Banffia

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Banffia
Fossil range: Cambrian

Banffia confusa & friends

Kingdom:	Animalia
Superphylum:	?Deuterostomia
Phylum:	?Vetulicolia Shu, et al. 2001
Family:	?Vetulicolidae
Genus:	Banffia

Species

B. confusa
B. constricta

Banffia is a genus of animals described from Early Cambrian fossils. The genus was named after Banff, Alberta, near where they were first discovered. Its placement in higher taxa is controversial. The species of Banffia were soft-bodied, and thus fossils are known primarily from the Burgess shales of Canada and the Chengjiang shales of China.

Banffia constricta is known from hundreds of fossils found in the Burgess shales. It is up to 10 cm in length, and divided equally into anterior and posterior parts. The entire body is twisted in a clockwise spiral, as seen from the front. This is believed to be a secondary adaptation from an initial bilateral condition. The anterior section is covered by two carapace-like un-mineralized shells that are fused together. A crown-like structure formed of three concentrate circular features surrounds the mouth. An antenna-form structure just posterior to the mouth may be a sensory organ. The posterior section is composed of 40 to 50 segments. The gut is straight, and the anus is at the terminal tip of the posterior section. The gut appears to have a series of diverticula or pouches. A possible circulatory system is visible in the fossils. Banffia constricta were probably filter or deposit feeders.

Banffia confusa is known from fossils from the Chengjiang shales. The animal also is divided, with segmentation visible in the posterior section. It is interpreted as either a swimming filter feeder or detrivore.

[edit] Classification

There is no agreement on the classification of Banffia. Banffia constricta was assigned to the annelids by Walcott in 1911. As of 2006, different proposals would place Banffia in Urochordata, Vetulicolia or Arthropoda. While the body plan (equal anterior and posterior sections with segmentation) resembles that of the Vetulicolians, it is argued that the absence of gills and an endostyle, and the presence of gut diverticula makes Banffia unlikely to be a member of the deuterostomes.