Four-eyed fish

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Four-eyed fishes

Anableps sp.

Scientific classification

Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cyprinodontiformes
Family:	Anablepidae
Genus:	Anableps Scopoli, 1777

Species

See text.

The four-eyed fishes are a genus, Anableps, of fishes in the family Anablepidae. They have eyes raised above the top of the head and divided in two different parts, so that they can see below and above the water surface at the same time. Like their relatives, the onesided livebearers, they only mate on one side, right-"handed" males with left-"handed" females and vice versa. These fish inhabitat freshwater and brackishwater and are only rarely coastal marine. They originate from lowlands in southern Mexico to Honduras and northern South America.^[1]

1 Behavior
2 Physical characteristics
- 2.1 Reproductive organs
- 2.2 Eye
3 Relationship to humans
4 Species
5 References
- 5.1 General References

[edit] Behavior

These fish spend most of their time at the surface of the water. Their diet mostly consists of terrestrial insects which are readily available at the surface, however they may consume other foods such as other invertebrates, diatoms, and small fishes.^[2]

These fish will group differently depending on the species. A. anableps commonly congegrates in schools.^[2] A. microlepis also is gregarious, but restricts its schools to about a dozen individuals; it is also recorded to be found singly or as couples.^[3]

A. anableps is also known for the ability to survive out of water exposed to air, such as during low tide.^[2]

[edit] Physical characteristics

The maximum length of these species is up to 32 cm TL in A. microlepis, making this species the largest in the order Cyprinidontiformes.^[3]

[edit] Reproductive organs

Main article: Anablepidae

These fish are one-sided livebearers and their reproductive organs are unique to its subfamily. This characteristic is only shared with the related genus Jenynsia.

[edit] Eye

The four-eyed fishes only have two eyes each, but the eyes are specially adapted for their surface-dwelling lifestyle. The eyes are positioned on the top of the head, and the fish floats at the water surface with only the lower half of each eye underwater. The two halves are divided by a band of tissue and the eye has two pupils, connected by part of the iris. The upper half of the eye is adapted for vision in air, the lower half for vision in water.^[1] The lens of the eye also changes in thickness top to bottom to account for the difference in the refractive indices of air versus water.

[edit] Relationship to humans

Four-eyed fish are often used in eye research.^[2]^[3]^[4]

This fish is occasionally found in pet stores.

[edit] Species

Largescale foureyes, Anableps anableps (Linnaeus, 1758).
Pacific foureyed fish, Anableps dowei Gill, 1861.
Foureyes, Anableps microlepis Müller & Troschel, 1844.

[edit] References

^ ^a ^b Nelson, Joseph, S. (2006). Fishes of the World. John Wiley & Sons, Inc.. ISBN 0471250317.
^ ^a ^b ^c ^d "Anableps anableps". FishBase. Ed. Ranier Froese and Daniel Pauly. Mar 2007 version. N.p.: FishBase, 2007.
^ ^a ^b ^c "Anableps microlepis". FishBase. Ed. Ranier Froese and Daniel Pauly. Mar 2007 version. N.p.: FishBase, 2007.
^ "Anableps dowei". FishBase. Ed. Ranier Froese and Daniel Pauly. Mar 2007 version. N.p.: FishBase, 2007.