconception is that the arthrodire body plan remained relatively conserved (that is, the majority of arthrodires were bullet or torpedo-shaped) during the Devonian period, save for increasing in size. However during their reign the arthrodires were one of the most diverse and numerically successful, if not the most successful, vertebrate orders of the Devonian, occupying a vast spectrum of roles from apex predator to detritus-nibbling bottom dweller. Despite their success, the arthrodires were one of many groups eliminated by the environmental catastrophes of the Late Devonian extinction, allowing other fish such as sharks to diversify into the vacated ecological niches during the Carboniferous period.

Contents

Systematics[1]

ORDER ARTHRODIRA

Genera incertae sedis

Arthrodira of unresolved relationships

Timeline of genera


Footnotes

  1. ^ For sources, see Haaramo (2005)

See Also

List of placoderms

References

External links