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An interesting-seeming article with likely relevent info & references: http://www.nature.com/cgi-taf/DynaPage.taf?file=/nature/journal/v425/n6953/full/nature01908_fs.html GREGORY J. VELICER AND YUEN-TSU N. YU (2003) "Evolution of novel cooperative swarming in the bacterium Myxococcus xanthus", Nature, 425, 75-78. It discusses how myxobacteria, and single-celled organisms in general, may have evolved mutually beneficial swarming behavior.