Camptosaurus

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Camptosaurus

Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Superorder: Dinosauria
Order: Ornithischia
Suborder: Cerapoda
Infraorder: Ornithopoda
Family: Camptosauridae
Genus: Camptosaurus
Marsh, 1885
Species
  • C. dispar (Marsh, 1879 [originally Camptonotus dispar]) (type)
  • C. aphanoecetes Carpenter & Wilson, 2008
  • ?C. hoggii (Owen, 1874 [originally Iguanodon hoggii]; shown by Carpenter and Wilson, 2008, not to be a Camptosaurus)
  • ?C. prestwichii (Hulke, 1880 [originally Iguanodon prestwichii])
Synonyms
  • Camptonotus Marsh, 1879 non Uhler, 1864 (preoccupied)
  • ?Brachyrophus Cope, 1878
  • ?Cumnoria Seeley, 1888
  • ?Symphyrophus Cope, 1878

Camptosaurus (pronounced /ˌkæmptəˈsɔrəs/ KAMP-to-SAWR-us) is a genus of plant-eating, beaked dinosaurs of the Late Jurassic and Early Cretaceous Periods. The name means 'bent lizard', because, when standing on all fours, its body must have been arched (Greek kamptos meaning 'bent' and sauros meaning 'lizard').

[edit] Discovery and species

Camptosaurus aphanoecetes skeleton, Carnegie Museum.
Camptosaurus aphanoecetes skeleton, Carnegie Museum.

Originally described by O. C. Marsh in 1879 as Camptonotus, or "bent back", it was renamed Camptosaurus in 1885 because the original name was already in use for a cricket. In 1879, Marsh named C. dispar (type species of the genus) and C. amplus for material he received from his collectors at Quarry 13 near Como Bluff, Wyoming. Throughout the 1880-1890s, he continued to receive specimens from Quarry 13 and named two additional species: C. medius and C. nanus based in part on size. Charles W. Gilmore named two additional species, C. browni and C. depressus in his 1909 [1] redescription of the Marsh specimens. Then in 1980, Peter Galton and H.P. Powell in their redescription of C. prestwichi (see following), considered C. nanus, C. medius and C. browni to be different growth stages or different gender of the larger C. dispar, and therefore only C. dispar was the valid species. They also considered the skull referred to C. amplus by Marsh and by Gilmore to belong to C. dispar, as well. Gilmore had used this skull to describe the skull of Camptosaurus, but the specimen was recently shown by Brill and Carpenter [2] not to belong to Camptosaurus and put it into its own genus and species, Theiophytalia kerri. A second species of Camptosaurus was recently named by Carpenter and Wilson [3] for specimens from Dinosaur National monument. It differs from C. dispar in the lower jaw, shorter neck vertebrae, and straighter ischium ending in a small "foot" among other features.

Cast of Camptosaurus skull from Bone Cabin Quarry West, Wyoming.
Cast of Camptosaurus skull from Bone Cabin Quarry West, Wyoming.

While Marsh was describing Camptosaurus species in North America, numerous species from Europe were also referred to the genus in the late 1800s and early 1900s, as well: C. inkeyi, C. hoggii, C. leedsi, C. prestwichii, and C. valdensis. C. inkeyi and C. leedsi consist of fragmentary material and species are no longer considered valid (nomen dubium). C. valdensis was moved to Valdosaurus canaliculatus by Galton in 1977 [4]. C. hoggii was originally named Iguanodon hoggii by Richard Owen in 1874 and was moved to Camptosaurus by Norman and Barrett in 2002[5].

The remaining European species Camptosaurus prestwichii was recovered from Chawley Brick Pits, Cumnor Hurst in Oxfordshire in England. The fossil was found when a tramway was driven into the side of a hill. It was described by Hulke in 1880 as Iguanodon prestwichii, and then placed in its own genus Cumnoria by Seeley in 1888. It was moved to Camptosaurus by Lydekker in 1889 where it remains today.[6] The holotype skeleton (see picture) remains the best skeleton of Camptosaurus from Europe.

[edit] Paleobiology

Camptosaurus (incomplete skeleton), Oxford University Museum of Natural History
Camptosaurus (incomplete skeleton), Oxford University Museum of Natural History

The largest adult camptosaurs were more than 7.9 meters long (26 ft), and 2.0 meters tall (6.7 ft) at the hips.[7] They had heavy bodies but, as well as walking on four legs (quadrupedal), they could rear up to walk on two legs (bipedal).

This genus is probably closely related to the ancestor of the later iguanodontid and hadrosaurid dinosaurs. It probably ate cycads with its parrot-like beak.

[edit] References

  1. ^ Gilmore, C.W.. Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings, U.S. National Museum, vol. 36:197-302. 
  2. ^ Brill, K. and K. Carpenter. 2006. A description of a new ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a reassessment of the skull of Camptosaurus. pp. 49-67. In K. Carpenter, ed., 2006, Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington.
  3. ^ Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Moument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum 76:227-263 PDF
  4. ^ Galton, P. M. 1977. The Upper Jurassic dinosaur Dryosaurus and a Laurasia-Gondwana connection in the Upper Jurassic. Nature 268(5617):230-232
  5. ^ Norman D.B. and Barrett, P.M. 2002. Ornithischian dinosaurs from the Lower Cretaceous (Berriasian) of England. Palaeontology 68:161-189
  6. ^ Benton MJ, Spencer PS (1995). Fossil Reptiles of Great Britain. Chapman & Hall. ISBN 0-412-62040-5. 
  7. ^ Erickson, Bruce R. (2003). Dinosaurs of the Science Museum of Minnesota. St. Paul, Minnesota: The Science Museum of Minnesota, 33. ISBN 911338-54-3.