Dionycha
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The Dionycha are a phylogenetic group of spiders (Araneomorphae:Entelegynae) with about 10,000 species. 17 families belong to this group, amongst others: Salticidae (jumping spiders), Gnaphosoidea, Thomisidae (crab spiders), and the Clubionidae. Spiders in this group have better senses (sight, hearing) than others, some even show courtship dances and songs.
The Dionycha probably evolved from the web-building Trionycha. Unlike these, the Dionycha have only two instead of three tarsal claws. The central third claw of the Trionycha is responsible for tracking the silk thread. Instead of this claw, Trionycha have dense tufts of scupula hairs that produce strong adhesion, enabling some species to climb glass. Most species hunt their prey instead of building webs, although some species of Trionycha (members of Lycosoidea) also hunt.
There are no cribellate members in the Dionycha. (Griswold et. al. 1999)
The monophyly of the Dionycha has been disputed. Today it is thought that the reduction of the third claw evolved several times independently.
[edit] Cladogram (after Coddington und Levi, 1991, simplified)
|-Araneomorphae |--| |-Haplogynae |__/-- Corinnidae |-Entelegynae |--| \-- Liocranidae |-Trionycha | |----- Gnaphosoidea (7 families) |-Dionycha-------|--| |-- Clubionidae, over 2000 species | |-----|-- Anyphaenidae | |-- Salticidae, > 4000 species |----------- Thomisidae, > 3000 species |----------- Philodromidae |----------- Sparassidae |----------- Selenopidae |----------- Zoridae
[edit] References
- J. Bosselaers and R. Jocqué. 2002. Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae. Zoologica Scripta 31:241-270.
- J. A. Coddington and H. W. Levi. 1991. Systematics and evolution of spiders (Araneae). Annual Review of Ecology and Systematics 22:565-592.
- Griswold, C.E., Coddington, J.A., Platnick, N.I. and Forster, R.R. (1999). Towards a Phylogeny of Entelegyne Spiders (Araneae, Araneomorphae, Entelegynae). Journal of Arachnology 27:53-63. PDF