Toothed whale
The toothed whales (systematic name Odontoceti) form a suborder of the cetaceans, including sperm whales, beaked whales, orca[1], dolphins, and others. As the name suggests, the suborder is characterized by having teeth, rather than baleen as do animals in the other suborder of cetaceans, Mysticeti. However, fossils indicate that early Baleen whales had teeth as well before evolving baleen, so defining the Odontoceti on teeth alone is problematic, and paleontologists have instead identified other features uniting fossil and modern odontocetes that are not shared by mysticetes. Toothed whales are active hunters, feeding on fish, squid, and in some cases marine mammals.
Anatomy
Toothed whales have a single blowhole on the top of the head (while the baleen whales possess two of them). The nostrils are not fused; one of them has become dominant over the other.
Toothed whales, as all whales, develop as an embryo in the womb, they are then born as calves, grow into juveniles and finally become adults.
As an adaptation for their echolocation, toothed whale skulls have become asymmetric. Their brains are relatively big, although real growth didn't occur before their echolocation started to evolve. Toothed whales' brains have a poor connection between the two hemispheres and an organ called a melon on their heads is used as a lens to focus sound waves. Vocal cords are not present; their sounds are produced in the blowhole system instead. Toothed whales have lost their sense of smell, as well as their saliva glands.
Except for the Sperm Whale, most toothed whales are smaller than the baleen whales. The teeth differ considerably between the species. They may be numerous, with some dolphins bearing over 100 teeth in their jaws. At the other extreme are the Narwhal with its single long tusk and the almost toothless beaked whales with bizarre teeth only in males. Not all species are believed to use their teeth for feeding. For instance, the Sperm Whale likely uses its teeth for aggression and showmanship.
Behaviour
Vocalizations
Vocalizations are of great importance to toothed whales. While many species also maintain a broad variety of calls to communicate, all species investigated so far use short click sounds for purposes of echolocation. Sperm whales use low frequencies (a few to perhaps 50 kHz), while other employ more narrow band high frequency sounds (porpoises, Cephalorhynchus species like Hector's dolphin). Most dolphin species use very broad band clicks.
Movement
Most toothed whales swim rapidly. The smaller species occasionally ride waves, such as the bow waves of ships. Dolphins can be frequently encountered this way. They are also famous for their acrobatic breaching from the water, e.g. the Spinner Dolphin.
Human impact
The Sperm Whale has been hunted commercially for a long time (see whaling). While small whales like the Pilot Whale today are still being pursued, the main threat for most species is accidental capture in fishing nets.
Keeping small whales (mostly Bottlenose Dolphins, Orcas, or Belugas) in captivity is a great attraction for ocean parks and zoos. However, it is controversial because of the marine mammals' need for large spaces.
Taxonomy
- Genus Cephalorhynchus
- Commerson's Dolphin, Cephalorhynchus commersonii
- Chilean Dolphin, Cephalorhynchus eutropia
- Heaviside's Dolphin, Cephalorhynchus heavisidii
- Hector's Dolphin, Cephalorhynchus hectori
- Genus Steno
- Rough-toothed Dolphin, Steno bredanensis
- Genus Sousa
- Atlantic Humpback Dolphin, Sousa teuszi
- Indian Humpback Dolphin, Sousa plumbea
- Chinese White Dolphin, Sousa chinensis
- Genus Sotalia
- Tucuxi, Sotalia fluviatilis
- Costero, Sotalia guianensis
- Genus Stenella
- Pantropical Spotted Dolphin, Stenella attenuata
- Atlantic Spotted Dolphin, Stenella frontalis
- Spinner Dolphin, Stenella longirostris
- Clymene Dolphin, Stenella clymene
- Striped Dolphin, Stenella coeruleoalba
- Genus Delphinus
- Short-beaked Common Dolphin, Delphinus delphis
- Long-beaked Common Dolphin, Delphinus capensis
- (Arabian Common Dolphin, Delphinus tropicalis)
- Genus Lagenodelphis
- Fraser's Dolphin, Lagenodelphis hosei
- Genus Lagenorhynchus
- White-beaked Dolphin, Lagenorhynchus albirostris
- Atlantic White-sided Dolphin, Lagenorhynchus acutus
- Pacific White-sided Dolphin, Lagenorhynchus obliquidens
- Dusky Dolphin, Lagenorhynchus obscurus
- Black-chinned Dolphin, Lagenorhynchus australis
- Hourglass Dolphin, Lagenorhynchus cruciger
- Genus Lissodelphis
- Northern Right Whale Dolphin, Lissodelphis borealis
- Southern Right Whale Dolphin, Lissodelphis peronii
- Genus Grampus
- Risso's Dolphin, Grampus griseus
- Genus Peponocephala
- Melon-headed Whale, Peponocephala electra
- Genus Feresa
- Pygmy Killer Whale, Feresa attenuata
- Genus Pseudorca
- False Killer Whale, Pseudorca crassidens
- Genus Globicephala
- Long-finned Pilot Whale, Globicephala melas
- Short-finned Pilot Whale, Globicephala macrorhynchus
- Genus Orcaella
- Irrawaddy Dolphin, Orcaella brevirostris
- Australian Snubfin Dolphin, Orcaella heinsohni
- Genus Neophocaena
- Finless Porpoise, Neophocaena phocaenoides
- Genus Phocoena
- Harbour Porpoise, Phocoena phocaena
- Vaquita, Phocoena sinus
- Spectacled Porpoise, Phocoena dioptrica
- Burmeister's Porpoise, Phocoena spinipinnis
- Genus Phocoenoides
- Dall's Porpoise, Phocoenoides dalli
- Genus Kogia
- Dwarf Sperm Whale, Kogia sima
- Pygmy Sperm Whale, Kogia breviceps
- Family Ziphidae: beaked whales
- Genus Ziphius
- Cuvier's Beaked Whale, Ziphius cavirostris
- Genus Berardius, giant beaked whales
- Arnoux's Beaked Whale, Berardius arnuxii
- Baird's Beaked Whale (North Pacific Bottlenose Whale), Berardius bairdii
- Genus Tasmacetus
- Tasman Beaked Whale (Shepherd's Beaked Whale), Tasmacetus shepherdi
- Sub-family Hyperoodontidae
- Genus Indopacetus
- Indo-Pacific Beaked Whale (Longman's Beaked Whale), Indopacetus pacificus
- Genus Hyperoodon
- Northern Bottlenose Whale, Hyperoodon ampullatus
- Southern Bottlenose Whale, Hyperoodon planifrons
- Genus Mesoplodon, mesoplodont whales
- Hector's Beaked Whale, Mesoplodon hectori
- True's Beaked Whale, Mesoplodon mirus
- Gervais' Beaked Whale, Mesoplodon europaeus
- Sowerby's Beaked Whale, Mesoplodon bidens
- Gray's Beaked Whale, Mesoplodon grayi
- Pygmy Beaked Whale, Mesoplodon peruvianus
- Andrew's Beaked Whale, Mesoplodon bowdoini
- Bahamonde's Beaked Whale, Mesoplodon bahamondi
- Hubbs' Beaked Whale, Mesoplodon carlhubbsi
- Ginko-toothed Beaked Whale, Mesoplodon ginkgodens
- Stejneger's Beaked Whale, Mesoplodon stejnegeri
- Layard's Beaked Whale, Mesoplodon layardii
- Blainville's Beaked Whale, Mesoplodon densirostris
- Perrin's Beaked Whale, Mesoplodon perrini
- Genus Inia
- Amazon River Dolphin, Inia geoffrensis
- Genus Platanista
- Ganges and Indus River Dolphin, Platanista gangetica
- Genus Pontoporia
- La Plata Dolphin, Pontoporia blainvillei
References
Extant Cetacea species |
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Kingdom Animalia · Phylum Chordata · Class Mammalia · Infraclass Eutheria · Superorder Laurasiatheria · (unranked) Cetartiodactyla · (unranked) Whippomorpha |
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Suborder Odontoceti (Toothed whales) (cont. below) |
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Delphinidae
(Oceanic dolphins) |
Peponocephala
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Melon-headed Whale (P. electra)
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Feresa
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Pygmy Killer Whale (F. attenuata)
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Pseudorca
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False Killer Whale (P. crassidens)
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Long-finned Pilot Whale (G. melas) · Short-finned Pilot Whale (G. macrorhynchus)
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Long-beaked Common Dolphin (D. capensis) · Short-beaked Common Dolphin (D. delphis)
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Lissodelphis
(Right whale dolphins)
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Northern Right Whale Dolphin (L. borealis) · Southern Right Whale Dolphin (L. peronii)
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Sotalia
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Tucuxi (S. fluviatilis) · Costero (Sotalia guianensis)
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Sousa
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Pacific Humpback Dolphin (S. chinensis) · Indian Humpback Dolphin (S. plumbea) · Atlantic Humpback Dolphin (S. teuszii)
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Stenella
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Atlantic Spotted Dolphin (S. frontalis) · Clymene Dolphin (S. clymene) · Pantropical Spotted Dolphin (S. attenuata) · Spinner Dolphin (S. longirostris) · Striped Dolphin (S. coeruleoalba)
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Steno
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Rough-toothed Dolphin (S. bredanensis)
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Cephalorhynchus
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Chilean Dolphin (C. eutropia) · Commerson's Dolphin (C. commersonii) · Heaviside's Dolphin (C. heavisidii) · Hector's Dolphin (C. hectori)
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Grampus
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Risso's Dolphin (G. griseus)
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Lagenodelphis
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Fraser's Dolphin (L. hosei)
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Lagenorhynchus
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Atlantic White-sided Dolphin (L. acutus) · Dusky Dolphin (L. obscurus) · Hourglass Dolphin (L. cruciger) · Pacific White-sided Dolphin (L. obliquidens) · Peale's Dolphin (L. australis) · White-beaked Dolphin (L. albirostris)
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Orcaella
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Irrawaddy Dolphin (O. brevirostris) · Australian Snubfin Dolphin (O. heinsohni)
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Suborder Odontoceti (Toothed whales) (cont. above) |
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Monodontidae |
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Phocoenidae
(Porpoises) |
Neophocaena
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Finless Porpoise (N. phocaeniodes)
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Harbor Porpoise (P. phocoena) · Vaquita (P. sinus) · Spectacled Porpoise (P. dioptrica) · Burmeister's Porpoise (P. spinipinnis)
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Phocoenoides
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Dall's Porpoise (P. dalli)
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Physeteridae |
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Kogiidae |
Kogia
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Pygmy sperm whale (K. breviceps) · Dwarf sperm whale (K. simus)
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Ziphidae
(Beaked whales) |
Berardius
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Arnoux's Beaked Whale (B. arnuxii) · Baird's Beaked Whale (B. bairdii)
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Hyperoodon
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Northern Bottlenose Whale (H. ampullatus) · Southern Bottlenose Whale (H. planifrons)
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Indopacetus
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Longman's Beaked Whale (I. pacificus)
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Mesoplodon
(Mesoplodont
whales)
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Sowerby's Beaked Whale (M. bidens) · Andrews' Beaked Whale (M. bowdoini) · Hubbs' Beaked Whale (M. carlhubbsi) · Blainville's Beaked Whale (M. densirostris) · Gervais' Beaked Whale (M. europaeus) · Ginkgo-toothed Beaked Whale (M. ginkgodens) · Gray's Beaked Whale (M. grayi) · Hector's Beaked Whale (M. hectori) · Layard's Beaked Whale (M. layardii) · True's Beaked Whale (M. mirus) · Pygmy Beaked Whale (M. peruvianus) · Perrin's Beaked Whale (M. perrini) · Stejneger's Beaked Whale (M. stejnegeri) · Spade-toothed Whale (M. traversii)
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Tasmacetus
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Shepherd's Beaked Whale (T. sheperdi)
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Ziphius
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Cuvier's Beaked Whale (Z. cavirostris)
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Iniidae |
Inia
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Amazon River Dolphin (I. geoffrensis)
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Lipotidae |
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Platanistidae |
Platanista
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Ganges and Indus River Dolphin (P. gangetica)
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Pontoporiidae |
Pontoporia
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La Plata Dolphin (P. blainvillei)
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