John Belling
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John Belling (1866-1933) born in Aldershot, England, was a cytogenetist who developed the iron-acetocarmine staining technique which is used in the study of chromosomes. He entered Mason College at the University of Birmingham in 1892 and received his BSc (Honours) from University College, London. He received an honorary DSc in 1922 from the University of Maine in recognition of his work.
In his work with Albert F. Blakeslee at Cold Spring Harbor on Datura (1920-1927) and at the University of California, Berkeley (1928-1933) Belling used plants such as lilies and hyacinths to demonstrate that segments between non-homogenous chromosomes can interchange. He was able to make accurate estimates of chromosome numbers and proposed that chromomeres, the small condensations along the chromosome, were individual genes.
Throughout his career Belling had many mental health problems that required frequent hospitalization. He died suddenly on February 28, 1933.
