Stegosauria

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Stegosauria
Fossil range: Middle Jurassic - Early Cretaceous

Fossil skeleton of a Stegosaurus,
National Museum of Natural History

Extinct (fossil)

Scientific classification

Kingdom:	Animalia
Phylum:	Chordata
Class:	Sauropsida
Superorder:	Dinosauria
Order:	Ornithischia
Suborder:	Thyreophora
Infraorder:	Stegosauria Marsh, 1877

Known colloquially as stegosaurs, the Stegosauria are a group of herbivorous dinosaurs of the Jurassic and Early Cretaceous Periods, being found mostly in the Northern Hemisphere, predominantly in what is now North America and China. Their geographical origins are unclear; the earliest stegosaurs have been found in China, although fragmentary material hails from southern England.

The genus Stegosaurus, from which the group acquires its name, is by far the most famous stegosaurian.

1 Paleobiology
- 1.1 Skull
- 1.2 Posture
2 Taxonomy
- 2.1 Classification
- 2.2 Phylogeny
3 Undescribed species
4 References
5 External links

[edit] Paleobiology

All stegosaurs have rows of special bones, called osteoderms, which develop into plates and spines along the back and tail. Many also have intermediate ones, called 'splates'.

[edit] Skull

They had characteristic long, narrow heads and a horn-covered beak or rhamphotheca, which covered the front of the upper jaw (premaxillary) and lower jaw (predentary) bones. Similar structures are seen in turtles and birds. Apart from Huayangosaurus, stegosaurs subsequently lost nearby premaxillary teeth.^[1]

[edit] Posture

All are quadrupedal, with hoof-like toes on all four limbs. All stegosaurians after Huayangosaurus have forelimbs much shorter than their hindlimbs. Given that their speed would have been limited by their shortest limb and their size is likely to have precluded them from being bipedal, this suggests that they were not able to run quickly.

[edit] Taxonomy

The Stegosauria was originally named as an order within Reptilia by O.C. Marsh in 1877,^[2] although today it is generally treated as an infraorder or suborder (or simply a clade) within Thyreophora, the armored dinosaurs. It includes the families Huayangosauridae and Stegosauridae.

The Huayangosauridae were an early family of stegosaurs which lived during the early to middle Jurassic Period. In general, they were smaller than later stegosaurs and had shorter and higher skulls. Currently, the only confirmed genus included is the type genus Huayangosaurus of China. The poorly-known remains of Regnosaurus from England, however, indicate it too could be a member. Its lower jaw is very similar to the former.

The vast majority of Stegosaurian dinosaurs thus far recovered being to the Stegosauridae, which lived in the later part of the Jurassic and early Cretaceous, with one possible genus (Dravidosaurus) from the late Cretaceous. It includes the well-known Stegosaurus. The family is widespread, with members across the Northern Hemisphere and Africa.

[edit] Classification

A fossil melee involving a stegosaurian (Tuojiangosaurus) and a mid-sized theropod (Monolophosaurus), Field Museum in Chicago

Following is a list of stegosaurian genera by classification and location:

Suborder Thyreophora

Infraorder Stegosauria

Gigantspinosaurus
Family Huayangosauridae
- Huayangosaurus (Sichuan,China)
- ?Regnosaurus (Sussex, United Kingdom)
- ?Tatisaurus (Yunnan,China)

Family Stegosauridae
- Paranthodon - (South Africa)
- Monkonosaurus - (Tibet, China)
- Dravidosaurus - (India)
- Chungkingosaurus - (Chongqing, China)
- Chialingosaurus - (Sichuan, China)
- Wuerhosaurus - (Xinjiang, Western China)
- Hesperosaurus - (Wyoming, USA)
- Dacentrurus- (United Kingdom, France & Spain)
- Subfamily Stegosaurinae
  - Tuojiangosaurus - (Sichuan, China)
  - Kentrosaurus - (Tanzania, Africa)
  - Lexovisaurus - (United Kingdom & France)
  - Stegosaurus - (Wyoming, USA)
  - Hypsirophus- (Colorado, USA)
- incertae sedis
  - ?Craterosaurus (Bedfordshire, England)

[edit] Phylogeny

Kenneth Carpenter of the Denver Museum of Nature and Science published a preliminary phyletic tree^[3] of stegosaurs, in the 2001 description of Hesperosaurus. Here, the basal stegosaur Huayangosaurus is used as the outgroup. The Stegosauridae are then defined as all stegosaurs closer to Stegosaurus than to Huayangosaurus. The position of Chungkingosaurus is uncertain due to lack of data.

Stegosauria
|--Huayangosaurus
`--Stegosauridae
   `--+-?Chungkingosaurus
      `--+--Chialingosaurus
         `--+--+--Wuerhosaurus
            |  `--+--Dacentrurus
            |     `--Hesperosaurus
            `--+--Tuojiangosaurus 
               `--+--+--Kentrosaurus            
                  |  `--Lexovisaurus
                  `--+--Stegosaurus stenops
                     `--S. ungulatus (=?S. armatus)

[edit] Undescribed species

To date, several genera from China bearing names have not been formally described: Gigantspinosaurus, Changdusaurus and Yingshanosaurus. Until formal descriptions are published, each of these genera is regarded as a nomen nudum.

[edit] References

^ Sereno, P & Z-M Dong (1992). The skull of the basal stegosaur Huayangosaurus taibaii and a cladistic diagnosis of Stegosauria. Journal of Vertebrate Paleontology 51: 318-343
^ Marsh, O.C. (1877). "New order of extinct Reptilia (Stegosauria) from the Jurassic of the Rocky Mountains." American Journal of Science, 14(ser.3):513-514.
^ Carpenter, K., Miles, C.A., and Cloward, K. (2001). "New Primitive Stegosaur from the Morrison Formation, Wyoming", in Carpenter, Kenneth(ed) The Armored Dinosaurs. Indiana University Press. ISBN 0-253-33964-2, 55–75.

Fastovsky DE, Weishampel DB (2005). "Stegosauria:Hot Plates", in Fastovsky DE, Weishampel DB: The Evolution and Extinction of the Dinosaurs (2nd Edition). Cambridge University Press, 107–130. ISBN 0-521-81172-4.