List of subgroups of the order Coleoptera

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This article classifies the subgroups of the order Coleoptera (beetles) down to the level of families, following the system in Volume 2 of American Beetles (2002) for all North American families (there were changes in the system between Vols. 1 & 2), and following Lawrence & Newton (1995) for extralimital taxa. A few subfamilies, tribes and synonyms are shown here where a family has been reclassified. Many of the common names listed below were newly-coined for these volumes, and are not in vernacular or widespread use.

Order Coleoptera (beetles)

• Suborder Adephaga Schellenberg 1806
  • Amphizoidae LeConte 1853 (trout-stream beetles)
  • Carabidae Latreille 1802 (ground beetles)
    • Cicindelinae, formerly Cicindelidae Latreille 1802 (tiger beetles)
    • Paussinae, formerly Paussidae Latreille 1807
  • Dytiscidae Leach 1815 (predaceous diving beetles)
    

    

    Dytiscus latissimus, a predaceous diving beetle
  • Gyrinidae Latreille 1802 (whirligig beetles)
  • Haliplidae Aube 1836 (crawling water beetles)
  • Hygrobiidae Régimbart 1878 (1837)
  • Noteridae C.G. Thomson 1860 (burrowing water beetles)
  • Rhysodidae Laporte 1840 (wrinkled bark beetles)
  • Trachypachidae C.G. Thomson 1857(false ground beetles)
• Suborder Archostemata Kolbe 1908
  • Crowsoniellidae Iablokoff-Khnzorian 1983
  • Cupedidae Laporte 1836 (reticulated beetles)
    

    

    Priacma serrata, a reticulated beetle
  • Micromalthidae Barber 1983 (telephone-pole beetles)
  • Ommatidae Sharp and Muir 1912
• Suborder Myxophaga Crowson 1955
  • Hydroscaphidae LeConte 1874 (skiff beetles)
  • Lepiceridae Hinton 1936
  • Sphaeriusidae Erichson 1845 (minute bog beetles) (= Microsporidae: Bulletin of Zoological Nomenclature 57(3): 182-184.)
  • Torridincolidae Steffan 1964
• Suborder Polyphaga Emery 1886
  • Infraorder Bostrichiformia Forbes 1926
    • Superfamily Bostrichoidea Latreille 1802
      • Anobiidae Fleming 1821 (death watch beetles)
        • Ptininae, formerly Ptinidae Latreille 1802 (spider beetles)
      • Bostrichidae Latreille 1802 (horned powder-post beetles)
        • Lyctinae, formerly Lyctidae Billberg 1820 (powder post beetles)
        • Endecatominae, formerly Endecatomidae LeConte 1861
      • Dermestidae Latreille 1804 (skin beetles)
        

        

        Anthrenus verbasci, a skin beetle
        • Thorictinae, formerly Thorictidae Agassiz 1846
      • Jacobsoniidae Heller 1926 (Jacobson's beetles)
      • Nosodendridae Erichson 1846 (wounded-tree beetles)
    • Superfamily Derodontoidea LeConte 1861
      • Derodontidae LeConte 1861 (tooth-necked fungus beetles)
  • Infraorder Cucujiformia Lameere 1938
    • Superfamily Chrysomeloidea Latreille 1802
      • Cerambycidae Latreille 1802 (long-horned beetles)
      • Chrysomelidae Latreille 1802 (leaf beetles)
        

        

        Colorado potato beetle, Leptinotarsa decemlineata, a leaf beetle
        • Cassidini, formerly Cassidinae
        • Bruchinae, formerly Bruchidae Latreille 1802
      • Megalopodidae Latreille 1802
      • Orsodacnidae C. G. Thomson 1869
    • Superfamily Cleroidea Latreille 1802
      • Acanthocnemidae Crowson 1964
      • Chaerosomatidae Crowson 1952
      • Cleridae Latreille 1802 (checkered beetles)
      • Melyridae Leach 1815 (soft-winged flower beetles)
      • Phloiophilidae Kiesenwetter 1863
      • Phycosecidae Crowson 1952
      • Prionoceridae Lacordaire 1857
      • Trogossitidae Latreille 1802 (bark-gnawing beetles)
    • Superfamily Cucujoidea Latreille 1802
      • Alexiidae Imhoff 1856
      • Biphyllidae LeConte 1861 (false skin beetles)
      • Boganiidae Sen Gupta and Crowson 1966
      • Bothrideridae Erichson 1845 (dry bark beetles)
      • Byturidae Jacquelin du Val 1858 (fruitworm beetles)
      • Cavognathidae Sen Gupta and Crowson 1966
      • Cerylonidae Billberg 1820 (minute bark beetles)
      • Coccinellidae Latreille 1807 (lady beetles)
        

        

        Asian multicolored lady beetle, Harmonia axyridis
      • Corylophidae LeConte 1852 (minute fungus beetles)
      • Cryptophagidae Kirby 1937 (silken fungus beetles)
      • Cucujidae Latreille 1802 (flat bark beetles)
      • Discolomatidae Horn 1878
      • Endomychidae Leach 1815 (handsome fungus beetles)
        • Merophysiinae, formerly Merophysiidae Seidlitz, 1872
      • Erotylidae Latreille 1802 (pleasing fungus beetles)
      • Helotidae Reitter 1876
      • Hobartiidae Sen Gupta and Crowson 1966
      • Kateretidae Erichson in Agassiz 1846 (short-winged flower beetles) (= Brachypteridae. ICZN Op. 1916, 1999).
      • Laemophloeidae Ganglbauer 1899 (lined flat bark beetles)
      • Lamingtoniidae Sen Gupta and Crowson 1966
      • Languriidae Crotch , 1873 (lizard beetles)
      • Latridiidae Erichson 1842 (minute brown scavenger beetles)
      • Monotomidae Laporte 1840 (root-eating beetles)
        • Rhizophaginae, formerly Rhizophagidae Redtenbacher 1845
      • Nitidulidae Latreille 1802 (sap-feeding beetles)
      • Passandridae Erichson 1845 (parasitic flat bark beetles)
      • Phalacridae Leach 1815 (shining flower beetles)
      • Phloeostichidae Reitter 1911
      • Propalticidae Crowson 1952
      • Protocucujidae Crowson 1954
      • Silvanidae Kirby 1937 (silvanid flat bark beetles)
      • Smicripidae Horn 1879 (palmetto beetles)
      • Sphindidae Jacquelin du Val 1860 (dry-fungus beetles)
    • Superfamily Curculionoidea Latreille 1802
      • Anthribidae Billberg 1820 (fungus weevils)
      • Attelabidae Billberg 1820 (tooth-nosed snout beetles)
      • Belidae Schönherr 1826 (primitive weevils)
        • Aglycyderinae, formerly Aglycyderidae Wollaston 1864
        • Oxycoryninae, formerly Oxycorynidae Schönherr 1840
      • Brentidae Billberg 1820 (straight-snouted weevils)
        • Apioninae, formerly Apionidae Schönherr, 1823
      • Caridae Thompson 1992
      • Curculionidae Latreille 1802 (snout beetles and true weevils)
        

        

        Citrus root weevil, Diaprepes abbreviatus
        • Scolytinae, formerly Scolytidae Latreille 1807 (typical bark beetles)
      • Ithyceridae Schönherr 1823 (New York weevils)
      • Nemonychidae Bedel 1882 (pine-flower snout beetles)
    • Superfamily Lymexyloidea Fleming 1821
      • Lymexylidae Fleming 1821 (ship-timber beetles)
    • Superfamily Tenebrionoidea Latreille 1802
      • Aderidae Winkler 1927 (antlike leaf beetles)
      • Anthicidae Latreille 1819 (antlike flower beetles)
      • Archeocrypticidae Kaszab 1964
      • Boridae C. G. Thomson 1859 (conifer bark beetles)
      • Chalcodryidae Watt 1974
      • Ciidae Leach 1819 (minute tree-fungus beetles) (= Cisidae)
      • Melandryidae Leach 1815 (false darkling beetles)
      • Meloidae Gyllenhal 1810 (blister beetles)
        

        

        Lytta magister, a blister beetle
      • Mordellidae Latreille 1802 (tumbling flower beetles)
      • Mycetophagidae Leach 1815 (hairy fungus beetles)
      • Mycteridae Blanchard 1845 (palm beetles and flower beetles)
        • Hemipeplinae, formerly Hemipeplidae Lacordaire, 1854
      • Oedemeridae Latreille 1810 (false blister beetles)
      • Perimylopidae St. George 1939
      • Prostomidae C. G. Thomson 1859 (jugular-horned beetles)
      • Pterogeniidae Crowson 1953
      • Pyrochroidae Latreille 1807 (fire-colored beetles)
        • Cononotini or Cononotidae
        • Pedilinae, formerly Pedilidae Lacordaire 1859
      • Pythidae Solier 1834 (dead log bark beetles)
      • Ripiphoridae Gemminger and Harold 1870 (wedge-shaped beetles) formerly Rhipiphoridae
      • Salpingidae Leach 1815 (narrow-waisted bark beetles)
        • Elacatini or Elacatidae
        • Inopeplinae, formerly Inopeplidae Grouvelle 1908
      • Scraptiidae Mulsant 1856 (false flower beetles)
      • Stenotrachelidae C. G. Thomson 1859 (false long-horned beetles)
        • Cephaloinae, formerly Cephaloidae LeConte 1852
      • Synchroidae Lacordaire 1859
      • Tenebrionidae Latreille 1802 (darkling beetles)
        

        

        Heliotautus ruficollis, a darkling beetle
        • Alleculinae, formerly Alleculidae Laporte 1840
        • Lagriinae, formerly Lagriidae Latreille 1825
        • Nilionini or Nilionidae Lacordaire 1859
        • Petriini or Petriidae
      • Tetratomidae Billberg 1820 (polypore fungus beetles)
      • Trachelostenidae Lacordaire 1859
      • Trictenotomidae Blanchard 1845
      • Ulodidae Pascoe 1869
      • Zopheridae Solier 1834 (ironclad beetles)
        • Colydiinae or Colydiini, formerly Colydiidae Erichson 1842
        • Monommatinae or Monommatini, formerly Monommatidae or Monommidae Blanchard 1845
  • Infraorder Elateriformia Crowson 1960
    • Superfamily Buprestoidea Leach 1815
      • Buprestidae Leach 1815 (metallic wood-boring beetles)
        

        

        Emerald ash borer, Agrilus planipennis, a metallic wood-boring beetle
      • Schizopodidae LeConte 1861
    • Superfamily Byrrhoidea Latreille 1804
      • Byrrhidae Latreille 1804 (pill beetles)
      • Callirhipidae Emden 1924 (cedar beetles)
      • Chelonariidae Blanchard 1845 (turtle beetles)
      • Cneoglossidae Champion 1897
      • Dryopidae Billberg 1820 (long-toed water beetles)
      • Elmidae Curtis 1830 (riffle beetles)
      • Eulichadidae Crowson 1973
      • Heteroceridae MacLeay 1825 (variegated marsh-loving beetles)
      • Limnichidae Erichson 1846 (minute marsh-loving beetles)
      • Lutrochidae Kasap and Crowson 1975 (robust marsh-loving beetles)
      • Psephenidae Lacordaire 1854 (water penny beetles)
      • Ptilodactylidae Laporte 1836 (toe-winged beetles)
    • Superfamily Dascilloidea Guerin-Meneville 1843
      • Dascillidae Guérin-Méneville 1843 (1834) (soft-bodied plant beetles)
      • Rhipiceridae Latreille 1834 (cicada parasite beetles)
    • Superfamily Elateroidea Leach 1815
      • Artematopodidae Lacordaire 1857 (soft-bodied plant beetles) = (Eurypogonidae)
      • Brachypsectridae Leconte and Horn 1883 (Texas beetles)
      • Cantharidae Imhoff 1856 (soldier beetles)
      • Cerophytidae Latreille 1834 (rare click beetles)
      • Drilidae Blanchard 1845
      • Elateridae Leach 1815 (click beetles)
        

        

        Alaus murina, a click beetle
        • Cebrioninae, formerly Cebrionidae Latreille 1802
      • Eucnemidae Eschscholtz 1829 (false click beetles)
      • Lampyridae Latreille 1817 (firefly beetles)
      • Lycidae Laporte 1836 (net-winged beetles)
      • Omalisidae Lacordaire 1857
      • Omethidae LeConte 1861 (false firefly beetles)
      • Phengodidae LeConte 1861 (glowworm beetles)
      • Plastoceridae Crowson 1972
      • Podabrocephalidae Pic 1930
      • Rhinorhipidae Lawrence 1988
      • Telegeusidae Leng 1920 (long-lipped beetles)
      • Throscidae Laporte 1840 (false metallic wood-boring beetles) = (Trixagidae)
    • Superfamily Scirtoidea Fleming 1821
      • Clambidae Fischer 1821 (minute beetles)
      • Decliniidae Nikitsky et al 1994
      • Eucinetidae Lacordaire 1857 (plate-thigh beetles)
      • Scirtidae Fleming 1821 (marsh beetles) (= Helodidae)
  • Infraorder Scarabaeiformia Crowson 1960
    • Superfamily Scarabaeoidea Latreille 1802
      • Belohinidae Paulian 1959
      • Bolboceratidae Laporte de Castelnau 1840
      • Ceratocanthidae White 1842 (= Acanthoceridae)
      • Diphyllostomatidae Holloway 1972 (false stag beetles)
      • Geotrupidae Latreille 1802 (earth-boring dung beetles)
      • Glaphyridae MacLeay 1819 (bumble bee scarab beetles)
      • Glaresidae Semenov-Tian-Shanskii and Medvedev 1932 (enigmatic scarab beetles)
      • Hybosoridae Erichson 1847 (scavenging scarab beetles)
      • Lucanidae Latreille 1804 (stag beetles)
        

        

        Lucanus cervus, a stag beetle
      • Ochodaeidae Mulsant and Rey 1871 (sand-loving scarab beetles)
      • Passalidae Leach 1815 (bess beetles)
      • Pleocomidae LeConte 1861 (rain beetles)
      • Scarabaeidae Latreille 1802 (scarab beetles)
        • Dynastinae, formerly Dynastidae MacLeay 1819 (rhinoceros beetles)
      • Trogidae MacLeay 1819 (hide beetles)
  • Infraorder Staphyliniformia Lameere 1900
    • Superfamily Hydrophiloidea Latreille 1802
      • Histeridae Gyllenhal 1808 (clown beetles)
      • Hydrophilidae Latreille 1802 (water scavenger beetles)
        • Georyssinae, formerly Georyssidae Laporte 1840
      • Sphaeritidae Schuckard 1839 (false clown beetles)
      • Synteliidae Lewis 1882
    • Superfamily Staphylinoidea Latreille 1802
      • Agyrtidae C.G. Thomson 1859
      • Hydraenidae Mulsant 1844
      • Leiodidae Fleming 1821 (round fungus beetles) = (Anisotomidae)
        • Platypsyllinae Ritsema 1869 or Leptinidae
      • Ptiliidae Erichson 1845 (feather-winged beetles)
        

        

        Devil's coach horse beetle, Staphylinus olens, a rove beetle
        • Cephaloplectinae, formerly Limulodidae Sharp 1883 (horse-shoe crab beetles)
      • Scydmaenidae Leach 1815 (antlike stone beetles)
      • Silphidae Latreille 1807 (carrion beetles)
      • Staphylinidae Latreille 1802 (rove beetles)
        • Scaphidiinae, formerly Scaphidiidae Latreille 1807
        • Pselaphinae, formerly Pselaphidae Latreille 1802

[edit] References

• Lawrence, J.F., Newton, A.F. Jr. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references, and data on family-group names), pp. 779-1006. In: Pakaluk, J., Slipinski, S.A. (eds.), Biology, phylogeny, and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson. Muzeum i Instytut Zoologii PAN, Warszawa.
• Ross H. Arnett, Jr. and Michael C. Thomas, American Beetles (CRC Press, 2001–2002)