The Structure of Evolutionary Theory

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Title The Structure of Evolutionary Theory
Author Stephen Jay Gould
Genre(s) Non-fiction, Science
Publisher Belknap Press
Released March 21, 2002
Pages 1,433
ISBN ISBN 0-674-00613-5
Preceded by The Lying Stones of Marrakech
Followed by I Have Landed

The Structure of Evolutionary Theory (2002) is a technical book on macroevolutionary theory by the Harvard paleontologist Stephen Jay Gould. The volume is divided into two parts. The first is a historical study and exegesis of classical evolutionary thought. The second is a constructive critique of contemporary Darwinian theory, and presents a case for a hierarchical interpretation of biological evolution based largely on the author's theory of punctuated equilibrium.

According to Gould, classical Darwinism encompasses three essential core commitments. These are: agency, efficacy, and scope. Agency is the unit upon which natural selection acts. For Darwin, this fundamental unit was the organism. Efficacy encompasses the power of natural selection—over all other forces—in shaping evolution at ecological scales. (Auxiliary forces include sexual selection, as well as historical, structural, and developmental constraints.) Scope is the degree to which natural selection can be extrapolated to explain biological diversity at the macroevolutionary level, including the evolution of higher taxonomic groups.

Gould described these three propositions as the "tripod" of Darwinian central logic. Each being so essential to the structure that if any branch were cut, it would either kill, revise, or superficially refurbish the whole structure (depending on the severity of the cut). According to Gould "substantial changes, introduced during the last half of the 20th century, have built a structure so expanded beyond the original Darwinian core, and so enlarged by new principles of macroevolutionary explanation, that the full exposition, while remaining within the domain of Darwinian logic, must be construed as basically different from the canonical theory of natural selection, rather than simply extended."

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