Tyrant flycatcher
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Tyrant flycatchers | ||||||||||||
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Scissor-tailed Flycatcher
Tyrannus forficatus |
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Scientific classification | ||||||||||||
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Distribution of tyrant flycatchers
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Some 100, see text |
The tyrant flycatchers (Tyrannidae) are a family of passerine birds which occur throughout North and South America, but are mainly tropical in distribution. They are now considered the largest family of birds on earth, with around 400 species. As could be expected from a family this large, the members vary greatly both in shape, patterns and colours. Some Tyrant flycatchers superficially resemble the Old World flycatchers. They are members of suborder Tyranni (suboscines) that do not have the sophisticated vocal capabilities of the songbirds.
Most, but not all, are rather plain, and many have erectile crests. As the name implies, most are insectivorous, but some will eat fruit or small vertebrates (e.g. small frogs). The smallest family members are the closely related Short-tailed Pygmy-Tyrant and Black-capped Pygmy-Tyrant. With a total length of a mere 6.5-6.8 cm (2.5-2.7 in) and a weight of 4-5 grams, they are the smallest passerines on earth. The largest tyrant flycatcher is the Great Shrike-Tyrant at 29 cm (11.5 in) and 88 grams (3.1 oz). A few species such as the Streamer-tailed Tyrant and Fork-tailed Flycatcher have a larger total length, but this is mainly due to their very long tails.
A number of species previously included in this family are now placed in the family Tityridae (see Systematics).
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[edit] Habitat and distribution
Species richness of Tyrannidae, when compared to habitat, is highly variable. The habitats of tropical lowland evergreen forest and montane evergreen forest have the highest single site species diversity while many habitats including rivers, palm forest, white sand forest, tropical deciduous forest edge, southern temperate forest, southern temperate forest edge, semihumid/humid montane scrub, and northern temperate grassland have the lowest single species diversity. The variation between the highest and the lowest is extreme; ninety species can be found in the tropical lowland evergreen forests while only one species can be found at the habitats listed above. This may be due in part to the fewer niches found in certain areas and therefore fewer places for the species to occupy.
Tyrannidae specialization among habitat is very strong in tropical lowland evergreen forests and montane evergreen forests. These habitat types therefore display the greatest specialization. The counts differ by three species (tropical lowland evergreen forests have 49 endemic species and montane evergreen forests have 46 endemic species). It can be assumed that they both have similar levels of specialization.
Regionally, the South Atlantic Coast has a significanltly higher species richness with the Manabí-Tumbes region following closely behind.
[edit] Protected status
The Northern Beardless Tyrannulet (Camptostoma imberbe) and the Rose-throated Becard (Pachyramphus aglaiae) are protected under the Migratory Bird Treaty Act of 1918.[1]. Both these species are common south of the US border. The situation for a number of other species from South and Central America is far more problematic. In 2007, BirdLife International (and consequently IUCN) considered two species, the Minas Gerais Tyrannulet and Kaempfer's Tody-Tyrant critically endangered. Both are endemic to Brazil. Additional, 7 species were considered endangered and 18 species vulnerable.[1]
[edit] Systematics
There are about 400 species in over 100 genera. A full list, sortable by common and binomial names is at list of tyrant flycatchers. Species in the genera Tityra, Pachyramphus, Laniocera and Xenopsaris have been placed in this family, but evidence strongly suggest they belong in Tityridae[2], where now placed by SACC.
- Genus Ornithion
- White-lored Tyrannulet, Ornithion inerme
- Yellow-bellied Tyrannulet, Ornithion semiflavum
- Brown-capped Tyrannulet, Ornithion brunneicapillum
- Genus Camptostoma
- Northern Beardless Tyrannulet, Camptostoma imberbe
- Southern Beardless Tyrannulet, Camptostoma obsoletum
- Genus Phaeomyias
- Mouse-colored Tyrannulet, Phaeomyias murina
- Genus Nesotriccus
- Cocos Island Flycatcher, Nesotriccus ridgwayi
- Genus Capsiempis - Yellow Tyrannulet
- Genus Tyrannulus
- Yellow-crowned Tyrannulet, Tyrannulus elatus
- Genus Myiopagis
- Forest Elaenia, Myiopagis gaimardii
- Foothill Elaenia, Myiopagis olallai
- Grey Elaenia, Myiopagis caniceps
- Pacific Elaenia, Myiopagis subplacens
- Yellow-crowned Elaenia, Myiopagis flavivertex
- Jamaican Elaenia, Myiopagis cotta
- Greenish Elaenia, Myiopagis viridicata
- Genus Pseudelaenia
- Gray-and-white Tyrannulet, Pseudelaenia leucospodia
- Genus Elaenia (18 species)
- Genus Serpophaga
- Torrent Tyrannulet, Serpophaga cinerea
- Sooty Tyrannulet, Serpophaga nigricans
- River Tyrannulet, Serpophaga hypoleuca
- White-crested Tyrannulet, Serpophaga subcristata
- White-bellied Tyrannulet, Serpophaga munda
- Genus Mionectes (5 species)
- Genus Leptopogon (4 species)
- Genus Pseudotriccus
- Bronze-olive Pygmy-tyrant, Pseudotriccus pelzelni
- Hazel-fronted Pygmy-tyrant, Pseudotriccus simplex
- Rufous-headed Pygmy-tyrant, Pseudotriccus ruficeps
- Genus Phylloscartes
- Marble-faced Bristle-tyrant, Phylloscartes ophthalmicus
- Venezuelan Bristle-tyrant, Phylloscartes venezuelanus
- Antioquia Bristle-tyrant, Phylloscartes lanyoni
- Spectacled Bristle-tyrant, Phylloscartes orbitalis
- Variegated Bristle-tyrant, Phylloscartes poecilotis
- Southern Bristle-tyrant, Phylloscartes eximius
- Black-fronted Tyrannulet, Phylloscartes nigrifrons
- Chapman's Bristle-tyrant, Phylloscartes chapmani
- Ecuadorian Tyrannulet, Phylloscartes gualaquizae
- Rufous-lored Tyrannulet, Phylloscartes flaviventris
- Cinnamon-faced Tyrannulet, Phylloscartes parkeri
- Minas Gerais Tyrannulet, Phylloscartes roquettei
- Sao Paulo Tyrannulet, Phylloscartes paulistus
- Oustalet's Tyrannulet, Phylloscartes oustaleti
- Restinga Tyrannulet, Phylloscartes kronei
- Serra do Mar Tyrannulet, Phylloscartes difficilis
- Alagoas Tyrannulet, Phylloscartes ceciliae
- Mottle-cheeked Tyrannulet, Phylloscartes ventralis
- Bahia Tyrannulet, Phylloscartes beckeri
- Yellow-green Tyrannulet, Phylloscartes flavovirens
- Olive-green Tyrannulet, Phylloscartes virescens
- Rufous-browed Tyrannulet, Phylloscartes superciliaris
- Bay-ringed Tyrannulet, Phylloscartes sylviolus
- Genus Phyllomyias
- Planalto Tyrannulet, Phyllomyias fasciatus
- White-fronted Tyrannulet, Phyllomyias zeledoni
- Rough-legged Tyrannulet, Phyllomyias burmeisteri
- Greenish Tyrannulet, Phyllomyias virescens
- Reiser's Tyrannulet, Phyllomyias reiseri
- Sclater's Tyrannulet, Phyllomyias sclateri
- Gray-capped Tyrannulet, Phyllomyias griseocapilla
- Sooty-headed Tyrannulet, Phyllomyias griseiceps
- Plumbeous-crowned Tyrannulet, Phyllomyias plumbeiceps
- Black-capped Tyrannulet, Phyllomyias nigrocapillus
- Ashy-headed Tyrannulet, Phyllomyias cinereiceps
- Tawny-rumped Tyrannulet, Phyllomyias uropygialis
- Genus Zimmerius
- Mistletoe Tyrannulet, Zimmerius vilissimus
- Venezuelan Tyrannulet, Zimmerius improbus
- Bolivian Tyrannulet, Zimmerius bolivianus
- Red-billed Tyrannulet, Zimmerius cinereicapillus
- Slender-footed Tyrannulet, Zimmerius gracilipes
- Peruvian Tyrannulet, Zimmerius viridiflavus
- Golden-faced Tyrannulet, Zimmerius chrysops
- Genus Sublegatus
- Amazonian Scrub-flycatcher, Sublegatus obscurior
- Northern Scrub-flycatcher, Sublegatus arenarum
- Southern Scrub-flycatcher, Sublegatus modestus
- Genus Suiriri (2 species)
- Genus Mecocerculus (6 species)
- Genus Inezia (4 species)
- Genus Stigmatura
- Lesser Wagtail-tyrant, Stigmatura napensis
- Greater Wagtail-tyrant, Stigmatura budytoides
- Genus Uromyias - sometimes included in Anairetes
- Agile Tit-tyrant, Uromyias agilis
- Unstreaked Tit-tyrant, Uromyias agraphia
- Genus Anairetes - typical tit-tyrants (6 species)
- Genus Tachuris - Many-colored Rush-tyrant
- Genus Culicivora - Sharp-tailed Grass-Tyrant
- Genus Polystictus
- Bearded Tachuri, Polystictus pectoralis
- Gray-backed Tachuri, Polystictus superciliaris
- Genus Pseudocolopteryx
- Crested Doradito, Pseudocolopteryx sclateri
- Dinelli's Doradito, Pseudocolopteryx dinellianus
- Subtropical Doradito, Pseudocolopteryx acutipennis
- Warbling Doradito, Pseudocolopteryx flaviventris
- Genus Euscarthmus (2 species)
- Genus Myiornis
- White-bellied Pygmy-tyrant, Myiornis albiventris
- Eared Pygmy-tyrant, Myiornis auricularis
- Black-capped Pygmy-tyrant, Myiornis atricapillus
- Short-tailed Pygmy-tyrant, Myiornis ecaudatus
- Genus Lophotriccus (5 species)
- Genus Oncostoma
- Northern Bentbill, Oncostoma cinereigulare
- Southern Bentbill, Oncostoma olivaceum
- Genus Poecilotriccus (12 species)
- Genus Taeniotriccus - Black-chested Tyrant
- Genus Hemitriccus - typical tody-tyrants (20 species)
- Genus Todirostrum - typical tody-flycatchers (7 species, others now in Poecilotriccus)
- Genus Corythopis - antpipits (2 species)
- Genus Cnipodectes (2 species)
- Genus Ramphotrigon (3 species)
- Genus Rhynchocyclus (4 species)
- Genus Tolmomyias
- Yellow-olive Flycatcher, Tolmomyias sulphurescens
- Yellow-margined Flycatcher, Tolmomyias assimilis
- Gray-crowned Flycatcher, Tolmomyias poliocephalus
- Orange-eyed Flycatcher, Tolmomyias traylori
- Yellow-breasted Flycatcher, Tolmomyias flaviventris
- Genus Platyrinchus - spadebills (7 species)
- Genus Onychorhynchus - Royal Flycatchers (1-4 species, depending on taxonomy)
- Genus Myiotriccus
- Ornate Flycatcher, Myiotriccus ornatus
- Genus Myiophobus
- Flavescent Flycatcher, Myiophobus flavicans
- Orange-crested Flycatcher, Myiophobus phoenicomitra
- Roraiman Flycatcher, Myiophobus roraimae
- Unadorned Flycatcher, Myiophobus inornatus
- Handsome Flycatcher, Myiophobus pulcher
- Orange-banded Flycatcher, Myiophobus lintoni
- Ochraceous-breasted Flycatcher, Myiophobus ochraceiventris
- Bran-colored Flycatcher, Myiophobus fasciatus
- Olive-chested Flycatcher, Myiophobus cryptoxanthus
- Genus Terenotriccus - Ruddy-tailed Flycatcher
- Genus Myiobius
- Tawny-breasted Flycatcher, Myiobius villosus
- Sulphur-rumped Flycatcher, Myiobius sulphureipygius
- Whiskered Flycatcher, Myiobius barbatus
- Yellow-rumped Flycatcher, Myiobius mastacalis
- Black-tailed Flycatcher, Myiobius atricaudus
- Genus Neopipo
- Cinnamon Tyrant, Neopipo cinnamomea
- Genus Pyrrhomyias
- Cinnamon Flycatcher, Pyrrhomyias cinnamomea
- Genus Hirundinea - Cliff Flycatcher and Swallow Flycatcher
- Genus Cnemotriccus - Fuscous Flycatcher
- Genus Lathrotriccus (2 species)
- Genus Aphanotriccus (2 species)
- Genus Xenotriccus
- Belted Flycatcher, Xenotriccus callizonus
- Pileated Flycatcher, Xenotriccus mexicanus
- Genus Mitrephanes (2 species)
- Genus Contopus - pewees (14 species)
- Genus Empidonax (15 species)
- Genus Sayornis - phoebes (3 species)
- Genus Pyrocephalus - Vermilion Flycatcher
- Genus Silvicultrix
- Jelski's Chat-tyrant, Silvicultrix jelskii
- Yellow-bellied Chat-tyrant, Silvicultrix diadema
- Golden-browed Chat-tyrant, Silvicultrix pulchella
- Genus Ochthoeca
- Crowned Chat-tyrant, Ochthoeca frontalis
- Peruvian Chat-tyrant, Ochthoeca spodionota
- Slaty-backed Chat-tyrant, Ochthoeca cinnamomeiventris
- Maroon-chested Chat-tyrant, Ochthoeca thoracica
- Piura Chat-tyrant, Ochthoeca piurae
- D'Orbigny's Chat-tyrant, Ochthoeca oenanthoides
- Rufous-breasted Chat-tyrant, Ochthoeca rufipectoralis
- Brown-backed Chat-tyrant, Ochthoeca fumicolor
- White-browed Chat-tyrant, Ochthoeca leucophrys
- Genus Tumbezia
- Tumbes Tyrant, Tumbezia salvini
- Genus Colorhamphus - Patagonian Tyrant
- Genus Ochthornis
- Drab Water-tyrant, Ochthornis littoralis
- Genus Cnemarchus - Red-rumped Bush-tyrant
- Genus Myiotheretes
- Streak-throated Bush-tyrant, Myiotheretes striaticollis
- Santa Marta Bush-tyrant, Myiotheretes pernix
- Smoky Bush-tyrant, Myiotheretes fumigatus
- Rufous-bellied Bush-tyrant, Myiotheretes fuscorufus
- Genus Xolmis
- Fire-eyed Diucon, Xolmis pyrope
- Gray Monjita, Xolmis cinerea
- Black-crowned Monjita, Xolmis coronata
- White-rumped Monjita, Xolmis velata
- White Monjita, Xolmis irupero
- Rusty-backed Monjita, Xolmis rubetra
- Salinas Monjita, Xolmis salinarum
- Genus Heteroxolmis
- Black-and-white Monjita, Heteroxolmis dominicana
- Genus Neoxolmis
- Chocolate-vented Tyrant, Neoxolmis rufiventris
- Genus Agriornis - shrike-tyrants (5 species)
- Genus Polioxolmis
- Rufous-webbed Tyrant, Polioxolmis rufipennis
- Genus Muscisaxicola - ground-tyrants (13 species)
- Genus Muscigralla
- Short-tailed Field-tyrant, Muscigralla brevicauda
- Genus Lessonia
- Andean Negrito, Lessonia oreas
- Austral Negrito, Lessonia rufa
- Genus Knipolegus - black-tyrants (11 species)
- Genus Hymenops - Spectacled Tyrant
- Genus Fluvicola - typical water-tyrants (3 species)
- Genus Arundinicola - White-headed Marsh-tyrant
- Genus Alectrurus (2 species)
- Genus Gubernetes - Streamer-tailed Tyrant
- Genus Satrapa
- Yellow-browed Tyrant, Satrapa icterophrys
- Genus Colonia - Long-tailed Tyrant
- Genus Machetornis - Cattle Tyrant
- Genus Muscipipra
- Shear-tailed Gray Tyrant, Muscipipra vetula
- Genus Attila - attilas (7-8 species)
- Genus Casiornis (2 species)
- Genus Sirystes
- Sirystes, Sirystes sibilator
- Genus Rhytipterna
- Rufous Mourner, Rhytipterna holerythra
- Grayish Mourner, Rhytipterna simplex
- Pale-bellied Mourner, Rhytipterna immunda
- Genus Myiarchus (22 species)
- Genus Deltarhynchus - Flammulated Flycatcher
- Genus Pitangus - kiskadees (2 species, includes Philohydor)
- Genus Megarynchus - Boat-billed Flycatcher
- Genus Myiozetetes (4-5 species)
- Genus Conopias (4 species)
- Genus Myiodynastes (5 species)
- Genus Legatus - Piratic Flycatcher
- Genus Phelpsia
- White-bearded Flycatcher, Phelpsia inornata
- Genus Empidonomus - Variegated Flycatcher
- Genus Griseotyrannus - Crowned Slaty Flycatcher (formerly in Empidonomus)
- Genus Tyrannopsis - Sulphury Flycatcher
- Genus Tyrannus - kingbirds (13 species)
[edit] References
- ^ BirdLife International (2007). Species factsheets. Accessed 12 December 2007 available online
- ^ Adopt the Family Tityridae - South American Classification Committee (2007)
[edit] External links
- Tyrant Flycatcher videos on the Internet Bird Collection