Dinosaur Park Formation
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The Dinosaur Park Formation is the uppermost member of the Judith River Group, a major geologic unit in southern Alberta. It was laid down roughly 75 million years ago. The formation is made up of deposits of a high-sinuosity (anastomosing) fluvial system, and is capped by the Lethbridge Coal Beds. The formation is bounded by the Oldman Formation below it and the marine Bearpaw Formation above it (Eberth, 2005).
It is best known for the dense concentrations of dinosaur skeletons, both articulated and disarticulated, that are found there. However, other animals such as fish, turtles, crocodilians are also abundant in the formation. The formation has been named after Dinosaur Provincial Park, a UNESCO World Heritage Site.
[edit] Stratigraphic Division of Ornithischians? (Ryan and Evans, 2005)
Centrosaurus, Chasmosaurus russelli, Gryposaurus, and Corythosaurus are most common at the base of the formation, to about the middle. Styracosaurus, Chasmosaurus belli, Prosaurolophus, and Lambeosaurus lambei are most common from the middle to near the top. A new group may be present at the top, as the inland sea transgresses onto land, but there are less remains here. An unnamed pachyrhinosaur, Chasmosaurus irvinensis, and Lambeosaurus magnicristatus may be more common here.
[edit] Flora and Fauna
Numerous types of plants and animals are known from the formation (taxonomy mostly after the relevant chapters of Currie and Koppelhus, 2005):
[edit] Palynomorphs (organic-walled microfossils, like spores, pollen, and algae) (Braman and Koppelhus, 2005)
Unknown producers
- at least 8 species
- at least 35 taxa
Chlorophyta (green algae and blue-green algae)
- at least 12 species
Pyrrhophyta (dinoflagellates, a type of marine algae)
- unassigned cysts
Bryophyta (mosses, liverworts, and hornworts)
- Anthocerotae (hornworts)
- at least 5 species
- Hepaticae (liverworts)
- at least 14 species
- Musci (mosses)
- at least 5 species
- Lycopodiaceae (club mosses)
- at least 11 species
- Selaginellaceae (small club mosses)
- at least 6 species
- Isoetaceae (quillworts)
- at least 1 species
- Osmundaceae (cinnamon ferns)
- at least 6 species
- Schizaeaceae (climbing ferns)
- at least 20 species
- Gleicheniaceae (Gleichenia and allies; coral ferns)
- at least 5 species
- Cyatheaceae (Cyathea and allies)
- at least 4 species
- Dicksoniaceae (Dicksonia and allies)
- at least 3 species
- Polypodiaceae (ferns)
- at least 4 species
- Matoniaceae
- at least 1 species
- Marsileaceae
- at least 1 species
- Cycadaceae (cycads)
- at least 3 species
- Caytoniaceae
- at least 1 species
- Pinaceae (pines)
- at least 4 species
- Cupressaceae (cypresses)
- at least 3 species
- Podocarpaceae (Podocarpus and allies)
- at least 4 species
- Cheirolepidiaceae
- at least 2 species
- Ephedraceae (Mormon teas)
- at least 6 species
Unknown gymnosperms: at least 3 species
- Magnoliopsida (dicots)
- Buxaceae (boxwood)
- at least 1 species
- Gunneraceae (gunneras)
- at least 1 species
- Salicaceae (willows, cottonwood, quaking aspen)
- at least 1 species
- Droseraceae (sundews)
- at least 1 species
- Olacaceae (tallowwood)
- at least 2 species
- Loranthaceae (showy mistletoes)
- at least 1 species
- Sapindaceae (soapberry)
- at least 1 species
- Aceraceae (maples)
- at least 1 species
- Proteaceae (proteas)
- at least 9 species
- Compositae (sunflowers)
- at least 1 species
- Fagaceae (beeches, oaks, chestnuts)
- at least 2 species
- Betulaceae (birches, alders)
- at least 1 species
- Ulmaceae (elms)
- at least 1 species
- Chenopodiaceae (goosefoots)
- at least 1 species
- Buxaceae (boxwood)
- Liliopsida (monocots)
- Liliaceae (lilies)
- at least 6 species
- Cyperaceae (sedges and reeds)
- at least 1 species
- Sparganiaceae (bur-reeds)
- possibly 1 species
- Liliaceae (lilies)
- Unknown angiosperms: at least 88 species
[edit] Plant body fossils (Koppelhus, 2005)
- various ferns
- Equisetum (Equisetaceae)
- Platyspiroxylon (Cupressaceae)
- Podocarpoxylon (Podocarpaceae)
- Elatocladus (Taxodiaceae)
- Sequoia (Taxodiaceae)
- Sequoiaxylon (Taxodiaceae)
- Taxodioxylon (Taxodiaceae)
- Baeria
- Ginkgoites
- Artocarpus (Moraceae)
- Cercidiphyllum (Cercidiphyllaceae)
- Dombeyopsis (Sterculiaceae)
- Menispermites (Menispermaceae)
- Pistia (Araceae)
- Platanus (Platanaceae)
- Vitis (Vitaceae)
- Trapa (Trapaceae)
[edit] Mollusks (Johnston and Hendy, 2005)
- Fusconaia
- Lampsilis
- Sphaerium (2 species)
Freshwater gastropods
- Campeloma (2 species)
- Elimia
- Goniobasis (3 species)
- Hydrobia
- Lioplacodes (2 species)
[edit] Fish
Chondrichthyans (Neuman and Brinkman, 2005)
- Hybodus (a shark)
- Myledaphus (a ray)
- indeterminate orectolobid
Acipenseriformes (sturgeons) (Neuman and Brinkman, 2005)
- unnamed sturgeon
- unnamed paddlefish
Holostean fish (Neuman and Brinkman, 2005)
- Belonostomus
- Lepisosteus (the gar)
- unnamed bowfin
- at least 2 other holosteans
Teleost fish (Neuman and Brinkman, 2005)
- Patatarpon (an elopomorph, like the tarpon)
- Cretophareodus (an osteoglossomorph)
- Coriops
- Estesesox
- Oldmanesox
- Paralbula (including Phyllodus)
- at least 8 other teleosts
[edit] Amphibians (Gardner, 2005)
Albanerpetonidae (extinct, salamander-like amphibians)
- Albanerpeton
- Habrosaurus
- Lisserpeton
- Opisthotriton
- Scapherpeton
- unnamed caudatan
- 2 indeterminate caudatans
Salienta (frogs)
- 2 unnamed salientans
[edit] Turtles (Brinkman, 2005)
- Adocus
- "Apalone"
- Aspideretoides (3 species)
- Basilemys
- Boremys
- Judithemys
- Neurankylus
- Plesiobaena
- 2 indeterminate taxa
[edit] Choristoderes (champsosaurs) (Keqin Gao and Brinkman, 2005)
- Champsosaurus (at least 3 species)
- Cteniogenys
[edit] Lizards (Caldwell, 2005)
- Labrodioctes
Necrosaurids
- Parasaniwa
- Glyptogenys
- Socognathus
- ?Exostinus
[edit] Plesiosaurs (Sato, Eberth, Nicholls, and Manabe, 2005)
- indeterminate elasmosaurids (very long-necked) (more common)
- indeterminate polycotylids (shorter-necked)
[edit] Crocodylians (Xiao-Chun Wu, 2005)
- Albertochampsa
- Leidysuchus
- at least 1 unnamed taxon
[edit] Pterosaurs (Godfrey and Currie, 2005)
- 1 large unnamed azhdarchid (giant, long-necked pterosaur)
- 1 smaller unnamed azdarchid
- 1 unnamed non-azdarchid pterosaur
[edit] Dinosaurs
Theropoda (Currie, 2005)
- Tyrannosauridae
- Daspletosaurus torosus
- D. sp.
- Gorgosaurus libratus (also called Albertosaurus libratus
- Ornithomimidae
- Caenagnathidae (all of these may be species of Chirostenotes)
- Caenagnathus collinsi
- C. sternbergi
- Chirostenotes pergracilis
- Elmisaurus rarus
- Avimimidae
- ?indeterminate avimimid
- Therizinosauridae
- ?indeterminate therizinosaurid
- Troodontidae
- Troodon inequalis or formosus
- Dromaeosauridae
- Family unknown
- Birds
- Baptornis sp.
- Cimolopteryx sp.
- Palintropus sp.
- additional indeterminate bird taxa
Ornithischia (Ryan and Evans, 2005)
- Ankylosauria
- Pachycephalosauridae
- Hanssuesia sternbergi
- Gravitholus albertae
- Ornatotholus browni (?juvenile Stegoceras)
- Stegoceras validum
- "S." breve (?Prenocephale)
- Ceratopsia
- Leptoceratops sp.
- Ceratopsidae
- Centrosaurinae (long nose horn, short brow horns, short spiky frill)
- Centrosaurus apertus
- Monoclonius lowei
- Styracosaurus albertensis
- unnamed Pachyrhinosaurus-like taxon
- Chasmosaurinae (short nose horn, long brow horns, long frill)
- Anchiceratops ornatus
- Chasmosaurus belli
- C. irvinensis
- C. russelli
- Centrosaurinae (long nose horn, short brow horns, short spiky frill)
- Ornithopoda
- at least 1 indeterminate hypsilophodont
- Hadrosauridae
- Hadrosaurinae (duckbills without hollow crests)
- Gryposaurus notabilis
- G. incurvimanus
- Prosaurolophus maximus
- Lambeosaurinae (duckbills with hollow crests)
- Corythosaurus casuarius
- Lambeosaurus lambei
- L. magnicristatus
- Parasaurolophus walkeri
- Hadrosaurinae (duckbills without hollow crests)
[edit] Mammals (Fox, 2005)
- Cimexomys sp.
- Cimolodon spp.
- Cimolomys clarki
- Meniscoessus major
- Mesodma primaeva
- unnamed multituberculates
- Alphadon halleyi
- Eodelphis browni
- E. cutleri
- 5 species of "Pediomys"
- Turgidodon russelli
- T. praesagus
- Cimolestes sp. (uncertain taxonomy)
- Gypsonictops lewisi
- Paranyctoides sternbergi
Unknown therians: at least 1 species
[edit] References
- Braman, D.R., and Koppelhus, E.B. 2005. Campanian palynomorphs. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 101-130.
- Brinkman, D.B. 2005. Turtles: diversity, paleoecology, and distribution. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 202-220.
- Caldwell, M.W. The squamates: origins, phylogeny, and paleoecology. In: Currie, P.J., and Koppelhus, E.B. (eds). 2005. ‘’Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed.’’ Indiana University Press: Bloomington and Indianapolis, 235-248.
- Currie, P.J. 2005. Theropods, including birds. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 367-397.
- Currie, P.J., and Koppelhus, E.B. (eds). 2005. Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 648 p.
- Eberth, D.A. 2005. The geology. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 54-82.
- Fox, R.C. 2005. Late Cretaceous mammals. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 417-435.
- K. Gao and Brinkman, D.B. 2005. Choristoderes from the Park and its vicinity. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 221-234.
- Gardner, J.D. 2005. Lissamphibians. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 186-201.
- Godfrey, S.J., and Currie, P.J. 2005. Pterosaurs. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 292-311.
- Johnston, P.A., and Hendy, A.J.W. 2005. Paleoecology of mollusks from the Upper Cretaceous Belly River Group. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 139-166.
- Koppelhus, E.B. 2005. Paleobotany. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 131-138.
- Neuman, A.G., and Brinkman, D.B. 2005. Fishes of the fluvial beds. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 167-185.
- Ryan, M.J., and Evans, D.C. 2005. Ornithischian dinosaurs. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 312-348.
- Sato, T., Eberth, D.A., Nicholls, E.L., and Manabe, M. 2005. Plesiosaurian remains from non-marine to paralic sediments. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed.’’ Indiana University Press: Bloomington and Indianapolis, 249-276.
- Xiao-Chun Wu. 2005. Crocodylians. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, 277-291.