Arctotherium

Arctotherium
Temporal range: Late Pliocene–Middle Pleistocene
Life restoration of Arctotherium bonariense
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Superfamily: Arctoidea
Family: Ursidae
Subfamily: Tremarctinae
Tribe: Tremarctini
Genus: Arctotherium
Burmeister, 1879
Species
  • A. angustidens
  • A. bonariense
  • A. brasilense
  • A. latidens
  • A. tarijense
  • A. vetustum
  • A. wingei

Arctotherium is an extinct genus of South American short-faced bears within Ursidae of the late Pliocene through the end of the Pleistocene.[1] Their ancestors migrated from North America to South America during the Great American Interchange, following the formation of the Isthmus of Panama. They were endemic to South America, living from ~2.0–0.01 Ma, existing for approximately 1.99 million years. Their closest relatives were the North American short-faced bears of genus Arctodus (A. pristinus and A. simus). The closest living relative would be the spectacled bear (Tremarctos ornatus).[2]

Arctotherium was named by Hermann Burmeister in 1879. It was assigned to Tremarctinae by Krause et al. 2008.[2] A humerus of A. angustidens from Buenos Aires indicates that the big males of this species would have weighed between 1588-1749 kg, standing at least 11 feet (3.4 meters) tall, making it the largest bear and the largest carnivorous land mammal yet known.[3][4] In contrast to their North American cousins, South American short-faced bears showed a trend of declining size and carnivory over time. This has been attributed to increased competition from other, later-arriving or evolving carnivorans, like jaguars or lions, following the early dispersal of short-faced bears to South America.[3][4][5] (The North American carnivorans that invaded South America, including short-faced bears and Smilodon, quickly dominated the predatory niches formerly occupied by South America's native metatherian sparassodont and avian phorusrhacid carnivores.)

References

  1. ^ Soibelzon, L. H.; Tonni, E. P.; Bond, M. (October 2005). "The fossil record of South American short-faced bears (Ursidae, Tremarctinae)". Journal of South American Earth Sciences 20 (1–2): 105–113. doi:10.1016/j.jsames.2005.07.005. 
  2. ^ a b Krause, J.; Unger, T.; Noçon, A.; Malaspinas, A.; Kolokotronis, S.; Stiller, M.; Soibelzon, L.; Spriggs, H.; Dear, P. H.; Briggs, A. W.; Bray, S. C. E.; O'Brien, S. J.; Rabeder, G.; Matheus, P.; Cooper, A.; Slatkin, M.; Pääbo, S.; Hofreiter, M. (2008-07-28). "Mitochondrial genomes reveal an explosive radiation of extinct and extant bears near the Miocene-Pliocene boundary". BMC Evolutionary Biology 8: 220. doi:10.1186/1471-2148-8-220. PMC 2518930. PMID 18662376. http://www.pubmedcentral.nih.gov/articlerender.fcgi?tool=pmcentrez&artid=2518930. 
  3. ^ a b Dell'Amore, C. (2011): Biggest Bear Ever Found, National Geographic News, Published February 3, 2011
  4. ^ a b Soibelzon, L. H.; Schubert, B. W. (2011-01). "The Largest Known Bear, Arctotherium angustidens, from the Early Pleistocene Pampean Region of Argentina: With a Discussion of Size and Diet Trends in Bears". Journal of Paleontology (Paleontological Society) 85 (1): 69–75. doi:10.1666/10-037.1. http://jpaleontol.geoscienceworld.org/cgi/content/abstract/85/1/69. Retrieved 2011-06-01. 
  5. ^ Hodge, A.-M. (2011-03-31). "Updated Range of Immensity for Arctotherium: New Record for Largest Known Bear". nature.com blogs. {Nature Publishing Group]]. http://blogs.nature.com/amch/2011/03/31/south-america-is-currently-home. Retrieved 2011-06-01.