Testudines

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Turtles
Fossil range: Late Triassic - Recent
"Chelonia" from Ernst Haeckel's Kunstformen der Natur, 1904
"Chelonia" from Ernst Haeckel's Kunstformen der Natur, 1904
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Superclass: Tetrapoda
Class: Reptilia
Subclass: Anapsida
Superorder: Chelonia
Macartney, 1802
Order: Testudines
Linnaeus, 1758
Diversity
14 extant families with ca. 300 species
blue: sea turtles, black: land turtles
blue: sea turtles, black: land turtles
Suborders

Cryptodira
Pleurodira
See text for families.

Turtles, Tortoises, and Terrapins are ectothermic reptiles, most of whose body is shielded by a special bony or cartilagenous shell developed from their ribs. They belong to an anapsid lineage, as can be seen from their solid skullcap. About 300 species are alive today; some are highly endangered.

The order to which the turtles belong is usually called Testudines. Sometimes Chelonia is given as a junior synonym, but this is strictly speaking incorrect. For most purposes, the differences are minor however: The Chelonia are simply the Testudines plus the "proto-turtle" Australochelys africanus, a turtlelike anapsid from the Jurassic which lacked some of the Testudines' apomorphies. The oldest fossils of the Testudines actually predate Australochelys. They are the basal turtle genus Proganochelys[1] which lived in Late Triassic Eurasia about 210 million years ago, making turtles one of the oldest reptile groups, and a much more ancient group than lizards and snakes.

Contents

[edit] Systematics[2]

[edit] Basal chelonians

[edit] Suborder Cryptodira

Basal genera

  • Genus Kayentachelys
  • Genus Indochelys

Infraorder †Paracryptodira

  • Basal and incertae sedis
    • Family †Kallokibotiidae
    • Family †Mongolochelyidae
    • Family †Pleurosternidae
    • Family †Solemydidae
  • Superfamily †Baenoidea
    • Family †Baenidae
    • Family †Macrobaenidae
    • Family †Neurankylidae

Infraorder Eucryptodira

  • "Sinemys" wuerhoensis
  • Genus †Chubutemys (Meiolaniidae?)
  • Genus †Hangaiemys (Macrobaenidae?)
  • Genus †Judithemys
  • Genus †Osteopygis
  • Genus †Planetochelys
  • Family Chelydridae (snapping turtles)
  • Family †Eurysternidae
  • Family †Macrobaenidae
  • Family †Meiolaniidae (horned turtles)
  • Family †Plesiochelyidae
  • Family †Sinemydidae
  • Family †Xinjiangchelyidae
  • Superfamily Chelonioidea (sea turtles)
The Western Hermann's Tortoise (Testudo hermanni hermanni) is a cryptodire.
The Western Hermann's Tortoise (Testudo hermanni hermanni) is a cryptodire.
The African Helmeted Turtle (Pelomedusa subrufa) is a pleurodire.
The African Helmeted Turtle (Pelomedusa subrufa) is a pleurodire.

[edit] Suborder Pleurodira

Basal and incertae sedis

  • Family †Araripemydidae
  • Family †Proterochersidae
  • Family Chelidae (Austro-American sideneck turtles)

Superfamily Pelomedusoidea

[edit] Footnotes

  1. ^ Formerly known as Triassochelys
  2. ^ See references in Haaramo (2008)

[edit] References

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