Drepanosauridae

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Drepanosaurids
Fossil range: Late Triassic
Megalancosaurus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Subclass: Diapsida
(unranked) Avicephala
Family: Drepanosauridae
Pinna, 1979
Genera

Dolabrosaurus
Drepanosaurus (type)
Hypuronector
Megalancosaurus
Vallesaurus

Synonyms

Megalancosauridae Renesto, 1994

Drepanosauridae is a family of strange reptiles that lived during the late Triassic Period, 220 million years ago. The various species of drepanosaurid were characterized by odd specialized grasping limbs and often prehensile tails, adaptions for arboreal (tree-dwelling), and/or possibly aquatic lifestyles. Fossils of drepanosaurids, also known as Simiosaurs ("monkey lizards"), have been found in New Mexico, New Jersey, England, and northern Italy. The name Drepanosauridae is taken from the family's namesake genus Drepanosaurus, which means "sickle lizard", a reference to their strongly curved claws.

Contents

[edit] Description

Drepanosaurids are notable for their distinctive, triangular skulls, which resemble the skulls of birds. Some drepanosaurids, such as Hypuronector, had pointed, toothless, bird-like beaks. This similarity to birds may have led to the possible mis-attribution of a drepanosaurid skull to the would-be "first bird", Protoavis.[1]

Drepanosaurids featured a suite of bizarre, almost chameleon-like skeletal features. Above the shoulders of most species was a specialized "hump" formed from fusion of the vertebrae, possibly used for advanced muscle attachments to the neck, and allowing for quick forward-striking movement of the head (perhaps to catch insects). Many had derived hands with two fingers opposed to the remaining three, an adaptation for grasping branches. Some individuals of Megalancosaurus (possibly exclusive to either males or females) had a primate-like opposable toe on each foot, perhaps used by one sex for extra grip during mating. Most species had broad, prehensile tails, sometimes tipped with a large "claw", again to aid in climbing. These tails, tall and flat like those of newts and crocodiles, have led some researches to conclude that they were aquatic rather than arboreal. In 2004, Senter dismissed this idea, while Colbert and Olsen, in their description of Hypuronector, state that while other drepanosaurs were probably arboreal, Hypuronector was uniquely adapted to aquatic life.[2] The tail of this genus was extremely deep and non-prehensile – much more fin-like than other drepanosaurids.[3]

[edit] Systematics

[edit] Taxonomy

[edit] Phylogeny

Drepanosaurids have been difficult to pin down in terms of their phylogenetic position. They have been assigned by some resarches to the prolacertiformes, though more recent studies place them together with the coelurosauravids and Longisquama in a clade called the Avicephala.[4][2]

Within Avicephala, Senter created the group Simiosauria to refer to all the species traditionally grouped together within the Drepanosauridae, reserving that name for a more exlusive sub-group. Senter found that Hypuronector, originally described as a drepanosaurid, actually lies just outside that clade, along with the primitive drepanosaur Vallesaurus. He also recovered a close relationship between the drepanosaurs Dolabrosaurus and Megalancosaurus.[2]

The following cladogram was found by Senter in his 2004 analysis.[2]


Simiosauria

Vallesaurus


unnamed

Hypuronector


Drepanosauridae

Drepanosaurus


unnamed

Dolabrosaurus



Megalancosaurus







[edit] References

  1. ^ Renesto, S. (2000). "Bird-like head on a chameleon body: new specimens of the enigmatic diapsid reptile Megalancosaurus from the Late Triassic of northern Italy." Rivista Italiana di Paleontologia e Stratigrafia, 106: 157–180. Abstract
  2. ^ a b c d Senter, P. (2004). "Phylogeny of Drepanosauridae (Reptilia: Diapsida)." Journal of Systematic Palaeontology, 2(3): 257-268.
  3. ^ Colbert, E. H., and Olsen, P. E. (2001). "A new and unusual aquatic reptile from the Lockatong Formation of New Jersey (Late Triassic, Newark Supergroup)." American Museum Novitates, 3334: 1-24.
  4. ^ Renesto, S. (1994). "Megalancosaurus, a possibly arboreal archosauromorph (Reptilia) from the Upper Triassic of northern Italy." Journal of Vertebrate Paleontology, 14(1): 38-52.

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