Chasmosaurus

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Chasmosaurus
Fossil range: Late Cretaceous

Conservation status
Extinct (fossil)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Superorder: Dinosauria
Order: Ornithischia
Suborder: Marginocephalia
Infraorder: Ceratopsia
Superfamily: Ceratopsoidea
Family: Ceratopsidae
Subfamily: Ceratopsinae
Genus: Chasmosaurus
Lambe, 1914
Species

C. belli
C. canadensis
C. irvinensis
C. russelli

Chasmosaurus (KAZ-mo-sawr-us) is a ceratopsid dinosaur genus from the Upper Cretaceous Period of North America. Its name means 'opening lizard', referring to the large openings (fenestrae) in its frill (Greek chasma meaning 'opening' or 'hollow' or 'gulf' and sauros meaning 'lizard'). With a length of 5 - 6 metres and a weight of 3.6 tonnes, Chasmosaurus was a ceratopsian of average size. Like all ceratopsians, it was purely herbivorous. It was initially to be called Protorosaurus, however this name had been previously published for another animal.

Contents

[edit] Discoveries and species

Skull of a Chasomosaurus belli, on display at the Royal Ontario Museum, Toronto.
Skull of a Chasomosaurus belli, on display at the Royal Ontario Museum, Toronto.

Chasmosaurus fossils were first recovered in 1902 but were thought to be from a previously-known short-frilled ceratopsian - Monoclonius. However, in 1913, Charles Sternberg and his sons found several complete skulls of what is now known as Chasmosaurus, in Alberta, Canada. These were finally described in 1914, by Lawrence M. Lambe of the Geological Survey of Canada. Since that date, more skulls have been found. There are some differences across these skulls, detailed below.

There are a number of known species of Chasmosaurus. Lambe's original C. belli ('Bell's cleft lizard') was joined by C. canadensis ('chasm lizard from Canada') in the same year. The latter species had been described as Eoceratops canadensis by Lambe but was later reclassified as a chasmosaur by Lehman. Lull named an unusual, short-muzzled skull, collected in 1926, C. brevirostris. C. M. Sternberg added C. russelli, in 1940, from southwestern Alberta. The most recently described species is C. irvinensis, which stems from the uppermost beds of the Dinosaur Park Formation.

Lehman described C. mariscalensis in 1989 from Texas, which has now been renamed Agujaceratops.

[edit] Characteristics

Ceratopsians are split into two subfamilies by taxonomists; those with short frills (centrosaurines), such as Centrosaurus and those with long frills (chasmosaurines), of which Chasmosaurus was one. In addition to the larger frill, the long-frilled ceratopsians typically had longer faces and jaws and it is suggested by some paleontologists that they were more selective about the plants they ate. Long frills were a relatively late development in dinosaur evolution, since even Chasmosaurus dates from the late Cretaceous Period, 76 to 70 million years ago. The frill of Chasmosaurus has been described as "heart-shaped", since its bone structure consists of two large 'loops' from a central bone.

Some finds include a number of smaller ossifications (called epoccipitals), which may have grown from the perimeter of the frill. The frill may also have been brightly coloured, to draw attention to its size or as part of mating display. However, the frill was so large and yet so flimsy (since it was mainly skin stretched between the bones) that it could not have provided much functional defence. It is possible that it was simply used to appear imposing or conceivably for thermoregulation. In the event of a chasmosaur herd being attacked by a predator (such as Tyrannosaurus), the males could have formed a ring and, with all the frills facing outwards, would have presented a formidable sight.

Like many ceratopsians, chasmosaurs had three main facial horns - one on the nose and two on the brow. Different fossil finds have produced inconclusive results - one species of Chasmosaurus, named C. kaiseni, bore long brow horns, while C. belli had only short ones. Although these were initially named as different species, it now seems possible that the long horns belonged to males and the shorter horns to females.

Interestingly, paleontologists have recovered some fossilized chasmosaur skin. The skin appears to have had many bony knobs (osteoderms), with five or six sides each. Unfortunately, nothing more can be learned from these samples - the colour of dinosaurs remains a mystery.

[edit] In popular culture

In a Calvin and Hobbes comic, Chasmosaurus was depicted as the main prey for tyrannosaurs in F-14 fighter jets.

Chasmosaurus appeared in When Dinosaurs Ruled the Earth.

[edit] External links